BIOMASS AND NUTRIENT DISTRIBUTION IN TWO OLD ... - Climate

BIOMASS AND N U T R I E N T D I S T R I B U T I O N I N TWO

OLD GROWTH FOREST ECOSYSTEMS I N

SOUTH COASTAL B R I T I S H COLUMBIA

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I n p r e s e n t i n g t h i s t h e s i s i n p a r t i a l f u l f i l l m e n t o f t h e r e q u i r e -

m e n t s f o r a n a d v a n c e d d e g r e e a t t h e U n i v e r s i t y o f B r i t i s h C o l u m b i a , I

a g r e e t h a t t h e L i b r a r y s h a l l m a ke i t f r e e l y a v a i l a b l e f o r r e f e r e n c e a n d

s t u d y . I f u r t h e r a g r e e t h a t p e r m i s s i o n f o r e x t e n s i v e c o p y i n g o f t h i s

t h e s i s f o r s c h o l a r l y p u r p o s e s may b e g r a n t e d b y t h e H e a d o f my D e p a r t m e n t

o r b y h i s r e p r e s e n t a t i v e s . I t i s u n d e r s t o o d t h a t c o p y i n g o r p u b l i c a t i o n

o f t h i s t h e s i s f o r f i n a n c i a l g a i n s h a l l n o t b e a l l o w e d w i t h o u t my w r i t t e n

p e r m i s s i o n .

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T h e d i s t r i b u t i o n o f t h e a b o v e - g r o u n d b i o m a s s a n d m a c r o n u t r i e n t

c o n t e n t o f t h e t r e e s w e r e s t u d i e d o n t w o s a m p l e p l o t s i n s o u t h c o a s t a l

B r i t i s h C o l u m b i a , n e a r V a n c o u v e r . T h e p l o t s d i f f e r e d i n e l e v a t i o n ( 4 , 6 0 0

a n d 2 , 2 0 0 f e e t ; 1,500 a n d 7 0 0 m ) , i n s o i l t y p e a n d d e p t h . T r e e a g e w a s

s i m i l a r i n b o t h p l o t s , r a n g i n g f r o m 1 5 0 t o 5 3 0 y e a r s . T h e t r e e c o v e r o n

t h e h i g h e l e v a t i o n p l o t c o n s i s t e d o f T s u g a m e r t e n s i a n a ( B o n g . ) C a r r . ( m o u n -

t a i n h e m l o c k ) a n d A b i e s a m a b i l i s ( D o u g l . ) F o r b e s ( P a c i f i c s i l v e r f i r ) w h i l e

t h e l o w e r e l e v a t i o n p l o t w a s o c c u p i e d b y T s u g a h e t e r o p h y l l a ( R a f . ) S a r g .

( w e s t e r n h e m l o c k ) , T h u j a p l i c a t a D. D o n ( w e s t e r n r e d c e d a r ) a n d C h a m a e c y -

p a r i s n o o t k a t e n s i s ( D . D o n ) S p a c h ( y e l l o w c e d a r ) . T w e n t y - f o u r t r e e s w e r e

s a m p l e d t o d e t e r m i n e t h e b i o m a s s a n d n u t r i e n t c o n t e n t o f w o o d , b a r k ,

b r a n c h e s , t w i g s , f o l i a g e a n d c o n e s . A n o t h e r n i n e t r e e s w e r e s a m p l e d f o r

t h e b i o m a s s a n d n u t r i e n t c o n t e n t o f w o o d a n d b a r k o n l y .

M u l t i p l e r e g r e s s i o n a n a l y s i s w a s u s e d t o e s t a b l i s h t h e r e l a t i o n -

s h i p b e t w e e n d . b . h . , t r e e l e n g t h , c r o w n l e n g t h a n d b i o m a s s o f t h e v a r i o u s

t r e e c o m p o n e n t s . T h e r e g r e s s i o n e q u a t i o n s o b t a i n e d w e r e u s e d t o e s t i m a t e

t h e t o t a l b i o m a s s o f w o o d , b a r k , b r a n c h e s , t w i g s a n d f o l i a g e c o n t a i n e d i n

t h e t r e e s o n e a c h o f t h e s a m p l e p l o t s . T h e d a t a t h u s o b t a i n e d w e r e com-

b i n e d w i t h d a t a o n c h e m i c a l c o n c e n t r a t i o n a n d u s e d t o e s t i m a t e t h e d i s t r i -

b u t i o n o f m a c r o n u t r i e n t e l e m e n t s i n d i f f e r e n t a b o v e - g r o u n d b i o m a s s c o m p o n -

e n t s o f t h e s t a n d s .

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T A B L E OF CONTENTS

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T I T L E PAGE i

ABSTRACT i i i

T A B L E OF CONTENTS i v

L I S T OF T A B L E S , v *

L I S T OF F I G U R E S , i x

L I S T OF A P P E N D I C E S t X

ACKNOWLEDGEMENTS , . x i

I INTRODUCTION 1

O b j e c t i v e s o f t h e s i s . . . . . . . 2

I I L I T E R A T U R E REVIEW 3

N u t r i e n t u p t a k e b y t r e e s a n d n u t r i e n t i m m o b i l i z a -t i o n i n t r e e b i o m a s s . . . . . 3

N u t r i e n t d i s t r i b u t i o n i n a b o v e - g r o u n d t r e e b i o m a s s 8

R e m o v a l o f n u t r i e n t s f r o m a s i t e a s t h e r e s u l t o f t r e e h a r v e s t i n g 17

I I I D E S C R I P T I O N OF SAMPLE P L O T S 19

L o c a t i o n a n d s i z e 19

C l i m a t e . . . . . . . . . . . . 24

S o i l s 2 6

V e g e t a t i o n . . . . . . . . . 26

I V METHODS 2 9

F i e l d s a m p l i n g . . . . . . . 29

L a b o r a t o r y a n a l y s e s 3 3

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P a g e

I V METHODS ( c o n t i n u e d )

D a t a a n a l y s e s . . . . . . f . . . . . . . . . . . . 3 8

L i m i t a t i o n o n d a t a i n t e r p r e t a t i o n . . . . . . . . . 47

M i s s i n g d a t a a n d a d j u s t m e n t o f i n c o r r e c t d a t a . . . 48

V R E S U L T S AND D I S C U S S I O N 5 0

B i o m a s s d i s t r i b u t i o n 5 0

N u t r i e n t d i s t r i b u t i o n w i t h i n t r e e l a y e r 57

R e m o v a l o f m a c r o n u t r i e n t s f r o m t h e e c o s y s t e m s d u e t o d i f f e r e n t l e v e l i o f t r e e u t i l i z a t i o n 73

V I SUMMARY AND CONCLUSIONS 77

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L I S T OF T A B L E S

T A B L E P a g e

1 T o t a l q u a n t i t i e s o f n u t r i e n t s i m m o b i l i z e d w i t h i n a l l o r g a n s o f t h e m a i n g r o w i n g c r o p o f v a r i o u s t r e e s t a n d s a t a g e 1 0 0 y e a r s ( R e n n i e , 1 9 5 5 ) . 4

2 T h e t o t a l q u a n t i t i e s o f n u t r i e n t s t a k e n u p b y 3 t y p e s o f t i m b e r - p r o d u c i n g f o r e s t a t a g e 1 0 0 y e a r s . ( R e n n i e , 1 9 5 5 ) . . 5

3 T h e p e r c e n t a g e d i s t r i b u t i o n o f n u t r i e n t s i m m o b i l i z e d i n t h e c o m p o n e n t o r g a n s o f t h e m a i n g r o w i n g c r o p o f t h r e e t y p e s o f s t a n d a t a g e 1 0 0 y e a r s ( R e n n i e , 1 9 5 5 ) . 9

4 C o n c e n t r a t i o n o f m a c r o n u t r i e n t s i n f o l i a g e o f u p p e r w h o r l s o f t r e e c r o w n f o r t h r e e s p e c i e s i n B. C. ( B e a t o n e t a l . , 1 9 6 5 ) . 11

5 A v e r a g e N - c o n c e n t r a t i o n s i n v a r i o u s l o b l o l l y p i n e t i s s u e s a t v a r i o u s a g e s o n g o o d s i t e s i n p e r c e n t o f o v e n - d r y w e i g h t ( S w i t z e r a n d N e l s o n , 1 9 6 4 ) . 12

6 T h e r a n g e o f n u t r i e n t c o n c e n t r a t i o n s ( i n p e r c e n t ) i n d i f f e r e n t t r e e c o m p o n e n t s i n v a r i o u s N o r w a y s p r u c e s t a n d s i n C z e c h o s l o v a k i a ( M a t e r n a , 1 9 6 4 ) . 13

7 D i s t r i b u t i o n o f N, P, K a n d C a among t h e n a j o r c o m p o n -e n t s o f a 3 6 - y e a r o l d s e c o n d - g r o w t h D o u g l a s - f i r f o r e s t ( C o l e , 1 9 6 7 ) . 14

8 D i s t r i b u t i o n o f b i o m a s s a n d n u t r i e n t s w i t h i n t h e a b o v e - g r o u n d t r e e c o m p o n e n t s o f a 1 2 - y e a r o l d s t a n d o f r a d i a t a p i n e , W h a k a r e w a r e w a F o r e s t ( W i l l , 1 9 6 4 ) . 15

9 A v e r a g e p e r c e n t a g e d i s t r i b u t i o n o f N , P, K, C a a n d Mg i n a e r i a l c o m p o n e n t s o f 12 n a t u r a l j a c k p i n e s t a n d s i n N o r t h e r n O n t a r i o ( M o r r i s o n , 1973)., 16

1 0 Summary o f c l i m a t i c i n f o r m a t i o n ( K r a j i n a , 1 9 6 5 ) . 25 <

11 D i a m e t e r c l a s s d i s t r i b u t i o n o n t h e s a m p l e p l o t s . 3 0

12 B a s i c s t a t i s t i c s o f o r i g i n a l m e a s u r e m e n t s f o r a l l t r e e s o v e r 6 i n c h e s o f d . b . h . o n t h e t w o s a m p l e p l o t s . 31

( v i i )

T A B L E P a g e

13 P e r c e n t l o s s o f w e i g h t i n s a m p l e s f o r c h e m i c a l a n a l y s e s u p o n r e - d r y i n g a f t e r 2^-6 m o n t h s o f s t o r a g e , 34

14 C o m p a r i s o n o f r e s u l t s o f c h e m i c a l a n a l y s e s f o r f i v e r a n d o m l y s e l e c t e d s a m p l e s . 3 8

15 A v e r a g e p e r c e n t d i f f e r e n c e b e t w e e n t r e e t r u n k v o l u m e s c a l c u l a t e d a c c o r d i n g t o B C F S v o l u m e e q u a t i o n s a n d t h o s e c a l c u l a t e d s e c t i o n - b y - s e c t i o n a c c o r d i n g t o N e w t o n ' s f o r m u l a , 4 0

16 B a s i c s t a t i s t i c s f o r s a m p l e d t r e e s , 4 1

17 B i o m a s s d i s t r i b u t i o n f o r f o u r t r e e s p e c i e s o n t h e S q u a m i s h a n d H a n e y s a m p l e p l o t s ( i n k g / h a ) . 51

18 A v e r a g e b i o m a s s p r o p o r t i o n s o f d i f f e r e n t t r e e c o m -p o n e n t s f o r f o u r t r e e s p e c i e s o n t h e S q u a m i s h a n d H a n e y s a m p l e p l o t s . 5 3

19 M e a n c o n c e n t r a t i o n ( p e r c e n t a g e o v e r - d r y w e i g h t ) o f N, P, K, C a a n d Mg i n c o m p o n e n t s a n d s u b - c o m p o n e n t s o f P a c i f i c s i l v e r f i r s a m p l e - t r e e s . 5 8

2 0 M e a n c o n c e n t r a t i o n ( p e r c e n t a g e o v e r - d r y w e i g h t ) o f N , P, K, C a a n d Mg i n c o m p o n e n t s a n d s u b - c o m p o n e n t s o f m o u n t a i n h e m l o c k s a m p l e t r e e s . 5 9

21 M e a n c o n c e n t r a t i o n ( p e r c e n t a g e o v e n - d r y w e i g h t ) o f N, P, K, C a a n d Mg i n c o m p o n e n t s a n d s u b - c o m p o n e n t s o f w e s t e r n h e m l o c k s a m p l e t r e e s , 6 0

22 M e a n c o n c e n t r a t i o n ( p e r c e n t a g e o v e n - d r y w e i g h t ) o f N, P, K, C a a n d Mg i n c o m p o n e n t s a n d s u b - c o m p o n e n t s o f y e l l o w c e d a r s a m p l e t r e e s , 6 1

23 C o n c e n t r a t i o n o f m a c r o n u t r i e n t s i n f o l i a g e w i t h i n u p p e r t h i r d o f t r e e c r o w n , 65

24 D i s t r i b u t i o n o f m a c r o n u t r i e n t s b y p a r t i c u l a r t r e e c o m p o n e n t s o n t h e S q u a m i s h s a m p l e p l o t . 67

25 D i s t r i b u t i o n o f m a c r o n u t r i e n t s b y p a r t i c u l a r t r e e c o m p o n e n t s o n t h e H a n e y s a m p l e p l o t . 6 8

( v i i i )

T A B L E P a g e

26 A v e r a g e p r o p o r t i o n s o f f i y e m a c r o n u t r i e n t s i n d i f V f e r e n t t r e e c o m p o n e n t s o n t h e S q u a m i s h a n d H a n e y s a m p l e p l o t s . 69

27 T h e a m o u n t o f m a c r o n u t r i e n t s i m m o b i l i z e d i n a b o v e -g r o u n d t r e e b i o m a s s b y v a r i o u s s p e c i e s i n d i f f e r e n t c o u n t r i e s . 72

28 R e m o v a l o f b i o m a s s a n d m a c r o n u t r i e n t s a c c o m p a n y i n g d i f f e r e n t l e v e l s o f u t i l i z a t i o n i n t h e S q u a m i s h s a m p l e p l o t . 74

29 R e m o v a l o f b i o m a s s a n d m a c r o n u t r i e n t s a c c o m p a n y i n g d i f f e r e n t l e v e l s o f u t i l i z a t i o n i n t h e H a n e y s a m p l e p l o t . 75

L I S T OF F I G U R E S

F I G U R E P a g e

V a n c o u v e r a n d v i c i n i t y - l o c a t i o n o f t h e s a m p l e p l o t s . 20

L I S T OF A P P E N D I C E S

P a g e

S c i e n t i f i c a n d common nam e s o f t r e e s p e c i e s . 3 8

B a s i c s t a t i s t i c s f o r t h e i n d e p e n d e n t v a r i a b l e s . 90

S i g n i f i c a n t i n d e p e n d e n t v a r i a b l e s i n s i m p l e l i n e a r a n d l o g a r i t h m i c r e g r e s s i o n e q u a t i o n s f o r b i o m a s s p r e d i c t i o n o f p a r t i c u l a r t r e e c o m p o n e n t s .

B i o m a s s d i s t r i b u t i o n f o r 2 0 s a m p l e t r e e s o n t h e S q u a m i s h s a m p l e p l o t ,

B i o m a s s d i s t r i b u t i o n f o r 13 s a m p l e t r e e s o n t h e H a n e y s a m p l e p l o t .

92

99

110

N u t r i e n t c o n c e n t r a t i o n a n d n u t r i e n t d i s t r i b u t i o n w i t h i n 2 0 s a m p l e t r e e s o n t h e S q u a m i s h s a m p l e p l o t . 1 1 8

N u t r i e n t c o n c e n t r a t i o n a n d n u t r i e n t d i s t r i b u t i o n w i t h i n 13 s a m p l e t r e e s o n t h e H a n e y s a m p l e p l o t . 139

P r o g r a m f o r c a l c u l a t i n g m o i s t u r e c o n t e n t o f s a m p l e s f o r c h e m i c a l a n a l y s e s . 1 5 3

S p e c i f i c g r a v i t y p r o g r a m . 157

P r o g r a m f o r c a l c u l a t i n g b i o m a s s d i s t r i b u t i o n f o r s a m p l e d t r e e s . 1 5 9

P r o g r a m f o r c a l c u l a t i n g n u t r i e n t d i s t r i b u t i o n . 1 6 3

171

P r o g r a m f o r c a l c u l a t i n g w e i g h t e d c o n c e n t r a t i o n s o f n u t r i e n t s w i t h i n d i f f e r e n t c o m p o n e n t s o f s a m p l e t r e e s .

P r o g r a m f o r c a l c u l a t i n g b i o m a s s f o r 2 5 6 t r e e s o n t h e S q u a m i s h s a m p l e p l o t a c c o r d i n g t o l o g a r i t h m i c r e g r e s s i o n e q u a t i o n s . 1 7 7

( x i )

ACKNOWLEDGEMENTS

T h e w r i t e r w i s h e s t o e x p r e s s h i s s i n c e r e t h a n k s t o D r . J . P.

K i m m i n s f o r p r o v i d i n g t h e o p p o r t u n i t y t o u n d e r t a k e t h i s p r o j e c t a n d f o r

h i s g u i d a n c e , a d v i c e a n d c r i t i c a l r e v i e w ; t o D r . T. B a l l a r d f o r n u m e r o u s

h o u r s o f d i s c u s s i o n , h e l p w i t h d a t a i n t e r p r e t a t i o n a n d d r a w i n g c o n c l u s i o n s ;

t o D r s . A. K o z a k a n d J . D e m a e r s c h a l k f o r h e l p w i t h s t a t i s t i c a l a n a l y s i s ;

t o D r s . P. H a d d o c k a n d J . W o r r a l l f o r c r i t i c a l r e v i e w o f t h e m a n u s c r i p t ;

t o D r . R. C. B r o o k e o f S i m o n F r a s e r U n i v e r s i t y a n d M r . K a r e l K l i n k a f o r

h e l p i n s i t e i d e n t i f i c a t i o n a n d s e l e c t i o n ; t o M e s s r s . J o h n P a r m i n t e r a n d

G r a n t S c h e i r m a n f o r h e l p w i t h f i e l d w o r k ; t o M r . A u s t i n P h i r i f o r s o i l

s a m p l i n g a n d l a b o r a t o r y a n a l y s e s ; t o M r . K e n T w a i t e s f o r h e l p w i t h l a b o r a -

t o r y a n a l y s e s a n d c o m p u t e r p r o g r a m m i n g ; t o M r . M i n T s z e f o r h e l p w i t h

l a b o r a t o r y a n a l y s e s ; t o M i s s L . C o w d e l l f o r h e l p w i t h c o m p u t e r p r o g r a m m i n g ;

t o M e s s r s . D o n G r a n t a n d T a g N e i g h b o u r o f t h e B C F S f o r h e l p i n s i t e l o c a -

t i o n a n d o t h e r p r a c t i c a l d e t a i l s ; t o t h e s t a f f o f t h e UBC R e s e a r c h F o r e s t

a t H a n e y ; t o M a c M i l l a n B l o e d e l l , S q u a m i s h O f f i c e , f o r c o o p e r a t i o n i n a c c e s s

t o t h e h i g h e l e v a t i o n s i t e a n d h e l p i n f a l l i n g t r e e s .

T h e h e l p o f t h e s e p e o p l e a n d o r g a n i z a t i o n s , t o g e t h e r w i t h f i n a n -

c i a l s u p p o r t o f t h e B C F S P r o d u c t i v i t y C o m m i t t e e made t h i s t h e s i s p o s s i b l e .

1

I INTRODUCTION

M u c h o f t h e l o g g i n g i n c o a s t a l B r i t i s h C o l u m b i a i n t h e p a s t w a s

i n t h e m o r e a c c e s s i b l e l o w e l e v a t i o n s i t e s . M o r e r e c e n t l y , t h e l o g g e r ' s

a t t e n t i o n h a s b e e n t u r n e d t o w a r d s h i g h e r e l e v a t i o n f o r e s t s , a n d b o t h o n

t h e c o a s t a n d i n t h e i n t e r i o r , s u b a l p i n e f o r e s t s a r e b e i n g c u t o n a n i n -

c r e a s i n g s c a l e . B e c a u s e o f t h e i r r e l a t i v e i n a c c e s s i b i l i t y a n d t h e i r r e l a -

t i v e l y r e c e n t u t i l i z a t i o n , m u c h l e s s i s k n o w n a b o u t t h e m t h a n t h e l o w e r

e l e v a t i o n f o r e s t s . L i t t l e i s k n o w n a b o u t t h e i r s o i l s , t h e i r h u m u s , t h e i r

c l i m a t e , t h e i r r e s p o n s e t o h a r v e s t i n g , t h e s u s t a i n a b i l i t y o f t h e f o r e s t s ,

t h e s u i t a b i l i t y o f t h e c l e a r c u t h a r v e s t i n g m e t h o d s , a n d t h e s o l u t i o n t o

t h e p r o b l e m s o f r e g e n e r a t i n g h i g h - e l e v a t i o n c u t o v e r a r e a s . S i n c e s u b a l p i n e

f o r e s t s r e p r e s e n t a v e r y s u b s t a n t i a l a r e a o f c o a s t a l f o r e s t s ( 1 1 . 3 m i l l i o n

a c r e s c o m p a r e d w i t h 2 6 . 3 m i l l i o n o f o t h e r f o r e s t t y p e s o n t h e c o a s t ( S m i t h ,

1 9 7 4 ) ) t h e r e i s a n u r g e n t n e e d f o r a l a r g e v a r i e t y o f s t u d i e s t o b e u n d e r -

t a k e n i n t h e s e f o r e s t s a s a f o u n d a t i o n f o r t h e i r r a t i o n a l m a n a g e m e n t .

A l t h o u g h r e l a t i v e l y l i t t l e i s k n o w n a b o u t t h e s e f o r e s t s , i n t e r e s t

i n t h e m h a s g r o w n o v e r t h e p a s t d e c a d e .

A s u b s t a n t i a l c o n t r i b u t i o n t o o u r u n d e r s t a n d i n g o f t h e e c o l o g y o f

t h e c o a s t a l s u b a l p i n e f o r e s t e x i s t s i n t h e f o r m o f t h e s t u d y b y B r o o k e ,

P e t e r s o n a n d K r a j i n a ( 1 9 7 0 ) , T h e B r i t i s h C o l u m b i a F o r e s t S e r v i c e h a s u n d e r -

t a k e n a r e v i e w o f h i g h e l e v a t i o n r e g e n e r a t i o n p r o b l e m s , a n d i s c u r r e n t l y

i n v e s t i g a t i n g a n u m b e r o f a s p e c t s o f h i g h e l e v a t i o n l o g g i n g . T h e w o r k r e -

p o r t e d h e r e i n w a s f u n d e d b y t h e B, C, F o r e s t S e r v i c e P r o d u c t i v i t y C o m m i t t e e

2

a n d r e f l e c t s i t s c o n c e r n f o r a b e t t e r u n d e r s t a n d i n g o f t h e e c o l o g i c a l a n d

p r a c t i c a l a s p e c t s o f h i g h - e l e v a t i o n c o a s t a l l o g g i n g .

T h i s s t u d y w a s a p a r t o f a n i n v e s t i g a t i o n o f t h e p o t e n t i a l f o r

s i t e d e g r a d a t i o n t h r o u g h n u t r i e n t d e p l e t i o n b y h a r v e s t i n g . T o i n v e s t i g a t e

t h e f u l l s i g n i f i c a n c e o f n u t r i e n t l o s s e s a c c o m p a n y i n g f o r e s t h a r v e s t i n g i s

b e y o n d t h e s c o p e o f t h i s s t u d y . T h i s t h e s i s w a s r e s t r i c t e d t o t h e f o l l o w i n g

o b j e c t i v e s :

1. T o q u a n t i f y t h e a b o v e - g r o u n d t r e e b i o m a s s b y d i f f e r e n t s t e m

a n d c r o w n b i o m a s s c o m p o n e n t s .

2. T o q u a n t i f y t h e m a c r o n u t r i e n t ( N , P, K, C a , Mg) c o n t e n t o f

t h e s e b i o m a s s c o m p o n e n t s .

3. To c a l c u l a t e t h e q u a n t i t y o f n u t r i e n t s r e m o v e d f r o m t h e s i t e

i n h a r v e s t e d m a t e r i a l s a t d i f f e r e n t l e v e l s o f u t i l i z a t i o n .

T h e s t u d y t h u s a d o p t s t h e s o - c a l l e d " s u b s t i t u t i o n p r i n c i p l e "

(Tamm, 1 9 6 4 ) m e t h o d o f a s s e s s i n g s i t e n u t r i e n t d e p l e t i o n a c c o m p a n y i n g t h e

r e m o v a l o f f o r e s t p r o d u c t s .

3

I I L I T E R A T U R E REVIEW

T h e r e v i e w e f t h e l i t e r a t u r e h a s b e e n o r g a n i z e d i n t o t h r e e m a j o r

t o p i c s :

1. N u t r i e n t u p t a k e b y t r e e s a n d n u t r i e n t i m m o b i l i z a t i o n

i n t r e e b i o m a s s .

2. N u t r i e n t d i s t r i b u t i o n i n a b o v e - g r o u n d t r e e b i o m a s s .

3. R e m o v a l o f n u t r i e n t s f r o m a s i t e a s t h e r e s u l t o f h a r -

v e s t i n g .

2 . 1 : N u t r i e n t u p t a k e b y t r e e s a n d n u t r i e n t i m m o b i l i z a t i o n i n t r e e

b i o m a s s .

S i n c e t h e m i d d l e o f t h e l a s t c e n t u r y t h e r e h a v e b e e n s e v e r a l a t -

t e m p t s t o e s t i m a t e t h e a m o u n t o f n u t r i e n t s r e q u i r e d f o r o p t i m a l t r e e g r o w t h

( E b e r m a y e r , 1 8 7 6 ) . M o r e r e c e n t l y e f f o r t s h a v e b e e n made t o d e t e r m i n e t h e

t o t a l n u t r i e n t d e m a n d o n a f o r e s t s i t e r e s u l t i n g f r o m n e t n u t r i e n t u p t a k e

b y t r e e s w h i c h a r e s u b s e q u e n t l y h a r v e s t e d . A v e r y c o m p r e h e n s i v e r e v i e w o f

t h e l i t e r a t u r e o n t h i s t o p i c , i n c l u d i n g n u t r i e n t u p t a k e a n d i m m o b i l i z a t i o n ,

w a s made b y R e n n i e ( 1 9 5 5 ) . He r e v i e w e d p u b l i c a t i o n s f r o m t h e s e c o n d h a l f

o f t h e l a s t c e n t u r y a n d f r o m t h e f i r s t h a l f o f t h i s c e n t u r y . B a s e d u p o n

t h i s l i t e r a t u r e t h e t o t a l q u a n t i t i e s o f n u t r i e n t s t a k e n u p a n d i m m o b i l i z e d

w i t h i n a l l o r g a n s o f t h e m a i n g r o w i n g c r o p o f t h r e e t y p e s o f f o r e s t s t a n d

w e r e e s t i m a t e d a n d a r e p r e s e n t e d i n T a b l e 1.

4

T A B L E 1: T o t a l q u a n t i t i e s o f n u t r i e n t s i m m o b i l i z e d w i t h i n a l l o r g a n s o f t h e m a i n g r o w i n g c r o p o f v a r i o u s t r e e s t a n d s a t a g e 1 0 0 y e a r s ( R e n r i i e , 1 9 5 5 ) .

T o t a l n u t r i e n t s i m m o b i l i z e d w i t h i n t h e m a i n g r o w i n g c r o p , k g e l e m e n t p e r a c r e .

S p e c i e s C r o p A f t e r 5 0 y e a r s ' g r o w t h A f t e r 1 0 0 y e a r s ' g r o w t h C a K P C a K P

P i n e s 98 5 7 1 3 1 1 5 5 6 1 2 O t h e r c o n i f e r s 2 0 8 1 1 3 2 5 2 7 4 1 5 2 28 H a r d w o o d s 3 0 2 8 2 19 5 2 0 1 2 9 2 8

C o m m e n t i n g o n t h e v a l i d i t y o f t h e s e n u t r i e n t e s t i m a t e s , R e n n i e

s t a t e d : " P a r t i c u l a r e s t i m a t e s m i g h t s h o w a p p r e c i a b l e c u m u l a t i v e e r r o r .

B u t f o r many i n d i v i d u a l e s t i m a t e s a n d f o r t h e m e a n e s t i m a t e s r e p r e s e n t i n g

t h e t h r e e s p e c i e s g r o u p s t h e v a r i a t i o n i n u p t a k e s e e n w o u l d , i n t h e m a i n , b e

g e n u i n e a n d d u e t o d i f f e r e n c e s i n s p e c i e s g r o w t h - r a t e a n d s i t e . " T h u s , a l -

t h o u g h R e n n i e ' s c o n c l u s i o n s w e r e b a s e d o n o l d a n d s c a t t e r e d d a t a , h e b e -

l i e v e d t h e y w e r e a t l e a s t a p p r o x i m a t e l y c o r r e c t .

C o n s i d e r a b l e q u a n t i t i e s o f n u t r i e n t s , p a r t i c u l a r l y C a , may b e

t a k e n u p a n d i m m o b i l i z e d w i t h i n t h e v a r i o u s t r e e o r g a n s o f a g r o w i n g f o r e s t .

T h e r e l a t i v e l y s m a l l i n c r e a s e i n n u t r i e n t i m m o b i l i z a t i o n f r e q u e n t l y o c c u r -

r i n g f r o m 5 0 t o 1 0 0 y e a r s ( s e e T a b l e 1 ) s u g g e s t s t h a t n u t r i e n t i m m o b i l i z a -

t i o n may r e a c h a n u p p e r l i m i t a n d t h a t t h e s o l e n u t r i e n t m o v e m e n t t h e r e -

a f t e r i s a c o m p a r a t i v e l y s m a l l n u t r i e n t c y c l e a s s o c i a t e d w i t h f o l i a r f l u s h

a n d f a l l . T h e same c o n c l u s i o n w a s r e a c h e d b y S w i t z e r a n d N e l s o n ( 1 9 6 4 ) .

5

T h e y f o u n d t h a t t h e r a t e s o f u p t a k e o f a l l n u t r i e n t s a r e g r e a t e s t d u r i n g

t h e e a r l y l i f e o f t h e s t a n d a n d t h a t b y a g e 4 5 y e a r s n u t r i t i o n i s m o s t l y

a m a t t e r o f c y c l i n g .

T a b l e 1 o n l y r e f e r s t o n u t r i e n t s i m m o b i l i z e d i n t h e f i n a l c r o p ,

h o w e v e r . I f we i n c l u d e t h e a m o u n t o f n u t r i e n t s r e m o v e d i n t h i n n i n g s , t h e

t o t a l a m o u n t o f n u t r i e n t s i m m o b i l i z e d o v e r t h e r o t a t i o n i n c r e a s e s c o n s i d e r -

a b l y . T h i s c a n b e s e e n b y a c o m p a r i s o n b e t w e e n T a b l e 1 a n d T a b l e 2, T h e

l a t t e r s h o w s t h e t o t a l q u a n t i t i e s o f n u t r i e n t s t a k e n u p b y t h e s t a n d t y p e s

r e f e r r e d t o i n T a b l e 1, T h u s , t h e q u a n t i t y o f n u t r i e n t s r e m o v e d i n h a r -

v e s t i n g i s l i k e l y t o i n c r e a s e a s u t i l i z a t i o n s t a n d a r d s a n d i n t e n s i t y o f

m a n a g e m e n t i n c r e a s e , a n d r o t a t i o n l e n g t h d e c r e a s e .

T A B L E 2: T h e t o t a l q u a n t i t i e s o f n u t r i e n t s t a k e n u p b y 3 t y p e s o f t i m b e r -p r o d u c i n g f o r e s t a t a g e 1 0 0 y e a r s ( R e n n i e , 1 9 5 5 ) .

T o t a l n u t r i e n t u p t a k e ( e x c l . f o l i a g e a n d r o o t s o f t h i n n i n g s a n d l i t t e r ) , k g . e l e m e n t p e r a c r e .

S p e c i e s C r o p C o m p o n e n t A f t e r 5 0 y e a r s ' g r o w t h A f t e r 1 0 0 y e a r s ' g r o w t l

C a K P C a K P

t i m b e r t h i n n i n g s 35 15 4 88 3 5 9

P i n e s m a i n c r o p 98 57 1 3 1 1 5 56 12 t o t a l 1 3 3 72 17 2 0 3 9 1 2 1

t i m b e r O t h e r t h i n n i n g s 56 26 5 1 6 4 8 2 1 3 c o n i f e r s m a i n c r o p 2 0 8 1 1 3 25 2 7 4 1 5 2 28

t o t a l 2 6 4 1 3 9 3 0 4 3 8 2 3 4 4 1

t i m b e r t h i n n i n g s 1 6 1 44 1 0 3 5 9 96 22

H a r d w o o d s m a i n c r o p 3 0 2 82 1 9 5 2 0 1 2 9 2 8 t o t a l 4 6 3 1 2 6 29 8 7 9 2 2 5 5 0

6

T h e r e a r e many o t h e r s t u d i e s i n w h i c h n u t r i e n t u p t a k e a n d i m -

m o b i l i z a t i o n o f n u t r i e n t s b y f o r e s t h a v e b e e n d o c u m e n t e d . V i r o ( 1 9 5 6 )

c a l c u l a t e d t h e a v e r a g e a n n u a l u p t a k e f o r v a r i o u s p i n e s t a n d s i n F i n l a n d

a v e r a g i n g a b o u t 1 0 0 y e a r s i n a g e t o b e 7.36 k g o f N, 0.58 k g o f P, 0.60 k g

o f K, 8.4 k g o f C a a n d 0.87 k g o f Mg p e r h e c t a r e . T h e g r e a t e r p a r t o f

t h e s e n u t r i e n t s i s r e t u r n e d t o t h e f o r e s t f l o o r i n t h e f o r m o f l i t t e r a n d

t h r o u g h f a l l , h o w e v e r , a n d o n l y p a r t o f t h e t o t a l u p t a k e i s i m m o b i l i z e d i n

t h e t r e e b i o m a s s .

W r i g h t a n d W i l l ( 1 9 5 8 ) c a l c u l a t e d t h e a m o u n t o f n u t r i e n t s b e i n g

i m m o b i l i z e d i n a 6 4 - y e a r o l d s t a n d o f S c o t s p i n e . * T o t a l d r y w e i g h t o f

t h e b i o m a s s w a s 1 1 9 , 0 0 0 k g / h a , c o n t a i n i n g 2 0 5 k g o f N, 2 3 k g o f P, 1 0 0 k g

o f K, 2 0 2 k g o f C a a n d 3 5 k g o f Mg p e r h e c t a r e .

W i l l ( 1 9 6 4 ) . c a l c u l a t e d t h e t o t a l n u t r i e n t r e m o v a l d u e t o t h i n n i n g

a n d c l e a r - f e l l i n g f o r a 3 5 - y e a r o l d s t a n d o f r a d i a t a p i n e . T h e a m o u n t o f

d r y b i o m a s s h a r v e s t e d w a s 3 6 2 , 0 0 0 k g / h a c o n t a i n i n g 2 3 0 k g o f N, 3 2 k g o f P,

2 8 3 k g o f K a n d 1 8 8 k g o f C a p e r h e c t a r e . I n c o n t r a s t , a 3 5 - y e a r o l d s t a n d

o f C o r s i c a n p i n e i n E n g l a n d ( C u l b i n f o r e s t , M o r a y s h i r e ) w a s f o u n d t o c o n -

t a i n c o n s i d e r a b l y m o r e n u t r i e n t s t h a n t h i s i n s p i t e o f i t s s m a l l e r a m o u n t

o f d r y b i o m a s s ( C r a i g a n d M i l l e r , 1 9 6 6 ) . T h e c a l c u l a t e d a m o u n t o f d r y b i o -

m a s s w a s o n l y 1 9 9 , 5 0 0 k g / h a b u t c o n t a i n e d 5 8 6 k g o f N, 75 k g o f P, 2 0 7 k g

o f K, 2 3 0 k g o f C a a n d 9 3 k g o f Mg p e r h e c t a r e .

* L a t i n n a m e s o f t r e e s p e c i e s a r e p r e s e n t e d i n A p p e n d i x 1.

7

T h e n u t r i e n t a c c u m u l a t i o n w i t h i n a l o b l o l l y p i n e p l a n t a t i o n i n

s o u t h - e a s t e r n U.S.A. w a s s t u d i e d b y S w i t z e r a n d N e l s o n ( 1 9 7 2 ) . A 2 0 - y e a r

o l d p l a n t a t i o n w a s f o u n d t o c o n t a i n 1 7 4 k g o f N, 1 9 k g o f P, 9 8 k g o f K,

9 0 k g o f C a a n d 23 k g o f Mg p e r h e c t a r e .

I n s t u d y i n g a 9 3 - y e a r o l d s p r u c e f o r e s t i n R u s s i a , S m i r n o v a ( 1 9 5 1 )

f o u n d 4 6 8 k g o f N, 85 k g o f P, 2 7 8 k g o f K, 5 5 2 k g o f C a a n d 9 0 k g o f Mg

p e r h e c t a r e i n a n u n d e f i n e d q u a n t i t y o f b i o m a s s .

T h e d i s t r i b u t i o n o f n u t r i e n t s w i t h i n a 3 6 - y e a r o l d s e c o n d * - g r o w t h

D o u g l a s - f i r e c o s y s t e m w a s s t u d i e d b y C o l e e t a l , ( 1 9 6 7 ) . T h e d r y w e i g h t

o f t r e e o r g a n i c m a t t e r ( i n c l u d i n g r o o t s ) w a s 2 0 4 , 5 2 9 k g / h a c o n t a i n i n g 3 2 0

k g o f N, 66 k g o f P, 2 2 0 k g o f K, 3 3 3 k g o f C a p e r h e c t a r e . T h e a n n u a l u p -

t a k e o f t h e f o r e s t s t a n d w a s : 3 8 . 8 , 7 , 2 , 2 9 . 4 , 2 4 . 4 k g p e r h e c t a r e f o r

n i t r o g e n , p h o s p h o r o u s , p o t a s s i u m a n d c a l c i u m , r e s p e c t i v e l y .

M o r r i s o n ( 1 9 7 2 ) m e a s u r e d t h e d i s t r i b u t i o n o f b i o m a s s a n d m a c r o -

n u t r i e n t s o f t w e l v e j a c k p i n e s t a n d s i n N o r t h e r n O n t a r i o . T h e a v e r a g e

a m o u n t o f d r y a b o v e - g r o u n d o r g a n i c m a t t e r w a s 1 0 5 , 7 8 3 k g / h a . ( S t a n d s 6 5

y e a r s o l d , s i t e c l a s s I , h i g h s t o c k i n g d e n s i t y . ) T h e a m o u n t o f n u t r i e n t s

i m m o b i l i z e d w i t h i n t h e a b o v e - g r o u n d t r e e b i o m a s s w a s 1 6 1 . 1 , 1 2 . 5 , 8 1 . 0 ,

1 1 5 . 6 a n d 1 7 . 5 k g p e r h e c t a r e o f n i t r o g e n , p h o s p h o r o u s , p o t a s s i u m , c a l c i u m

a n d m a g n e s i u m , r e s p e c t i v e l y .

L j a m e b o r s a j ( 1 9 6 8 ) s t u d i e d i m m o b i l i z a t i o n o f n i t r o g e n a n d a s h

e l e m e n t s i n a n u m b e r o f t r e e s p e c i e s . I n v e s t i g a t i o n s o f t h e b i o m a s s w e r e

8

made o n 3 0 0 s a m p l e t r e e s o n 35 p l o t s i n s t a n d s o f 7 s p e c i e s a t t h e a g e o f

m aximum c u r r e n t a n n u a l i n c r e m e n t . F r o m t h e d a t a o b t a i n e d i t w a s c a l c u l a t e d

t h a t S c o t s p i n e r e q u i r e d 1.7 k g o f N , 0.6 k g o f P, 1.1 k g o f K a n d 1.9 k g

o f C a t o p r o d u c e o n e c u b i c m e t e r o f m e r c h a n t a b l e w o o d ; c o r r e s p o n d i n g f i g -

u r e s f o r N o r w a y s p r u c e w e r e 2 . 1 , 1 . 1 , 1.8 a n d 2.5 k g ; f o r b i r c h 3 . 8 , 1 . 0 ,

1.5 a n d 2.8 k g ; f o r o a k 6.7, 1.4, 4.4 a n d 1 3 . 4 k g ; f o r a s p e n 3 . 2 , 0 . 8 , 1.8

a n d 5.3 k g ; f o r l i m e 5 . 9 , 0 . 8 , 3.4 a n d 7.2 k g ; a n d f o r l a r c h 1 . 3 , 1 . 1 , 0.8

a n d 8.1 k g ; r e s p e c t i v e l y .

T h e a m o u n t o f n u t r i e n t s i m m o b i l i z e d w i t h i n t h e t r e e s t a n d d e p e n d s

m a i n l y o n t h e w e i g h t o f d r y t r e e b i o m a s s a n d s e c o n d l y o n t h e c o n c e n t r a t i o n

o f n u t r i e n t s i n p a r t i c u l a r t r e e s p e c i e s . T h e w e i g h t o f i m m o b i l i z e d e l e m e n t s

a c c o r d i n g t o t h e a b o v e - p r e s e n t e d l i t e r a t u r e c a n b e a s h i g h a s 5 8 0 k g N, 85

k g P, 2 8 0 k g K, 5 5 0 k g C a a n d 9 0 k g Mg p e r h e c t a r e . T h e a n n u a l u p t a k e o f

n u t r i e n t s d u r i n g t h e e a r l y s t a g e o f s t a n d g r o w t h ( f i r s t 3 0 - 4 0 y e a r s ) c a n b e

a s h i g h a s 4 0 k g N , 8 k g P, 3 0 k g K a n d 25 k g C a p e r h e c t a r e ( C o l e e t a l . ,

1 9 6 7 ; W i l l 1 9 6 4 ) .

2 , 2 : N u t r i e n t d i s t r i b u t i o n i n a b o v e - g r o u n d t r e e b i o m a s s .

T h e d i s t r i b u t i o n o f n u t r i e n t s among t h e c o m p o n e n t o r g a n s o f t h e

m a i n g r o w i n g c r o p ( T a b l e 1 ) i s s h o w n i n T a b l e 3.

T h e p e r c e n t a g e d i s t r i b u t i o n s o f n u t r i e n t s among t h e f i v e c o m p o n -

e n t s f o r e a c h s p e c i e s g r o u p r e p r e s e n t m e a n s o f t h e p e r c e n t a g e d i s t r i b u t i o n s

o f t h e s e v e r a l s p e c i e s w i t h i n t h a t g r o u p . T h e s e d a t a a r e i n a g r e e m e n t w i t h

9

T A B L E 3: T h e p e r c e n t a g e d i s t r i b u t i o n o f n u t r i e n t s i m m o b i l i z e d i n t h e c o m -p o n e n t o r g a n s o f t h e m a i n g r o w i n g c r o p o f t h r e e t y p e s o f s t a n d a t a g e 1 0 0 y e a r s ( R e n n i e , 1 9 5 5 ) .

M a i n M e a n s o f t h e p e r c e n t a g e d i s t r i b u t i o n o f n u t r i -G r o w i n g e n t s a mong t h e m a i n c r o p c o m p o n e n t s

S p e c i e s G r o u p C r o p C o m p o n e n t A f t e r 5 0 y e a r s ' g r o w t h A f t e r 1 0 0 y e a r s ' g r o w t h

C a K P C a K P

n e e d l e s 2 3 3 9 4 3 14 2 8 32 b r u s h 25 1 8 2 1 1 8 14 1 8

P i n e s s t e m b a r k 17 8 9 2 5 1 2 14 s t e m w o o d 22 22 1 4 3 0 3 3 2 3 r o o t s 1 3 1 3 1 3 1 3 1 3 13

n e e d l e s 26 3 0 4 0 1 5 17 27 b r u s h 3 0 27 25 2 1 2 0 2 2

O t h e r s t e m b a r k 2 0 14 1 3 3 2 23 2 3 c o n i f e r s s t e m w o o d 1 1 16 9 1 9 27 1 5

r o o t s 1 3 1 3 1 3 13 1 3 13

l e a v e s 1 1 2 0 22 5 1 0 1 2 b r u s h 22 2 5 28 16 2 1 24

H a r d w o o d s * s t e m b a r k 3 8 14 17 46 18 2 3 s t e m w o o d 16 2 8 2 0 2 0 3 8 28 r o o t s 1 3 1 3 1 3 1 3 13 1 3

* e x c l u d i n g t h o s e w i t h o u t l e a v e s s a m p l e d i n w i n t e r .

d a t a r e v i e w e d b y O v i n g t o n ( 1 9 6 2 ) f o r a l a r g e n u m b e r o f e c o s y s t e m s i n d i f -

f e r e n t p a r t s o f t h e w o r l d . D u v i g n e a u d a n d D e n a e y e r - D e S m e t ( 1 9 6 4 ) a n a l y s e d

t h e v a r i o u s o r g a n s o f t h r e e d e c i d u o u s s p e c i e s a n d f o u n d t h e f o l l o w i n g s e -

q u e n c e w h e n r e s u l t s a r e e x p r e s s e d a s t h e c o n t e n t o f e a c h n u t r i e n t i n p e r -

c e n t a g e o f t h e t o t a l t r e e c o n t e n t o f t h a t e l e m e n t :

1 0

N - f o l i a g e > b o l e w o o d > b o l e b a r k = b r a n c h b a r k > -

b r a n c h w o o d t w i g s

C a - b o l e b a r k > b r a n c h b a r k > f o l i a g e > b o l e w o o d

t w i g s > b r a n c h w o o d

K - f o l i a g e > b o l e w o o d > b r a n c h w o o d ; > t w i g s >

b o l e b a r k = b r a n c h b a r k

S l i g h t c h a n g e s i n t h i s s e q u e n c e w e r e n o t e d a mong some s p e c i e s d u e t o i n -

h e r e n t m o r p h o l o g i c a l d i f f e r e n c e s . F o r e x a m p l e , r e p r o d u c t i v e o r g a n s a r e

n o t a b l y r i c h i n some e l e m e n t s . D u v i g n e a u d a n d D e n a e y e r - D e S m e t s u m m a r i z e d

t h e l i t e r a t u r e o n m i n e r a l c o n t e n t o f p o l l e n a n d e s t a b l i s h e d t h a t t h e e l e -

m e n t s a r e p r e s e n t i n t h e o r d e r o f N > K J > P > - C a o n a p e r c e n t a g e b a s i s .

T h e c o n c e n t r a t i o n s o f N, P, K, C a , Mg a n d S i n n e e d l e s o f P a c i f i c

s i l v e r f i r , a l p i n e f i r , w e s t e r n r e d c e d a r , D o u g l a s - f i r , w e s t e r n h e m l o c k ,

l o d g e p o l e p i n e , E n g e l m a n n s p r u c e a n d S i t k a s p r u c e o f d i f f e r e n t a g e s a n d

l o c a t i o n s i n B r i t i s h C o l u m b i a w e r e s t u d i e d b y B e a t o n e_t a l . ( 1 9 6 5 ) . H i s

r e s u l t s f o r w e s t e r n h e m l o c k , a m a b i l i s f i r a n d r e d c e d a r a r e s u m m a r i z e d i n

T a b l e 4.

N u t r i e n t c o n c e n t r a t i o n s i n f o l i a g e o f D o u g l a s - f i r i n W a s h i n g t o n

S t a t e w e r e s t u d i e d b y G e s s e l eX a l . ( 1 9 6 0 ) f o r s t a n d s o f d i f f e r e n t g r o w t h

v i g o r . H i s r e s u l t s c a n b e s u m m a r i z e d : N - 0.6 t o 2.3 p e r c e n t ; P - 0.1

t o 0.25 p e r c e n t ; K - 0.3 t o 1.0 p e r c e n t : C a - 0.2 t o 0.75 p e r c e n t , a n d

1 1

T A B L E 4: C o n c e n t r a t i o n o f m a c r o n u t r i e n t s i n f o l i a g e o f t h e u p p e r w h o r l s o f t r e e c r o w n f o r t h r e e t r e e s p e c i e s i n B. C. ( B e a t o n e t •. ml.'» 1 9 6 5 ) .

T y p e , l o c a t i o n D a t e N o . o f Y e a r P l a n t n u t r i e n t c o n c e n t r a t i o n & a g e o f s t a n d o f T r e e s o f p e r c e n t i n d r y m a t t e r

S a m p l e S a m p l e d G r o w t h N P K C a Mg

A m a b i l i s f i r A u g u s t 5 1 9 6 3 0.99 0.16 1.20 0.30 0.08 T e r r a c e ( 8 y r . ) 1 9 6 3 1 9 6 2 0 . 9 1 0.12 1.10 0 . 4 1 0.07

W e s t e r n r e d A u g u s t c e d a r , T e r r a c e 1 9 6 3 5 1 9 6 3 0 .73 0.13 0.52 1.16 0.10 ( 5 8 y e a r s )

W e s t e r n h e m l o c k F e b . 1 9 6 3 1.17 0.16 0.43 0.20 0.08 Q u a d r a I s l a n d 1 9 6 4 1 2 1 9 6 2 1 . 2 1 0.18 0.40 0.24 0.10 ( 6 0 y e a r s ) 1 9 6 1 1.22 0.18 0.38 0.30 0.08

G o l d R i v e r A p r i l ( 1 4 0 - 1 5 0 y e a r s ) 1 9 6 4 5 1 9 6 3 0 . 9 1 0 . 1 1 0.28 0.18 0.13

T e r r a c e ( 8 , 5 7 , A u g u s t 1 9 6 3 0.86 0.19 0.57 0.27 0.12 7 3 , 1 1 5 , 1 5 3 , 1 9 6 3 2 0 1 9 6 2 0.84 0 . 2 1 0 . 5 1 0.32 0.14 1 8 5 y e a r s ) 1 9 6 1 0.88 0.22 0.47 0.35 0.14

Mg - 0.05 t o 0.15 p e r c e n t .

T h e i m p o r t a n c e o f h a v i n g a c o r r e c t b a l a n c e b e t w e e n n u t r i e n t s i n

t h e f o l i a g e a n d e s p e c i a l l y o f t h e r a t i o s N /P, N/K, a r i d K/P h a s b e e n s t r e s -

s e d b y a n u m b e r o f i n v e s t i g a t o r s . F o r e x a m p l e , L e y t o n ( 1 9 5 7 ) c a l c u l a t e d

N/P r a t i o s f o r J a p a n e s e l a r c h i n B r i t a i n a n d f o u n d t h a t t h e y g e n e r a l l y l i e

b e t w e e n 1 2 a n d 1 5 . He c i t e s o t h e r a u t h o r s a s f i n d i n g N/P r a t i o s o f b e t w e e n

9 t o 1 2 .

T h e t o t a l a m o u n t o f n i t r o g e n c o n t a i n e d i n a t r e e i n c r e a s e s w i t h

1 2

a g e a s a f u n c t i o n o f b i o m a s s a c c u m u l a t i o n a n d n i t r o g e n c o n c e n t r a t i o n w i t h i n

a t r e e . S w i t z e r a n d N e l s o n ( 1 9 6 4 ) g i v e a v e r a g e N c o n c e n t r a t i o n s i n v a r i o u s

l o b l o l l y p i n e t i s s u e s a t v a r i o u s a g e s o n g o o d s i t e s ( i n p e r c e n t o f o v e n -

d r y w e i g h t ) , ( T a b l e 5)..

T A B L E 5: A v e r a g e N - c o n c e n t r a t i o n s i n v a r i o u s l o b l o l l y p i n e t i s s u e s a t v a r i o u s a g e s o n g o o d s i t e s i n p e r c e n t o f o v e n d r y w e i g h t ( S w i t z e r a n d N e l s o n , 1 9 6 4 ) .

T r e e P L A N T T I S S U E

A g e C u r r e n t f o l i a g e O l d e r b r a n c h e s S t e m b a r k S t e m w o o d

4 1.00 0 . 4 1 0.42 ' 0.16

8 0.95 0.24 0.24 0.06

18 1,08 0.23 0.23 0.06

3 0 1.22 0.22 0.19 0.04

56 1.16 0 . 2 1 0.17 0.03

A v e r a g e c o n c e n t r a t i o n s o f m a c r o n u t r i e n t s i n t h e a b o v e - g r o u n d o r g a n s

o f N o r w a y s p r u c e s t a n d s i n C z e c h o s l o v a k i a w e r e c a l c u l a t e d b y M a t e r n a ( 1 9 6 4 ) ,

T h e r e s u l t s a r e s u m m a r i z e d i n T a b l e 6,

T h e d i s t r i b u t i o n o f m a c r o n u t r i e n t s b e t w e e n t h e m a j o r c o m p o n e n t s

o f t h e 3 6 - y e a r o l d s e c o n d - g r o w t h D o u g l a s - f i r f o r e s t w a s c a l c u l a t e d b y C o l e

e t a l , ( 1 9 6 7 ) . T h e r e s u l t s a r e s u m m a r i z e d i n T a b l e 7.

1 3

T A B L E 6: T h e r a n g e o f n u t r i e n t c o n c e n t r a t i o n s ( i n p e r c e n t ) i n t h e d i f -f e r e n t t r e e c o m p o n e n t s i n v a r i o u s N o r w a y s p r u c e s t a n d s i n C z e c h o s l o v a k i a ( s i t e c l a s s c l a s s i f i c a t i o n a f t e r m o d i f i e d S c h w a p p a c h t a b l e s f o r C z e c h o s l o v a k i a ) , ( M a t e r n a , 1 9 6 4 ) .

S i t e C l a s s N P K C a Mg S

WOOD

1 0 , 1 3 3 - 0 , 0 1 5 - 0 , 0 3 0 - 0 , 0 6 6 - t r a c e 0 , 0 1 3 -0,217 0 , 0 3 0 0 , 0 5 1 0 , 0 8 6 0 , 0 3 5

5 0 , 1 4 7 - 0 , 0 2 4 - 0 , 0 3 4 - 0 , 0 4 4 - 0 , 0 0 7 - 0 , 0 1 3 -0 , 1 8 9 0 , 0 2 6 0 , 0 5 1 0 , 0 9 0 0 , 0 1 0 0 , 0 3 1

9 0 , 1 4 0 - 0 , 0 2 6 - 0 , 0 1 2 - 0 , 0 3 2 - 0 , 0 1 0 - 0 , 0 1 1 -0 , 2 1 0 0 , 0 4 8 0 , 0 4 2 0 , 0 6 6 0 , 0 1 8 0 , 0 2 9

BARK

1 0 , 3 6 7 - 0 , 0 3 8 - 0 , 2 0 2 - 0 , 5 0 4 - 0 , 0 1 4 - 0 , 0 4 3 -0 , 5 9 5 0 , 0 7 6 0 , 4 0 4 1 , 0 6 4 0 , 0 3 9 0 , 0 6 1

5 0 , 2 4 5 - 0 , 0 4 0 - 0 , 1 1 7 - 0 , 4 5 0 - 0 , 0 0 8 - 0 , 0 2 7 -0 , 4 8 3 0,087 0,216 1 , 0 2 0 0 , 0 2 9 0 , 0 6 0

9 0 , 2 9 4 - 0 , 0 3 3 - 0 , 0 5 2 - 0 , 4 9 4 - 0 , 0 3 5 - 0 , 0 1 5 -0 , 5 6 0 0 , 0 6 1 0,247 1 , 0 6 8 0 , 0 8 8 0 , 0 4 5

F O L I A G E

1 1 , 3 0 - 2 , 1 0 0 , 1 3 - 0 , 2 7 0 , 4 8 - 0 , 9 8 0 , 2 1 - 0 , 5 8 0 , 0 4 - 0 , 1 1 0 , 1 1 - 0 , 1 4

5 1 , 0 5 - 1 , 4 1 0 , 1 1 - 0 , 1 8 0 , 1 6 - 0 , 6 6 0 , 1 7 - 0 , 4 6 0 , 0 4 - 0 , 1 9 0 , 1 0 - 0 , 1 5

9 0 , 8 3 - 1 , 3 1 0 , 1 0 - 0 , 1 6 0 , 1 1 - 0 , 7 9 0 , 0 8 - 0 , 4 2 0 , 0 4 - 0 , 0 9 0 , 1 0 - 0 , 1 5

TWIGS

0 , 2 3 - 0 , 8 6 0 , 0 3 - 0 , 1 4 0 , 1 1 - 0 , 6 4 0 , 2 9 - 0 , 4 1 0 , 0 4 - 0 , 0 8 0 , 0 5 - 0 , 0 9

1 4

T A B L E 7: D i s t r i b u t i o n o f N, P, K a n d C a among t h e m a j o r c o m p o n e n t s o f a 3 6 - y e a r - o l d s e c o n d - g r o w t h D o u g l a s - f i r f o r e s t ( C o l e e t a l . , 1 9 6 7 )

E c o s y s t e m N P K C a C o m p o n e n t k g / h a % o f k g / h a % o f k g / h a % o f k g / h a % o f

t o t a l t o t a l t o t a l t o t a l

F o l i a g e 1 0 2 3 1 . 9 29 4 3 . 9 6 2 2 8 . 2 73 2 1 . 9

B r a n c h e s 6 1 1 9 . 1 1 2 1 8 . 2 3 8 1 7 . 3 1 0 6 3 1 . 8

Wood 77 2 4 . 0 9 1 3 . 6 5 2 2 3 . 6 47 1 4 . 1

B a r k 4 8 1 5 . 0 1 0 1 5 . 2 4 4 2 0 . 0 70 2 1 . 0

R o o t s 3 2 1 0 . 0 6 9.1 24 1 0 . 9 37 1 1 . 2

T o t a l 3 2 0 66 2 2 0 3 3 3

A s i m i l a r s t u d y w a s u n d e r t a k e n b y W i l l ( 1 9 6 4 ) i n a 1 2 - y e a r o l d

s t a n d o f r a d i a t a p i n e o n t h e N o r t h I s l a n d o f New Z e a l a n d ( T a b l e 8 ) .

T h e p e r c e n t a g e d i s t r i b u t i o n o f m a c r o n u t r i e n t s i n t h e a e r i a l c o m -

p o n e n t s o f j a c k p i n e w a s c a l c u l a t e d b y M o r r i s o n ( 1 9 7 3 ) f o r t w e l v e n a t u r a l

s t a n d s i n N o r t h e r n O n t a r i o . T a b l e 9 s u m m a r i z e s h i s r e s u l t s f o r t h r e e a g e

c l a s s e s .

A s a g e n e r a l r u l e , o n e c a n s a y t h a t t h e c o n c e n t r a t i o n o f n u t r i e n t s

i s t h e h i g h e s t i n r e p r o d u c t i v e o r g a n s a n d s e e d s b e i n g f o l l o w e d b y f o l i a g e ,

t w i g s , b a r k o r b r a n c h e s a n d t r e e b o l e , s a p w o o d a n d h e a r t w o o d i n t h a t

o r d e r . T h e n u t r i e n t c o n c e n t r a t i o n o f a p a r t i c u l a r t r e e c o m p o n e n t i s v e r y

T A B L E 8: D i s t r i b u t i o n o f b i o m a s s a n d n u t r i e n t s w i t h i n a b o v e - g r o u n d t r e e c o m p o n e n t s o f 1 2 - y e a r - o l d s t a n d o f r a d i a t a p i n e , W h a k a r e w a r e w a F o r e s t ( W i l l , 1 9 6 4 ) .

D r y N P K C a Mg_ M a t t e r % l b / % l b / % l b / % l b / % l b / l b / a c D.M.* a c D.M. a c D.M. a c D.M. a c D.M. a c

S t a n d i n g t r e e s

F o l i a g e 8 , 0 0 0 1.47 1 1 8 0.15 1 2 . 0 0.99 79 0.33 26 0.12 1 0 T w i g s 2 , 0 0 0 0.90 1 8 0.10 2.0 0.79 16 0 . 2 1 4 0.13 3 D e a d B r . 6,000 0.28 17 0 . 0 1 0.6 0.08 5 0.18 1 1 0.08 5 L i v e B r . 7 , 0 0 0 0.38 27 0.03 2.1 0.27 19 0.19 13 0.07 5 D e a d S t . 7 , 3 0 0 0.12 9 0 . 0 1 0.7 0.16 12 0 . 0 8 5 6 0 . 0 3 5 3 S m a l l L i v e

S t e m s 9 , 3 0 0 0.12 1 1 0 . 0 1 0.9 0.13 1 2 0.09 8 0 . 0 3 5 3 L a r g e L i v e

S t e m s 1 0 9 , 2 0 0 0.10 1 0 9 0 . 0 0 9 9.8 0.11 1 2 0 0.08 89 0 . 0 3 5 37

T o t a l 1 4 8 , 0 0 0 3 0 9 2 8 . 1 2 6 3 1 5 7 6 6

* D.M. - d r y m a t t e r .

16

T A B L E 9: A v e r a g e p e r c e n t a g e d i s t r i b u t i o n o f N, P, K, C a a n d Mg i n a e r i a l c o m p o n e n t s o f 12 n a t u r a l j a c k p i n e s t a n d s i n N o r t h e r n O n t a r i o ( M o r r i s o n , 1 9 7 3 ) .

A g e L i v e D e a d S t e m S t e m L i v e C l a s s F o l i a g e B r a n c h e s B r a n c h e s B a r k Wood C o n e s C r o w n

N I T R O G E N

20 6 1 . 8 a 1 9 . 9 a 1.8 a 5.2 a 1 0 . 9 a 0.4 a 8 1 . 7 a 3 0 3 6 . 5 b 1 9 . 4 a 1 1 . 8 b 9.5* b 2 2 . 4 b 0.4 a 5 5 . 9 b 65 2 8 . 0 c 2 1 . 0 a 2.7 a 9.3 b 3 6 . 3 c 2.7 b 4 9 . 0 b

PHOSPHOROUS

20 6 6 . 6 a 1 5 . 0 a 0.6 a 5.4 a 1 1 . 9 a 0.5 a 8 1 . 6 a 3 0 4 2 . 8 b 1 7 . 8 a 4.1 b 9.9 a 2 4 . 7 b 0.7 a 6 0 . 6 b 65 4 2 . 6 b 1 9 . 0 a 1.6 a b 8.8 a 2 4 . 2 b 3.8 b 6 1 . 6 b

P O T A S S I U M

20 4 2 . 0 a 3 1 . 6 a 0.7 a 6.7 a 1 8 . 6 a 0.4 a 7 3 . 6 a 3 0 2 3 . 3 b 2 6 . 8 b 3.9 b 1 0 . 0 a 3 5 . 7 b 0.3 a 5 0 . 1 b 6 5 1 5 . 8 c 2 5 . 4 b 1.3 a 8.2 a 4 7 . 9 c 1.4 b 4 1 . 2 b

C A L C I U M

20 3 7 . 3 a 3 3 . 7 a 2.2 a 1 1 . 0 a 1 5 . 7 a 0.1 a 7 1 . 0 a 30 1 6 . 1 b 2 4 . 2 b 7.8 b 2 1 . 1 b 3 0 . 7 b 0.1 a 4 0 . 3 b 6 5 9.4 c 1 9 . 4 b 2.3 a 2 3 . 3 b 4 5 . 2 c 0.4 b 2 8 . 8 c

MAGNESIUM

20 4 0 . 6 a 2 9 . 4 a 1.0 a 1 0 . 5 a 1 8 . 2 a 0.3 a 7 0 . 0 a 3 0 2 0 . 4 b 2 3 . 9 a b 3.0 a 1 3 . 3 a 3 9 . 2 b 0.2 a 4 4 . 3 b 6 5 1 5 . 6 b 2 0 . 0 b 2.2 a 9.0 a 5 1 . 1 c 2.1 b 3 5 . 6 b

N o t e : C o r r e s p o n d i n g s u p e r s c r i p t l e t t e r s i n v e r t i c a l c o l u m n s i n d i c a t e n o s i g n i f i c a n t ( 5 % ) d i f f e r e n c e s b e t w e e n v a l u e s , a s c a l c u l a t e d b y a t - t e s t .

17

v a r i a b l e a n d d e p e n d s o n t h e e l e m e n t , s p e c i e s , a g e a n d n u t r i e n t a v a i l a b i l i t y

i n a p a r t i c u l a r e c o s y s t e m . D i s t r i b u t i o n o f n u t r i e n t s b e t w e e n t h e v a r i o u s

a b o v e - g r o u n d t r e e c o m p o n e n t s o f a s t a n d v a r i e s f o r d i f f e r e n t n u t r i e n t e l e -

m e n t s , t r e e s p e c i e s a n d t r e e a g e s . M o s t o f t e n t h e p r o p o r t i o n s o f n u t r i e n t s

o f y o u n g s t a n d s a p p e a r i n t h e s e q u e n c e : f o l i a g e a n d t w i g s > b r a n c h e s >

s t e m b a r k > s t e m w o o d . T h e p r o p o r t i o n s o f n u t r i e n t s o f o l d s t a n d s a r e r e -

v e r s e d : s t e m w o o d > s t e m b a r k > f o l i a g e a n d t w i g s >* b r a n c h e s , T h i s i s o n l y

a g e n e r a l r u l e , h o w e v e r , a n d t h e a c t u a l d i s t r i b u t i o n o f p a r t i c u l a r n u t r i -

t i o n a l e l e m e n t s w i t h i n a p a r t i c u l a r t r e e s t a n d may v a r y c o n s i d e r a b l y f r o m

t h i s b a s i c p a t t e r n .

2 . 3 : R e m o v a l o f n u t r i e n t s f r o m a s i t e a s a r e s u l t o f t r e e h a r v e s t i n g .

T h e a m o u n t o f n u t r i e n t s r e m o v e d f r o m a n e c o s y s t e m d u e t o l o g g i n g

I s e q u a l t o t h e a m o u n t o f n u t r i e n t s i m m o b i l i z e d w i t h i n t h o s e a b o v e - g r o u n d

c o m p o n e n t s t h a t a r e p h y s i c a l l y r e m o v e d f r o m t h e s i t e p l u s t h e a m o u n t o f

n u t r i e n t s l o s t t h r o u g h t h e i n c r e a s e d s o i l l e a c h i n g w h i c h may a c c o m p a n y t h e

i n c r e a s e i n t h e r a t e o f o r g a n i c m a t t e r d e c o m p o s i t i o n a n d t h e d i s r u p t i o n o f

t h e n u t r i e n t c y c l e . I n m o u n t a i n t e r r a i n c o n s i d e r a b l e a m o u n t s o f n u t r i e n t s

may a l s o b e l o s t a s t h e r e s u l t o f s o i l e r o s i o n ( C u r r y , 1 9 7 3 ) . T h e r e i s a

c o n t i n u o u s i n p u t o f n u t r i e n t s t o a n e c o s y s t e m , h o w e v e r , i n t h e f o r m o f

p r e c i p i t a t i o n , d u s t f a l l a n d b e d r o c k w e a t h e r i n g . T h e c a p i t a l o f a v a i l a b l e

n u t r i e n t s r e s u l t s f r o m t h e s e i n p u t s a n d o u t p u t s a n d f r o m e x c h a n g e s b e t w e e n

t h e a v a i l a b l e a n d u n a v a i l a b l e p o o l s i n t h e s o i l . T h e s i g n i f i c a n c e o f a n y

1 8

n u t r i e n t r e m o v a l i n h a r v e s t e d m a t e r i a l s m u s t b e v i e w e d t o g e t h e r w i t h o t h e r

l o s s e s a n d t h e v a r i o u s i n p u t s b y w h i c h t h e l o s s e s may b e r e p l a c e d .

T h e f o l l o w i n g d i s c u s s i o n w i l l b e l i m i t e d t o o u t p u t s i n l o g g e d

m a t e r i a l s h o w e v e r , s i n c e t h i s p a r a m e t e r w a s , m e a s u r e d i n t h e r e s e a r c h f o r

t h i s t h e s i s . T h e a m o u n t o f n u t r i e n t s r e m o v e d f r o m a n e c o s y s t e m i n t r e e

b i o m a s s i s d e t e r m i n e d b y t h e l e v e l o f t r e e u t i l i z a t i o n . D i s t r i b u t i o n o f

n u t r i e n t s b e t w e e n d i f f e r e n t c o m p o n e n t s o f t h e t r e e b i o m a s s h a s a l r e a d y b e e n

d i s c u s s e d i n S e c t i o n 2 . 2 . F r o m t h i s r e v i e w i t i s a p p a r e n t t h a t f o r t h e

m a j o r i t y o f t r e e s p e c i e s , a p p r o x i m a t e l y 5 0 p e r c e n t o r e v e n m o r e o f t h e

t o t a l n u t r i e n t s i m m o b i l i z e d i n t h e a b o v e - g r o u n d t r e e b i o m a s s i s r e t a i n e d

w i t h i n c o m p o n e n t s o f t h e t r e e c r o w n . T h e e x a c t p r o p o r t i o n v a r i e s f o r d i f -

f e r e n t t r e e s p e c i e s a n d d i f f e r e n t n u t r i e n t e l e m e n t s , b u t i t i s g e n e r a l l y a

s u b s t a n t i a l p r o p o r t i o n o f t h e t o t a l . T r e e b a r k i s a n o t h e r n u t r i t i o n a l l y

r i c h c o m p o n e n t o f t r e e b i o m a s s . A s a g e n e r a l r u l e , o n e c a n s a y t h a t a p p r o x -

i m a t e l y h a l f o f t h e n u t r i e n t s i n t h e a b o v e - g r o u n d t r e e b i o m a s s c a n b e r e -

t a i n e d o n t h e s i t e a n d b e a v a i l a b l e f o r t h e g r o w t h o f t h e n e x t t r e e r o t a t i o n

i f o n l y t h e t r e e b o l e s a r e h a r v e s t e d . Up t o 6 0 - 8 0 p e r c e n t o f t r e e n u t r i -

e n t s c a n b e r e t a i n e d i f t h e l o g s a r e d e - b a r k e d a t s t u m p a n d o n l y t h e t r e e

b o l e w o o d i s h a r v e s t e d . W e e t m a n a n d W e b e r c a l c u l a t e d t h a t h a r v e s t i n g o f

t r e e b o l e s o n l y , a s c o m p a r e d w i t h w h o l e t r e e l o g g i n g , c a n r e d u c e r e m o v a l

o f m a c r o n u t r i e n t s f r o m a n e c o s y s t e m b y a s m u c h a s 75 p e r c e n t o f N a n d P,

a n d 6 0 p e r c e n t o f K, C a a n d Mg.

19

I I I D E S C R I P T I O N OF SAMPLE P L O T S

L o c a t i o n a n d s i z e :

T h e s t u d y w a s c o n d u c t e d a t t w o s i t e s . T h e f i r s t w a s i n i t i a l l y

s e l e c t e d a s a p i l o t s t u d y f o r t h e s e c o n d s i t e , b u t , o w i n g t o t h e e x c e p t i o n -

a l l y l a t e s n o w m e l t w h i c h r e s t r i c t e d a c c e s s t o t h e h i g h e r e l e v a t i o n s i t e

u n t i l m i d - s u m m e r , a r e a s o n a b l y c o m p l e t e s t u d y w a s a c c o m p l i s h e d o n b o t h

s i t e s . T h e r e w a s n o i n t e n t i o n t o m a k e a c o m p a r a t i v e s t u d y , h o w e v e r , a n d

t h e s a m p l e p l o t s w e r e n o t s e l e c t e d t o b e c o m p a r a b l e .

T h e f i r s t p l o t w a s l o c a t e d o n a n o r t h - f a c i n g a s p e c t o n a r o c k y

r i d g e a t 7 2 0 m ( 2 , 2 0 0 f t . ) e l e v a t i o n i n t h e n o r t h - e a s t e r n c o r n e r o f t h e

U n i v e r s i t y o f B r i t i s h C o l u m b i a R e s e a r c h F o r e s t a t H a n e y , B. C. I t l a y a p -

p r o x i m a t e l y h a l f a m i l e e a s t o f t h e n o r t h e r n e n d o f C a t h e r i n e L a k e . T h e

p l o t w a s 0.79 h a ( a b o u t 2 a c r e s ) i n s i z e , e l o n g a t e i n s h a p e , a n d i n c l u d e d

1 1 4 t r e e s .

T h e s e c o n d p l o t w a s l o c a t e d o n a s o u t h - f a c i n g a s p e c t a t 1 , 5 0 0 m

( 4 , 5 0 0 f t . ) e l e v a t i o n o n t h e n o r t h b a n k o f t h e Mamquam R i v e r b e t w e e n

S k o o k u m C r e e k a n d C r a w f o r d C r e e k . T h e p l o t w a s l o c a t e d o n a g e n t l y ( 2 5 p e r

c e n t ) s l o p i n g b e n c h , w i t h s t e e p e r s l o p e s i m m e d i a t e l y b e l o w a n d s t e e p , b r o -

k e n , r o c k y t e r r a i n a b o v e . T h e p l o t w a s 1.16 h a ( a b o u t 3 a c r e s ) i n s i z e ,

t r i a n g u l a r i n s h a p e , a n d i n c l u d e d 2 5 6 t r e e s .

T h e l o c a t i o n o f t h e t w o p l o t s may b e s e e n i n m o r e d e t a i l i n „

F i g u r e 1.

FIGURE 1: Location of the sample plots.

l i t '

21

V i e w o f t h e l o g g e d a r e a n e a r t h e S q u a m i s h s a m p l e p l o t . A r r o w i n d i c a t e s l o c a t i o n o f t h e p l o t .

2 3

l

24

A c c o r d i n g t o H a l l i d a y ' s c l a s s i f i c a t i o n ( H a l l i d a y , 1 9 3 7 ) b o t h

p l o t s a r e l o c a t e d i n t h e S o u t h e r n C o a s t S e c t i o n . R o w e ' s ( 1 9 5 9 ) c l a s s i f i -

c a t i o n , w h i c h i s b a s i c a l l y a n e l a b o r a t i o n u p o n t h a t o f H a l l i d a y , p l a c e s

t h e S q u a m i s h p l o t i n t h e C o a s t a l S u b a l p i n e S e c t i o n a n d t h e H a n e y p l o t i n

t h e S o u t h e r n P a c i f i c C o a s t S e c t i o n . A c c o r d i n g t o K r a j i n a ' s c l a s s i f i c a t i o n

( 1 9 6 9 ) , t h e S q u a m i s h p l o t i s i n t h e s u b a l p i n e f o r e s t s u b z o n e o f t h e M o u n -

t a i n H e m l o c k Z o n e w h i l e t h e H a n e y p l o t i s i n t h e u p p e r e l e v a t i o n o f t h e

w e t s u b z o n e o f t h e C o a s t a l W e s t e r n H e m l o c k Z o n e . A c c o r d i n g t o t h e

F o r e s t P r o d u c t i v i t y Z o n e s o f t h e B C F S , t h e S q u a m i s h p l o t w o u l d b e d e s c r i b e d

a s i n t e r m e d i a t e b e t w e e n C o a s t R a n g e S q u a m i s h - F r a s e r a n d S u b a l p i n e - A l p i n e

T u n d r a - G l a c i e r s , w h i l e t h e H a n e y p l o t w o u l d b e C o a s t R a n g e S q u a m i s h -

F r a s e r .

C l i m a t e :

No c l i m a t i c d a t a w e r e c o l l e c t e d o n t h e s a m p l e p l o t s . T a b l e 1 0

s u m m a r i z e s c l i m a t i c d a t a r e p r e s e n t a t i v e o f t h e a r e a o f b o t h s a m p l e p l o t s

a c c o r d i n g t o K r a j i n a ( 1 9 6 5 ) , C l i m a t e i n t h e f o r e s t s u b z o n e o f t h e M o u n -

t a i n H e m l o c k Z o n e i s r a t h e r u n f a v o u r a b l e t o t r e e g r o w t h . T h e n u m b e r o f

f r o s t - f r e e d a y s r a n g e s f r o m 4 0 t o 1 2 0 p e r annum, w h i l e t h e mean d a i l y t e m p -

e r a t u r e i s a b o v e 50 ° F f o r o n l y 2 o r 3 m o n t h s i n d i c a t i n g a v e r y s h o r t

g r o w i n g s e a s o n . Snow p e r s i s t s f o r 8 t o 9 m o n t h s e v e r y y e a r , r e t r e a t i n g i n

t h e m i d d l e o f J u l y a n d r e t u r n i n g i n t h e m i d d l e o f O c t o b e r . A n n u a l t o t a l

p r e c i p i t a t i o n i s v e r y h i g h , r a n g i n g f r o m 2 , 2 0 0 t o 4 , 3 0 0 mm m a i n l y i n t h e

T A B L E 1 0 :

P l o t

S q u a m i s h

H a n e y

Summary o f c l i m a t i c i n f o r m a t i o n ( K r a j i n a , 1 9 6 5 ) . D a t a r e p r e s e n t c h a r a c t e r i s t i c r a n g e s f o r t h e M o u n t a i n H e m l o c k a n d t h e W e s t e r n H e m l o c k ( w e t s u b z o n e ) B i o g e o c l i m a t l c z o n e s . No c l i m a t i c d a t a w e r e o b t a i n e d o n t h e p l o t s .

M e a n A n n u a l

3 8 - 4 4

4 1 - 4 9

M e a n M o n t h l y J A N J U L

1 6 - 3 2 5 0 - 5 5

2 4 - 4 1 5 5 - 6 4

TEMPERATURE (°F)

N u m b e r o f M o n t h s A n n u a l A b o v e B e l o w R a n g e 50°F 32°F

2-3 (-4) 1-6 3 0 - 4 0

4-6 0-3 (-4) 1 6 - 3 7

N u m b e r o f F r o s t - f r e e

D a y s

4 0 - 1 2 0

( 1 2 0 - ) 1 5 0 - 2 5 0

A b s o l u t e

M a x . M i n .

7 5 - 8 6 - 3 1 - 1 0

7 8 - 1 0 4 - 2 2 - 1 9

P R E C I P I T A T I O N

P l o t

S q u a m i s h

H a n e y

A n n u a l T o t a l (mm)

2 2 0 0 -4 3 0 0

1 7 8 0 -6 6 5 0

A n n u a l S n o w f a l l

(mm)

2 5 8 0 -2 0 0 0 0

1 2 6 -7 5 0 0

D r i e s t M o n t h (mm)

3 3 - 8 4

3 2 - 1 6 5

W e t t e s t M o n t h (mm)

3 0 0 -4 0 0

2 8 0 -1 1 6 0

C l i m a t e C l a s s i f i c a t i o n ( K o p p e n )

D f b - D f c

C f b ( a n d t h e m i l d e s t D f b )

D a t a o r i g i n a l l y e x p r e s s e d i n i n c h e s .

26

f o r m o f s n o w . Summer i s u s u a l l y v e r y s h o r t (.6-8 w e e k s ) a n d i s h o t a n d d r y .

I n t h e w e t s u b z o n e o f t h e W e s t e r n H e m l o c k Z o n e t h e c l i m a t e i s

m o r e f a v o u r a b l e . T h e n u m b e r o f f r o s t - f r e e d a y s r a n g e s f r o m 1 5 0 t o 2 5 0

d a y s p e r a n n u m a n d t h e m e an d a i l y t e m p e r a t u r e i s a b o v e 5 0 ° F f o r 4 t o 6

m o n t h s , p r o v i d i n g a r e a s o n a b l y l o n g p e r i o d f o r g r o w t h . Snow c o v e r r e m a i n s

f o r 3 t o 4 m o n t h s e v e r y y e a r . A n n u a l t o t a l p r e c i p i t a t i o n r a n g e s b e t w e e n

1 , 7 0 0 - 2 , 2 0 0 mm, m a i n l y i n t h e f o r m o f r a i n .

S o i l s ;

B e c a u s e o f t i m e a n d f i n a n c i a l c o n s t r a i n t s s o i l a n a l y s e s w e r e n o t

c o n d u c t e d a s p a r t o f t h i s t h e s i s s t u d y . S o i l c a t e g o r i e s f o r t h e S u b a l p i n e

M o u n t a i n H e m l o c k Z o n e i n g e n e r a l h a v e b e e n d e s c r i b e d b y B r o o k e e t a l . ( 1 9 7 0 ) .

S o i l s o n t h e S q u a m i s h a n d H a n e y s a m p l e p l o t s w e r e d e s c r i b e d a n d a n a l y s e d b y

P h i r i ( 1 9 7 3 ) .

V e g e t a t i o n : *

A c c o r d i n g t o t h e c l a s s i f i c a t i o n o f B r o o k e e t a l . ( 1 9 7 0 ) , ( w h i c h

i s b a s e d o n t h e Z u r i c h - M o n t p e l l i e r s y n e c o l o g i c a l c l a s s i f i c a t i o n s c h e m e ) f o r

t h e M o u n t a i n H e m l o c k z o n e , t h e S q u a m i s h p l o t i s l o c a t e d i n t h e a b i e t o -

t s u g o s u m m e r t e n s i a n a e v a r i a n t o f t h e A b i e t o - T s u g e t u m m e r t e n s i a n a e a s s o c i a -

* L a t i n n ames o f t r e e s a r e t o b e f o u n d i n A p p e n d i x 1.

27

t i o n . T h e t r e e l a y e r i s f o r m e d . b y t w o s p e c i e s , P a c i f i c s i l v e r f i r a n d

m o u n t a i n h e m l o c k . Y e l l o w c e d a r w h i c h i s a r e p r e s e n t a t i v e s p e c i e s f o r t h i s

a s s o c i a t i o n ( B r o o k e , e t a l . , 1 9 7 0 ) w a s n o t p r e s e n t o n t h e p l o t . P a c i f i c

s i l v e r f i r c o m p r i s e d 69 p e r c e n t o f t h e s t a n d w i t h m o u n t a i n h e m l o c k m a k i n g

up t h e r e m a i n i n g 3 1 p e r c e n t .

L i s t e d i n o r d e r o f a b u n d a n c e , t h e u n d e r s t o r e y v e g e t a t i o n o n t h e

S q u a m i s h p l o t c o n s i s t e d o f V e r a t r u m e s c h s c h o l t z i i A . G r a y , V a c c i n i u m mem-

b r a n a e e u m D o u g l . e x H o o k , a l a s k a e n s e H o w e l l , V_. o v a l i f o l i u m S m i t h , L u e t k e a

p e c t i n a t a ( P u r s h ) K u n t z e , R u b u s p e d a t u s J . E. S m i t h , a n d L u z u l a s p p . V a c c i n i u m

o v a l i f o l i u m a n d L u e t k e a p e c t i n a t a w e r e o b s e r v e d o n l y i n t h e d e p r e s s i o n a t

t h e l o w e r e n d o f t h e p l o t . T h e V a c c i n i u m s p p . a n d R u b u s p e d a t u s a r e i n d i c a -

t i v e o f e x t r e m e l y a c i d s o i l c o n d i t i o n s ( B r o o k e , p e r s o n a l c o m m u n i c a t i o n ) ,

w h i l e V e r a t r u m e s c h s c h o l t z i i i s i n d i c a t i v e o f t e m p o r a r y s o i l s e e p a g e ( K r a j i n a ,

p e r s o n a l c o m m u n i c a t i o n ) . T h e m o s s l a y e r w h i c h w a s p r e s e n t i n s c a t t e r e d p a t -

c h e s w a s d o m i n a t e d b y D i c r a n u m f u s c e s c e n s T u r n , a n d D_. s c o p a r i u m Hedw. r e -

f l e c t i n g t h e p r e s e n c e o f a c i d r a w h u m u s .

U s i n g t h e same c l a s s i f i c a t i o n s c h e m e a s f o r t h e S q u a m i s h p l o t ,

t h e H a n e y p l o t i s a s s i g n e d t o t h e o r t h i c V a c c i n i u m - P l a g i o t h e c i u m e c o s y s t e m

t y p e o f t h e A b i e t o - T s u g e t u m h e t e r o p h y l l a e f o r e s t p l a n t a s s o c i a t i o n ( O r l o c i ,

1 9 6 5 ) . T h e t r e e s p e c i e s o n t h e p l o t w e r e w e s t e r n r e d c e d a r , w e s t e r n hem-

l o c k a n d y e l l o w c e d a r c o m p r i s i n g 2 8 % , 5 9 % a n d 1 3 % o f t h e s t a n d , r e s p e c t i v e l y .

S i z e c l a s s f r e q u e n c i e s a r e s h o w n b y s p e c i e s i n T a b l e 1 1 .

28

L i s t e d i n o r d e r o f a b u n d a n c e t h e u n d e r s t o r e y v e g e t a t i o n o n t h e

H a n e y p l o t c o n s i s t e d o f M e n z i e s i a f e r r u g i n e a S m i t h , V a c c i n i u m p a r v i f o l i u m

S m i t h , V_. a l a s k a e n s e H o w e l l , V . o v a l i f o l i u m S m i t h , G o o d y e r a o b l o n g i f o l i a

R a f . , a n d G a u l t h e r i a s h a l l o n P u r s h , t h e l a t t e r b e i n g a s s o c i a t e d w i t h t h i c k

a c c u m u l a t i o n s o f o r g a n i c m a t t e r o n t h e l i t h i c f o l i s o l s . T h e m o s s l a y e r w a s

a l m o s t e x c l u s i v e l y R h y t i d i o p s i s r o b u s t a ( H o o k . ) B r o t h .

29

I V METHODS

F i e l d S a m p l i n g : *

A l l t r e e s o n t h e s a m p l e p l o t s w e r e n u m b e r e d a n d m e a s u r e d f o r d . b . h . ,

t o t a l t r e e l e n g t h , c r o w n l e n g t h , a n d c r o w n d i a m e t e r ( T a b l e 1 2 ) . T h e d . b . h .

d a t a w e r e g r o u p e d i n t o 4 - i n c h ( 1 0 . 1 6 cm) s i z e c l a s s e s b y s p e c i e s , r e s u l t i n g

i n s i x s u c h s i z e c l a s s e s ( 6 " t o 3 0 " ; 1 5 . 2 4 cm t o 7 6 . 2 0 cm) f o r t h e H a n e y

p l o t a n d e l e v e n f o r t h e S q u a m i s h p l o t ( 6 " t o 5 0 " ; 1 5 . 2 4 cm t o 1 2 7 c m ) ,

( T a b l e 1 1 ) . T r e e s f o r s a m p l i n g w e r e s e l e c t e d f r o m m o s t o f t h e s i z e c l a s s e s ,

t h e s e l e c t i o n b e i n g m ade a t r a n d o m w i t h i n e a c h s i z e c l a s s s a m p l e d . A t o t a l

o f 13 t r e e s w e r e s a m p l e d a t t h e H a n e y p l o t a n d 2 0 a t t h e S q u a m i s h p l o t . B e -

c a u s e o f t i m e l i m i t a t i o n s , e i g h t o f t h o s e a t t h e S q u a m i s h p l o t w e r e s a m p l e d

f o r s t e m b i o m a s s o n l y , b r a n c h e s a n d f o l i a g e b e i n g e x c l u d e d .

S a m p l e t r e e s w e r e f e l l e d a n d t h e f o l l o w i n g d a t a r e c o r d e d : a g e a t

s t u m p ( a b o u t 1 f t . h i g h ; 3 0 . 5 c m ) , t o t a l l e n g t h o f s t e m , l e n g t h t o s i x i n c h

d i a m e t e r , l e n g t h t o o n e i n c h d i a m e t e r , d . b . h . , c r o w n l e n g t h , a n d t h e a v e r a g e

l e n g t h o f t h e t w o l a r g e s t b r a n c h e s .

B r a n c h e s w e r e s e v e r e d f r o m t h e s t e m a n d d i v i d e d m a n u a l l y i n t o

t h r e e g r o u p s : b i g b r a n c h e s ( > 1 i n c h d i a m e t e r ; 2.54 c m ) , s m a l l b r a n c h e s

(1-H i n . d i a m e t e r ) , a n d t w i g s (<Ch i n , d i a m e t e r ; 0.64 c m ) , w i t h f o l i a g e .

S e p a r a t i n g s m a l l b r a n c h e s f r o m t w i g s w i t h f o l i a g e w a s t h e m o s t t i m e - c o n s u m i n g

* F i e l d w o r k r e q u i r e d 1 3 m a n - m o n t h s .

3 0

T A B L E 1 1 : D i a m e t e r c l a s s d i s t r i b u t i o n o n t h e s a m p l e p l o t s ,

F r e q u e n c y o f t r e e s

S q u a m i s h H a n e y DBH *

C l a s s ( i n . )

M o u n t a i n h e m l o c k

P a c i f i c s i l v e r f i r

W e s t e r n h e m l o c k

W e s t e r n r e d c e d a r

Y e l l o w c e d a r

6-10 4 3 25 ( 3 ) 2 6 ( 2 )

1 0 - 1 4 6 ( 0 + 1 ) 19 ( 1 + 1 ) 12 ( 2 ) 7 2

1 4 - 1 8 1 0 ( 1 ) 2 3 ( 1 + 1 ) 18 ( 2 ) 7 5 ( 1 )

1 8 - 2 2 1 2 ( 1 ) 2 9 ( 1 + 1 ) 1 0 ( 1 ) 1 1 ( 0 + 1 ) -2 2 - 2 6 1 2 ( 1 ) 27 ( 1 ) - 5 2 ( 1 )

2 6 - 3 0 15 ( 1 + 2 ) 27 ( 1 ) 2 - -3 0 - 3 4 6 ( 1 ) 3 1 ( 1 + 1 ) - - -3 4 - 3 8 3 15 (1+D - - -3 8 - 4 2 2 4 - - -4 2 - 4 6 1 3 - - -> 4 6 3 - - - -

F i r s t n u m b e r i n b r a c k e t s s t a t e s n u m b e r o f c o m p l e t e l y s a m p l e d t r e e s . W h e r e a s e c o n d n u m b e r i s s h o w n i t i n d i c a t e s t h e n u m b e r o f t r e e s s a m p l e d f o r t r e e b o l e o n l y .

* ( d i a m e t e r b r e a t h e i g h t m e a s u r e d 4,5 f e e t a b o v e g r o u n d . )

T A B L E 1 2 : B a s i c s t a t i s t i c s o f o r i g i n a l m e a s u r e m e n t s f o r a l l t r e e s o v e r 6 i n c h e s o f d . b . h . o n t h e t w o s a m p l e p l o t s , n ; m o u n t a i n hem-l o c k = 7 4 ; P a c i f i c s i l v e r f i r = 1 8 2 ; w e s t e r n h e m l o c k = 6 7 ; y e l l o w c e d a r = 1 5 ; w e s t e r n r e d c e d a r = 3 2 .

T r e e T r e e M e a n S t a n d . M i n i m u m M a x i m u m P a r a m e t e r S p e c i e s U n i t V a l u e D e v . V a l u e V a l u e

T r e e M.h. m 0 . 6 2 1 0 . 2 4 6 0 . 2 3 1 1 . 4 1 0 d i a m e t e r P . s . f . 0 . 6 2 1 0 . 2 1 0 0 . 1 8 8 1 . 1 3 8

W.h. 0 . 3 3 2 0 . 1 3 3 0.157 0 . 7 0 6 Y . c . 0 . 3 4 8 0 . 1 3 6 0 . 1 8 0 0 . 6 3 5 W . r . c . 0 . 4 3 9 0 . 1 2 1 0 . 1 5 5 0 . 6 5 5

T r e e M.h. m 2 8 . 2 1 0 . 2 7.0 4 5 . 0 h e i g h t P . s . f . 3 0 . 8 8.9 9.0 4 6 . 0

W.h. 2 0 . 8 4.8 1 2 . 0 3 1 . 0 Y . c . 2 1 . 1 4.7 1 2 . 5 2 8 . 0 W . r . c . 2 0 . 9 4.8 7.0 3 0 . 0

C r o w n M.h. m 1 9 . 8 8.9 1.5 3 6 . 0 l e n g t h P . s . f . 2 0 . 8 7.8 1.5 3 8 . 0

W.h. 1 4 . 4 4.2 5.0 2 4 . 5 Y . c . 1 4 . 0 3.8 7.0 2 0 . 0 W . r . c . 1 4 . 4 3.8 4.0 2 5 . 0

C r o w n M.h. m 6.5 1.8 3.2 1 1 . 0 d i a m e t e r P . s . f . 6.2 1.5 1.8 1 0 . 2

W.h. 6.5 1.9 2.9 1 1 . 7 Y . c . 6.6 1.6 3.7 9.1 W . r . c . 5.0 2.5 0.5 8.5

V o l u m e o f M.h. m 3 4 . 0 4 8 — — —

t r e e b o l e P . s . f . 4 . 6 8 2 - - -W.h. 0 . 8 9 6 - - -Y . c . 1 . 0 3 5 - - -W . r . c . 1 . 3 0 3 - - -

j o b , a n d f o r t r e e s w i t h a l a r g e a m o u n t o f b r a n c h e s a s u b - s a m p l e ( a p p r o x i -

m a t e l y 1 0 p e r c e n t ) w a s t a k e n f r o m w h i c h t h e p r o p o r t i o n s o f t h e s e t w o

32

b i o m a s s c o m p o n e n t s w a s d e t e r m i n e d , T w i g s a n d f o l i a g e w e r e s a m p l e d s e p a r -

a t e l y f o r t h e t o p , m i d d l e a n d l o w e r p a r t o f t h e t r e e c r o w n .

T h e f r e s h w e i g h t o f e a c h b r a n c h g r o u p w a s t a k e n a s q u i c k l y a s

p o s s i b l e t o m i n i m i z e l o s s o f m o i s t u r e . Two r e p r e s e n t a t i v e s a m p l e s w e r e

t a k e n f r o m e a c h g r o u p : ( a ) t o d e t e r m i n e o v e n d r y w e i g h t ( d r i e d a t 1 5 0 ° C)

a n d m o i s t u r e c o n t e n t , ( b ) t o d e t e r m i n e c o n c e n t r a t i o n o f m a c r o n u t r i e n t s

( d r i e d a t 70° C ) , T h e s a m p l e s w e r e t a k e n a c c o r d i n g t o a t a b l e o f r a n d o m

n u m b e r s w h i c h s e l e c t e d t h e p o s i t i o n o f t h e s a m p l e a l o n g t h e b r a n c h . T h e

s a m p l e s c o n s t i t u t e d b e t w e e n 0.5 p e r c e n t a n d 5 p e r c e n t o f t h e t o t a l b r a n c h

b i o m a s s , a n d w e i g h e d 2.0 - 4.0 k g f o r b i o m a s s d e t e r m i n a t i o n , a n d 1.0 - 1.5

k g f o r c h e m i c a l a n a l y s i s .

T h e s a m p l e s o f b r a n c h e s u n d e r % - i n c h d i a m e t e r ( w h i c h i n c l u d e d

f o l i a g e ) w e r e d r i e d a t 70° C t h e d a y t h e y w e r e c u t i n o r d e r t o p r e v e n t

c h a n g e o f n u t r i e n t c o n t e n t i n t h e n e e d l e s . H e m l o c k f o l i a g e w a s e a s i l y s e p -

a r a t e d f r o m t w i g s f o l l o w i n g d r y i n g . T h e f o l i a g e a n d t w i g s o f P a c i f i c s i l -

v e r f i r , o n t h e o t h e r h a n d , d i d n o t s e p a r a t e s o e a s i l y a n d t h e r e f o r e t h e s e

t w o b i o m a s s c a t e g o r i e s w e r e p o o l e d . F o l i a g e o f y e l l o w c e d a r w a s s e p a r a t e d

m a n u a l l y f r o m t h e b r a n c h e s a t t h e p o i n t a t w h i c h t h e b r a n c h e s t u r n e d b r o w n .

T h e t w i g s a n d f o l i a g e b i o m a s s s a m p l e s w e i g h e d b e t w e e n 2,0 - 4,0 k g . T h e

w e i g h t o f c h e m i c a l s a m p l e s w a s b e t w e e n 1.0 - 1.5 k g .

T r e e b o l e s w e r e t r i m m e d a t 1 - i n c h ( 2 , 5 4 cm) d i a m e t e r , a n d t h e n

d i v i d e d u p i n t o s e c t i o n s e q u a l t o 1 0 p e r c e n t o f t h e t r i m m e d l e n g t h . Two

3 3

m e a s u r e m e n t s w e r e made o f b o t h o u t s i d e a n d i n s i d e b a r k d i a m e t e r a t r i g h t

a n g l e s t o e a c h o t h e r a t t h e m i d d l e a n d e a c h e n d o f t h e s e c t i o n s . Two d i s c s ,

a p p r o x i m a t e l y 5 cm i n t h i c k n e s s , w e r e c u t f r o m t h e l a r g e e n d o f e a c h s t e m

s e c t i o n , a n d t h e f r e s h w e i g h t o f t h e b a r k a n d w o o d f r o m o n e o f t h e m d e t e r -

m i n e d . One d i s c w a s f o r b i o m a s s d e t e r m i n a t i o n a n d t h e o t h e r f o r c h e m i c a l

a n a l y s i s . D i s c s s a t u r a t e d w i t h w a t e r ( b y i m m e r s i o n i n c r e e k f o r 24 h o u r s )

f o r b i o m a s s d e t e r m i n a t i o n w e r e p l a c e d i n p l a s t i c b a g s p a r t i a l l y f i l l e d w i t h

w a t e r t o p r e v e n t m o i s t u r e l o s s a n d t h e a c c o m p a n y i n g s h r i n k a g e .

L a b o r a t o r y A n a l y s e s : *

B i o m a s s

T h e v o l u m e o f b a r k a n d w o o d w a s d e t e r m i n e d o n o n e o f e a c h o f t h e

p a i r s o f s t e m d i s c s b y t h e d i s p l a c e m e n t m e t h o d ( H e i n r i c h s 1 9 5 4 ) . E a c h s a m -

p l e w a s c o m p l e t e l y i m m e r s e d i n w a t e r , ( b u t w i t h o u t t o u c h i n g t h e b o t t o m o r

w a l l s o f t h e c o n t a i n e r ) a n d i t s v o l u m e d e t e r m i n e d b y s u b t r a c t i n g t h e w e i g h t

o f w a t e r f r o m t h e w e i g h t o f w a t e r w i t h t h e s a m p l e f u l l y i m m e r s e d . T h e s a m -

p l e s w e r e t h e n d r i e d a t 105° C f o r 4 8 h o u r s a n d w e i g h e d . S p e c i f i c g r a v i t y

w a s c a l c u l a t e d f r o m t h e v o l u m e a n d w e i g h t o f e a c h d i s c ( A p p e n d i x 9 ) . B r a n c h

b i o m a s s s a m p l e s w e r e d r i e d a t 1 0 5 ° C f o r 48 h o u r s a n d w e i g h e d . M o i s t u r e

c o n t e n t w a s c a l c u l a t e d a s t h e d i f f e r e n c e b e t w e e n f r e s h w e i g h t a n d o v e n d r y

w e i g h t .

* L a b o r a t o r y a n a l y s e s r e q u i r e d 7 m a n - m o n t h s .

34

S a m p l e s f o r c h e m i c a l a n a l y s i s w e r e d r i e d a t 70° C ± 5 ° C f o r

4 8 h o u r s . I m m e d i a t e l y f o l l o w i n g d r y i n g , s a m p l e s w e r e g r o u n d i n a W i l e y

m i l l t o p a s s a 2-mm m e s h a n d s e a l e d i n p o l y t h e n e b a g s t o a w a i t d r y - a s h i n g

f o r t h e d e t e r m i n a t i o n o f K, C a a n d Mg. P o r t i o n s o f t h i s m a t e r i a l w e r e r e -

g r o u n d t o p a s s a #40 m e s h a n d s e a l e d i n b a g s a s a b o v e t o a w a i t K j e l d a h l

d i g e s t i o n f o r t h e d e t e r m i n a t i o n o f N a n d P. S a m p l e s w e r e s t o r e d f o r b e -

t w e e n 2 a n d 1 0 m o n t h s p r i o r t o a n a l y s e s .

S i n c e i t w a s s u s p e c t e d t h a t t h e s a m p l e s m i g h t h a v e a b s o r b e d

m o i s t u r e w h i l s t b e i n g g r o u n d , 1 1 6 s a m p l e s c o v e r i n g a l l t y p e s o f m a t e r i a l

w e r e s e l e c t e d a t r a n d o m i n O c t o b e r 1 9 7 2 ( a b o u t 2 m o n t h s a f t e r g r i n d i n g ) a n d

t h e m o i s t u r e c o n t e n t d e t e r m i n e d b y r e - d r y i n g a t b o t h 70° C a n d 1 0 5 ° C. T h e

r e s u l t s ( T a b l e 1 3 ) c o n f i r m t h a t m o i s t u r e h a d i n d e e d b e e n a b s o r b e d b e f o r e

t h e s a m p l e s w e r e s e a l e d i n t h e b a g s . A p p r o p r i a t e c o r r e c t i o n s w e r e made t o

t h e n u t r i e n t c o n c e n t r a t i o n d a t a .

T A B L E 1 3 : P e r c e n t l o s s o f w e i g h t i n s a m p l e s f o r c h e m i c a l a n a l y s e s u p o n -r e - d r y i n g a f t e r 2-6 m o n t h s o f s t o r a g e .

N o . o f 70°C D r y i n g 105°C D r y i n g S a m p l e S p e c i f i c a t i o n S a m p l e s T e m p e r a t u r e T e m p e r a t u r e

A n a l y s e d M e a n SD M e a n SD

Wood 20 5.06 1.01 5.63 0.88 B a r k 2 0 4.77 0.67 5.84 0.66 B i g B r a n c h e s 18 5 . 5 0 2.24 6.32 2.30 S m a l l B r a n c h e s 2 0 4.93 1.64 5.86 1.64 T w i g s 20 4.83 0,85 5.94 0 . 8 1 F o l i a g e 1 8 2.98 0.86 4 . 3 1 0.89

A v e r a g e f r o m a l l g r o u p s 4.66 0 . 0 2 3 5.65 0.03

3 5

P a n d N A n a l y s i s

T h e m e t h o d u s e d w a s a m o d i f i e d K j e l d a h l d i g e s t i o n ( T w i n e a n d

W i l l i a m s , 1 9 7 1 ) u s i n g s e l e n i u m a s a c a t a l y s t w i t h s u b s e q u e n t a n a l y s i s f o r

N a n d P u s i n g a T e c h n i c o n A u t o ^ - A n a l y s e r . B e t w e e n 0.15 a n d 0.25 gm o f s a m -

p l e w a s a d d e d t o a b o i l i n g t u b e c o n t a i n i n g 5 m l o f d i g e s t i o n m i x t u r e a n d

s e l e n i z e d H e n g a r g r a n u l e s t o f a c i l i t a t e e v e n b o i l i n g . T h e t u b e s w e r e p l a c e d

i n a 7 7 - p l a c e h e a t i n g b l o c k a n d d i g e s t e d a t 8 0 t o 1 0 0 ° C f o r 1 2 h o u r s , t h e n

s l o w l y r a i s e d t o t h e b o i l i n g t e m p e r a t u r e o f t h e d i g e s t i o n m i x t u r e a n d h e l d

t h e r e u n t i l c o m p l e t e l y c o l o u r l e s s a n d t r a n s p a r e n t . G l a s s b a l l s p l a c e d o v e r

t h e m o u t h s o f t h e t u b e s a c t e d a s r e f l u x c o n d e n s e r s , w a s h i n g u n d i g e s t e d

m a t e r i a l s p a t t e r e d o n t o t h e w a l l s o f t h e t u b e b a c k down i n t o t h e d i g e s t i o n

m i x t u r e . When t h e d i g e s t s w e r e c o o l t h e y w e r e d i l u t e d t o 1 0 0 m l w i t h d e -

i o n i z e d w a t e r , a n d l e f t t o s t a n d o v e r n i g h t t o p e r m i t s e t t l i n g o f a n y u n d i -

g e s t e d s o l i d s . T h e s u p e r n a t a n t d i g e s t a t e w a s d e c a n t e d i n t o p l a s t i c b o t t l e s

t o a w a i t a n a l y s i s .

N i t r o g e n w a s d e t e r m i n e d u s i n g a n a m m o n i a c a r t r i d g e o f a T e c h n i c o n

A u t o - A n a l y s e r w h i c h m a k e s u s e o f t h e B e r t h e l o t r e a c t i o n ( r e a c t i o n o f a m m o n i a

w i t h s o d i u m p h e n a t e a n d s o d i u m h y p o c h l o r i t e t o y i e l d a b l u e i n d o p h e n o l c om -

p l e x w h i c h i s q u a n t i f i e d i n a c o l o r i m e t e r ) .

P h o s p h o r u s w a s d e t e r m i n e d u s i n g a n o r t h o - p h o s p h a t e c a r t r i d g e o f

a T e c h n i c o n A u t o - A n a l y s e r , T h i s i n v o l v e s f o r m a t i o n o f a r e d u c e d p h o s p h o -

m o l y b d a t e c o m p l e x , w h i c h i s q u a n t i f i e d i n a c o l o r i m e t e r .

36

C a t i o n A n a l y s i s

A n a l y s i s f o r K, C a a n d Mg w a s c o n d u c t e d o n a V a r i a n T e c h t r o n

A t o m i c A b s o r p t i o n S p e c t r o p h o t o m e t e r ( M o d e l A A 5 ) f o l l o w i n g d r y - a s h i n g . One

gm o f s a m p l e w a s w e i g h e d i n t o a s i l i c a c r u c i b l e a n d a s h e d i n a m u f f l e f u r -

n a c e a t 200° C f o r 1 % - 2 h o u r s , f o l l o w e d b y 4 7 5 ° C ± 5 ° C f o r 1 2 h o u r s .

T h e w e i g h t o f a s h f o r e a c h s a m p l e w a s d e t e r m i n e d a n d t h e n t a k e n up i n 5 m l

o f 2 0 % H C 1 . A f u r t h e r 1 5 m l o f h y d r o c h l o r i c a c i d w a s t h e n a d d e d a n d t h e

c r u c i b l e h e a t e d o n a s a n d b a t h t o f a c i l i t a t e s o l u t i o n . T h e s o l u t i o n w a s

t h e n t r a n s f e r r e d t o a v o l u m e t r i c f l a s k , made up t o 1 0 0 m l , a n d t r a n s f e r r e d

t o a p l a s t i c b o t t l e f o r s t o r a g e p r i o r t o a n a l y s i s . A n a i r - a c e t y l e n e f l a m e

w a s u s e d f o r a l l d e t e r m i n a t i o n s .

B e c a u s e t h e r e a r e a n u m b e r o f i n t e r f e r e n c e s p o s s i b l e i n t h e m e t h o d

u s e d , t h e f o l l o w i n g i n v e s t i g a t i o n s w e r e u n d e r t a k e n :

1. T o c h e c k t h e i n f l u e n c e o f p y r o p h o s p h a t e s a n d h y d r a t e d s i l i c a

o n d e t e r m i n a t i o n o f C a , Mg a n d K, 2 8 s a m p l e s w e r e p r o c e s s e d a s f o l l o w s

( A l l a n , 1969).. T h e a s h o b t a i n e d f r o m t h e m u f f l e f u r n a c e w a s d i s s o l v e d i n

5 m l o f 2 0 % HC1 a n d s l o w l y t a k e n t o d r y n e s s o n a s a n d h e a t e r ( t o h y d r o l y z e

t h e p y r o p h o s p h a t e s a n d t o d e h y d r a t e t h e s i l i c a ) . T h i s o p e r a t i o n w a s r e p e a -

t e d w i t h a f u r t h e r 5 m l o f 2 0 % H C 1 . T h e r e s i d u e w a s t h e n d i s s o l v e d i n 2 0 m l

o f 2 0 % H C 1 , a n d a n a l y s e d o n a n A t o m i c A b s o r p t i o n S p e c t r o p h o t o m e t e r . No s i g -

n i f i c a n t d i f f e r e n c e s w e r e f o u n d b e t w e e n t h e c o n c e n t r a t i o n s o f C a , Mg a n d K

o b t a i n e d b y t h i s m e t h o d a n d t h o s e o b t a i n e d b y t h e b a s i c m e t h o d w h i c h vras

t h e r e f o r e u s e d t h r o u g h o u t t h e s t u d y .

3 7

2, T o d e t e r m i n e t h e c h e m i c a l i n t e r f e r e n c e o f p h o s p h o r o u s , t w o

r e l e a s i n g a g e n t s w e r e t e s t e d : s t r o n t i u m c h l o r i d e a n d l a n t h a n u m c h l o r i d e ,

t h e f o r m e r a t 3 , 0 0 0 ppm a n d t h e l a t t e r a t 1 0 , 0 0 0 ppm ( D a v i d , I 9 6 0 ) , No

s i g n i f i c a n t d i f f e r e n c e w a s f o u n d i n t h e c o n c e n t r a t i o n o f C a a n d Mg u s i n g

e i t h e r o f t h e s e r e l e a s i n g a g e n t s a s c o m p a r e d t o t h e b a s i c m e t h o d w h i c h w a s

t h e r e f o r e u s e d t h r o u g h o u t t h e s t u d y .

C h e c k o n t h e a c c u r a c y o f t h e c h e m i c a l a n a l y s i s

T o v e r i f y t h e r e l i a b i l i t y o f t h e c h e m i c a l a n a l y s e s , some s a m p l e s

w e r e s e n t t o o t h e r l a b o r a t o r i e s f o r a n a l y s i s . F o u r s a m p l e s w e r e s e n t t o

t h e P a c i f i c F o r e s t R e s e a r c h C e n t r e i n V i c t o r i a , B. C., a n d f i v e s a m p l e s w e r e

s e n t t o t h e R e s e a r c h D i v i s i o n o f t h e B r i t i s h C o l u m b i a F o r e s t S e r v i c e i n

V i c t o r i a , B. C. T h e r e s u l t s f r o m a l l t h r e e l a b o r a t o r i e s a r e s u m m a r i z e d i n

T a b l e 1 4 , T h e r e s u l t s o f t h i s s t u d y s h o w c o n s i s t e n t l y l o v e r f i g u r e s t h a n

t h o s e f r o m t h e o t h e r l a b o r a t o r i e s , s u g g e s t i n g a g e n e r a l u n d e r - e s t i m a t i o n i n

my d a t a . O n l y o n e s a m p l e ( t r e e n u m b e r 6 8 ) s h o w s s e r i o u s ( 4 0 p e r c e n t ) u n d e r -

e s t i m a t i o n h o w e v e r . F r o m t h i s t a b l e i t a p p e a r s t h a t t h e d a t a o n b i o m a s s

c h e m i c a l c o n t e n t s may b e u n d e r - e s t i m a t e d b u t i n g e n e r a l , my r e s u l t s a r e

r e a s o n a b l y a c c u r a t e .

3 8

T A B L E 1 4 : C o m p a r i s o n o f r e s u l t s o f c h e m i c a l a n a l y s e s f o r f i v e r a n d o m l y s e l e c t e d s a m p l e s ( r e s u l t s a r e i n p e r c e n t ) ,

T r e e R e s u l t S a m p l e S p e c i f i c a t i o n N o , o f

P. s i l , f i r , f o l i a g e 22 UBC 1

( l o w t h i r d o f c r o w n ) B C F S 2

P F R C 3

M o u n . h e m l o c k , t w i g s 6 8 UBC ( l o w t h i r d o f c r o w n ) B C F S

P F R C

M o u n , h e m l o c k , t w i g s 1 1 9 UBC ( l o w t h i r d o f c r o w n ) B C F S

P F R C

P. s i l . f i r , b a r k 1 7 8 UBC ( d i s c N o . 7) B C F S

P F R C

P. s i l , f i r , b a r k 2 1 5 UBC ( d i s c N o . 5 ) B C F S

P F R C

1 - U n i v e r s i t y o f B r i t i s h C o l u m b i a , L a b o r a t o r y .

2 - B r i t i s h C o l u m b i a F o r e s t S e r v i c e , 3 - C a n a d i a n F o r e s t r y S e r v i c e , P a c i f

N P K C a Mg

0.76 0.77 0.75

0.107 0 . 1 0 8 0 . 0 8 6

0 . 3 1 0.39 0.22

0,56 0.69 0.66

0 . 0 7 3 0 . 0 7 6 0 . 0 7 6

0.24 0.55 0.46

0.034 0 . 0 7 1 0 . 0 6 0

0.17 0.24 0.14

0.12 0.20 0.24

0 . 0 4 5 0 . 0 5 3 0 . 0 5 6

0,40 0.42 0.43

0 . 0 6 1 0 . 0 6 5 0 . 0 6 0

0.14 0.23 0.13

0.19 0.28 0.23

0 . 0 4 6 0 . 0 5 5 0 . 0 5 6

0.18 0.22 0.17

0 . 0 3 5 0 . 0 3 6 0 . 0 3 0

0 . 1 1 0.15 0.10

0 . 3 1 0.36 0.30

0 . 0 3 1 0.034 0 . 0 2 2

0 . 2 1 0.22

0.037 0.037

0.08 0.10

0.29 0.35

0 . 0 2 2 0. 0 2 5

F a c u l t y o f F o r e s t r y , F o r e s t E c o l o g y

R e s e a r c h D i v i s i o n , V i c t o r i a , i c F o r e s t R e s e a r c h C e n t r e , V i c t o r i a .

D a t a A n a l y s i s : *

B i o m a s s D a t a

D r y b i o m a s s o f w o o d a n d b a r k w a s c a l c u l a t e d f r o m s p e c i f i c g r a v i t y

a n d v o l u m e d a t a f o r i n d i v i d u a l t r e e b o l e s e c t i o n s , V o l u m e o f e a c h s e c t i o n

* D a t a a n a l y s i s r e q u i r e d 5 m a n - m o n t h s .

3 9

( w o o d a n d b a r k ) w a s c a l c u l a t e d a c c o r d i n g t o N e w t o n ' s f o r m u l a :

ft Ik ( d 2 + 4 d | + d 2 )

w h e r e d ^ , d ^ a n d d 2 a r e d i a m e t e r s a t b o t h e n d s a n d t h e m i d d l e o f t h e s e c t i o n ,

r e s p e c t i v e l y ; S i s t h e l e n g t h o f t h e s e c t i o n . V o l u m e o f w o o d w a s c a l c u l a t e d

i n t h e same m a n n e r . V o l u m e o f b a r k w a s o b t a i n e d b y s u b t r a c t i n g t h e v o l u m e

o f w o o d f r o m t h e v o l u m e o f t h e w h o l e s e c t i o n . T h e v o l u m e s o f t r e e b o l e s

c a l c u l a t e d a s t h e sum o f t h e v o l u m e s o f t h e t e n s e c t i o n s u s i n g N e w t o n ' s

f o r m u l a w e r e c o m p a r e d w i t h v o l u m e s o b t a i n e d u s i n g B. C. F o r e s t S e r v i c e v o l -

ume e q u a t i o n s ( T a b l e 1 5 ) . V o l u m e s c a l c u l a t e d b y t h e f o r m e r m e t h o d w e r e

f o u n d t o b e s m a l l e r t h a n v o l u m e s o b t a i n e d u s i n g t h e B, C. F o r e s t S e r v i c e

v o l u m e e q u a t i o n s . T h e d i f f e r e n c e m i g h t b e e x p l a i n e d b y a g r e a t e r t a p e r t h a n

i s a s s u m e d i n t h e F o r e s t S e r v i c e e q u a t i o n s o r i t c a n b e t h e r e s u l t o f s a m -

p l i n g a s m a l l n u m b e r , o f t r e e s .

D r y b i o m a s s o f w o o d f o r a n y s e c t i o n w a s o b t a i n e d b y m u l t i p l y i n g

t h e v o l u m e o f w o o d f o r t h a t s e c t i o n b y t h e m e a n o f t h e v a l u e s o f s p e c i f i c

g r a v i t y o f w o o d f o r t h a t s e c t i o n a n d t h e n e x t s e c t i o n u p t h e s t e m . B a r k

b i o m a s s w a s e s t i m a t e d i n t h e same w a y .

B ( ± ) „ sgU) Y S s C i + i) . v ( i )

w h e r e B i s b i o m a s s , S g i s s p e c i f i c g r a v i t y a n d V i s v o l u m e o f t h e i - t h s e c -

t i o n . O n l y o n e s p e c i f i c g r a v i t y v a l u e w a s u s e d f o r t h e u p p e r m o s t s e c t i o n .

4 0

T A B L E 1 5 : A v e r a g e p e r c e n t d i f f e r e n c e b e t w e e n t r e e t r u n k v o l u m e s c a l c u l a -t e d a c c o r d i n g t o B C F S v o l u m e e q u a t i o n s a n d t h o s e c a l c u l a t e d s e c t i o n - b y - s e c t i o n a c c o r d i n g t o N e w t o n ' s f o r m u l a . N o , o f o b s e r -v a t i o n s : M o u n t a i n h e m l o c k = 8; P a c i f i c s i l v e r f i r = 1 2 ; W e s t e r n h e m l o c k = 8; Y e l l o w c e d a r = 4,

P E R CENT D I F F E R E N C E = V S N - V B F S # 1 0 0

V S N

VSN = v o l u m e o f t r e e b o l e a c c o r d i n g t o N e w t o n ' s f o r m u l a .

V B C F S = v o l u m e o f t r e e b o l e a c c o r d i n g t o B C F S v o l u m e e q u a t i o n .

T r e e S p e c i e s P e r C e n t D i f f e r e n c e *

M o u n t a i n h e m l o c k - 8.304

P a c i f i c s i l v e r f i r - 1 0 . 8 7 8

W e s t e r n h e m l o c k - 8 . 1 8 3

Y e l l o w c e d a r - 1 4 . 0 0 6

1 - R e a l v o l u m e ( s e c t i o n - b y - s e c t i o n ) i s s m a l l e r t h a n t a b u l a r v o l u m e .

B i o m a s s o f b i g b r a n c h e s , s m a l l b r a n c h e s a n d t w i g s a n d n e e d l e s

w a s c a l c u l a t e d b y m u l t i p l y i n g t h e f r e s h w e i g h t o f t h e p a r t i c u l a r c o m p o n e n t

b y t h e a p p r o p r i a t e f i g u r e f o r p e r c e n t a g e d r y w e i g h t ( A p p e n d i x 1 0 ) . B a s i c

s t a t i s t i c s f o r s a m p l e d t r e e s a r e s u m m a r i z e d i n T a b l e 1 6 . B i o m a s s d i s t r i b u -

t i o n f o r a l l s a m p l e d t r e e s i s p r e s e n t e d i n A p p e n d i c e s 4 a n d 5.

R e g r e s s i o n e q u a t i o n s w e r e d e v e l o p e d t o c a l c u l a t e t h e b i o m a s s b y

c o m p o n e n t s f o r a l l t r e e s o n t h e s a m p l e p l o t s w i t h t h e e x c e p t i o n o f y e l l o w

T A B L E 1 6 : B a s i c s t a t i s t i c s f o r s a m p l e d t r e e s , n ; M o u n t a i n h e m l o c k = 5; P a c i f i c s i l v e r f i r = 7; W e s t e r n h e m l o c k = 8; Y e l l o w c e d a r = 4.

T r e e T r e e M e a n S t a n d a r d M i n i m u m M a x i m u m P a r a m e t e r S p e c . U n i t V a l u e D e v i a t i o n V a l u e V a l u e

d . b . h . M.h. m 0 . 6 0 2 0 . 1 3 3 0.437 0 . 7 6 2 P . s . f . 0.587 0 . 2 2 1 0 . 3 1 0 0 . 9 0 4 W.h. 0 . 3 1 4 0.116 0 . 1 6 0 0 . 4 9 0 Y . c . 0 . 3 5 3 0 . 1 9 1 0 . 1 8 0 0 . 6 0 2

T r e e M.h. m 3 2 . 1 4.7 2 5 . 1 3 8 . 0 l e n g t h P . s . f . 3 0 . 9 8.7 1 7 . 8 4 1 . 2

W.h. 2 0 . 8 4.3 1 5 . 5 2 7 . 7 Y . c . 2 1 . 6 5.8 1 4 . 8 2 7 . 2

C r o w n M.h. m 2 3 . 9 2.0 2 0 . 6 2 5 . 8 l e n g t h P . s . f . 2 2 . 4 6.4 1 1 . 5 2 8 . 9

W.h. 1 5 . 3 4.3 1 0 . 3 2 3 . 2 Y . c . 1 3 . 2 2.9 9.1 1 5 . 4

C r o w n M.h. m 6.6 2.1 4.7 9.4 d i a m e t e r P . s . f . 7.1 1.3 5.5 8.5

W.h. 5.0 1.6 2.9 7.3 Y . c . 7.4 1.7 5.8 9.1

W e i g h t o f M.h. k g 1 3 5 2 . 6 5 7 5 . 6 7 0 2 . 1 9 9 4 , w o o d P . s . f . 1 3 3 2 . 5 1 0 7 0 . 9 1 7 5 3 0 6 6

W.h. 2 8 0 . 6 1 9 5 . 0 66 6 0 1 Y . c . 4 2 1 . 3 - 4 0 1 . 8 6 8 9 3 8

W e i g h t o f M.h. k g 3 6 4 . 1 1 6 3 . 6 1 7 8 0 5 7 5. b a r k P . s . f . 2 7 7 . 3 2 2 6 . 0 37„ 6 8 6

W.h. 6 6 . 2 5 2 . 5 1 1 , 1 5 3 Y . c . 3 5 . 7 3 2 . 8 6 79

W e i g h t o f M.h. k g 1 5 0 . 4 9 9 . 5 53; 2 9 0 b i g P . s . f . 1 6 1 . 9 1 5 3 . 8 33 4 1 1 b r a n c h e s W.h. 4 0 . 4 4 3 . 4 lo 1 2 2

Y . c . 4 6 . 1 5 7 . 4 4 126.

W e i g h t o f M.h. k g 3 6 . 5 1 1 . 1 2 4 5 4 s m a l l P . s . f . 5 3 . 3 2 4 . 7 1 8 86 b r a n c h e s W.h. 4 5 . 0 2 5 . 8 •16 74-

Y . c . 3 3 . 8 1 8 . 2 14 53-

T A B L E 16 ( c o n t i n u e d ) :

T r e e P a r a m e t e r

T r e e S p e c . U n i t

M e a n V a l u e

S t a n d a r d D e v i a t i o n

M i n i m u m V a l u e

M a x i m u m V a l u e

W e i g h t o f t w i g s

M.h. P . s . f . W.h. Y . c .

k g 4 9 . 6

1 2 . 6

1 8 . 9

8.1

24 ,

5,

70

24

W e i g h t o f f o l i a g e

M.h. P . s . f . W.h. Y . c .

k g 7 0 . 6

2 3 . 2

2 1 . 3

1 7 . 4

35.

3,

90 ,

4 8

W e i g h t o f t w i g s a n d f o l i a g e

M.h. P . s . f . W.h. Y . c .

k g 1 2 0 . 2 1 2 0 . 9 3 5 . 8 5 9 . 4

3 9 . 5 7 3 . 4 2 5 . 2 4 0 . 2

5 9 c

3 0 9

15

1 6 1 1 9 8 72.

1 1 0

W e i g h t o f w h o l e t r e e

M.h. P . s . f . W.h. Y . c .

k g 2 0 2 4 . 2 1 9 4 9 . 2 4 6 8 . 0 5 7 5 . 0

8 7 5 . 2 1 5 1 9 . 0 3 3 4 . 0 5 5 3 . 0

1 0 1 6 r

2 9 5 , 1 0 9 1 0 7 c

3 0 0 7 4 3 5 4 1 0 2 1 1 3 0 5

A g e M.h. P . s . f . W.h. Y . c .

Y e a r 4 3 7 4 0 5 2 8 3 1 7 9

3 2 5 3 0 0 1 1 5 1 1 5

5 3 0 5 3 0 4 1 0 2 2 1

4 3

c e d a r a t t h e H a n e y p l o t , Two r e g r e s s i o n m o d e l s w e r e t e s t e d ( F r e e s e , 1 9 6 4 ) .

A ) L i n e a r m o d e l :

Y = a + b x ( 1 )

B) E x p o n e n t i a l m o d e l :

Y = a X b C X ( 2 )

T r a n s f o r m e d t o l i n e a r l o g a r i t h m i c f o r m u s i n g

l o g a r i t h m i c t r a n s f o r m a t i o n

l o g Y = b Q + b1 l o g X + b 2 X ( 3 )

w h e n C = 1 , t h e n ( 2 ) b e c o m e s :

l o g Y = b 0 + b L l o g X ( 4 )

w h e n b = 0, t h e n ( 2 ) b e c o m e s :

l o g Y = b Q + b 2 X ( 5 )

T w e n t y - f i v e i n d e p e n d e n t v a r i a b l e s w e r e t e s t e d i n t h e s e r e g r e s s i o n

m o d e l s :

D D.CL + ( D . C L ) 2 B A . H.CL

H C L 2 B A.H + C L

C L D.CW2 D 2 , H + C L

D 2 . H D.CW.CL 2 B A

BA.II D . C L . B T L CW 2

D.CL D 2 . H . C L CW 2 . CL

D . C L 2 D 2.H.CW D,CW 2.CL

D . H . C L 2 ( B A . H ) + (CW.CL) C r o w n v o l u m e

C r o w n s u r f a c e

44

w h e r e D = d . b . h ,

H = t r e e l e n g t h

C L = c r o w n l e n g t h

CW = c r o w n w i d t h

BA = c r o s s s e c t i o n a l a r e a a t b r e a s t h e i g h t

B T L = t r e e l e n g t h m i n u s c r o w n l e n g t h

C r o w n v o l u m e = CW 2 ,CL

C r o w n s u r f a c e = 7V . CW . "1 4 C L + CW:

O f t h e s e i n d e p e n d e n t v a r i a b l e s , 14 w e r e f o u n d s i g n i f i c a n t a t 1 p e r c e n t

p r o b a b i l i t y l e v e l f o r d e s c r i b i n g b i o m a s s o f d i f f e r e n t t r e e c o m p o n e n t s i n

l i n e a r , l o g a r i t h m i c o r i n b o t h m o d e l s .

12

4

S i g n i f i c a n t v a r i a b l e s w e r e :

D 2 . H CR.VOL CW 2

B A C R .SURF D.CW2

BA.H D D.CL

D2.H.CW H D.CL 2

( BA.H). + (CL.CW). C L

B a s i c s t a t i s t i c s f o r i n d e p e n d e n t v a r i a b l e s a r e p r e s e n t e d i n A p p e n d i x 2.

I n d e p e n d e n t v a r i a b l e s , a s t h e y a p p e a r i n p a r t i c u l a r m o d e l s f o r d i f f e r e n t

t r e e c o m p o n e n t s a r e p r e s e n t e d i n A p p e n d i x 3,

4 5

L o g a r i t h m i c r e g r e s s i o n e q u a t i o n s w e r e u s e d t o e s t i m a t e b i o m a s s

f o r p a r t i c u l a r t r e e c o m p o n e n t s a n d t h e e n t i r e t r e e l a y e r b i o m a s s o n b o t h

s a m p l e p l o t s ( A p p e n d i x 1 3 ) , R e s u l t s a r e s u m m a r i z e d i n T a b l e 17, R e s i d u a l s

w e r e p l o t t e d f o r a l l r e g r e s s i o n e q u a t i o n s u s e d f o r c a l c u l a t i n g b i o m a s s o n

t h e s a m p l e p l o t s i n o r d e r t o t e s t f o r l a c k o f f i t .

N u t r i e n t C o n c e n t r a t i o n D a t a :

C o n c e n t r a t i o n o f e l e m e n t s w a s r e a d i n ppm ( p a r t s p e r m i l l i o n )

e i t h e r o n t h e T e c h n i c o n A u t o - A n a l y s e r ( f o r N a n d P ) . o r o n t h e A t o m i c A b s o r p -

t i o n S p e c t r o p h o t o m e t e r ( f o r C a , Mg, K ) . R e s u l t s w e r e t r a n s f e r r e d i n t o p e r -

c e n t a g e c o n c e n t r a t i o n . T h e a m o u n t o f n u t r i e n t s w i t h i n t h e s a m p l e d t r e e s

was o b t a i n e d b y m u l t i p l y i n g t h e b i o m a s s o f t h e v a r i o u s t r e e c o m p o n e n t s b y

t h e a p p r o p r i a t e f i g u r e f o r c o n c e n t r a t i o n o f n u t r i e n t e l e m e n t s a n d s u m m i n g

t h e p r o d u c t s . T h e d i s t r i b u t i o n o f n u t r i e n t s w i t h i n t h e s a m p l e d t r e e s a n d

c o n c e n t r a t i o n o f n u t r i e n t s w i t h i n d i f f e r e n t t r e e c o m p o n e n t s a r e l i s t e d i n

A p p e n d i c e s 6 a n d 7.

T h e t o t a l a m o u n t o f n u t r i e n t s c o n t a i n e d w i t h i n t h e a b o v e - g r o u n d

p o r t i o n o f t h e t r e e l a y e r o n t h e s a m p l e p l o t s w a s c a l c u l a t e d b y m u l t i p l y i n g

t h e w e i g h t o f i n d i v i d u a l b i o m a s s c o m p o n e n t s o f e a c h t r e e b y t h e n u t r i e n t

c o n c e n t r a t i o n s f o r t h e s e c o m p o n e n t s m e a s u r e d o n a s a m p l e d t r e e o f t h e same

d . b . h . c l a s s .

F o r t h o s e t r e e s i n d . b . h . c l a s s e s f o r w h i c h n o t r e e w a s s a m p l e d ,

n u t r i e n t c o n c e n t r a t i o n d a t a f r o m t h e n e a r e s t s a m p l e d s i z e c l a s s w a s u s e d .

46

B y s u m m a t i o n , t h e t o t a l a b o v e - g r o u n d b i o m a s s n u t r i e n t c o n t e n t w a s o b t a i n e d

f o r t h e p l o t s , w i t h a b r e a k d o w n b y i n d i v i d u a l b i o m a s s c o m p o n e n t s .

No s i g n i f i c a n t r e g r e s s i o n e q u a t i o n s w e r e f o u n d f o r t h e b i o m a s s

o f s m a l l b r a n c h e s 0-~H i n c h d i a m e t e r ) f o r m o u n t a i n h e m l o c k , p o s s i b l y b e -

c a u s e o f t h e s m a l l n u m b e r o f s a m p l e d t r e e s ( 5 ) . T h e a m o u n t o f b i o m a s s a n d

t h e a m o u n t o f i n d i v i d u a l n u t r i e n t s i n p a r t i c u l a r d i a m e t e r c l a s s e s w e r e

t h e r e f o r e c a l c u l a t e d i n t h e f o l l o w i n g m a n n e r : c l a s s e s 1 a n d 2 w e r e c a l c u -

l a t e d w i t h c l a s s t h r e e ( T a b l e 1 1 ) u s i n g b i o m a s s a n d n u t r i e n t c o n t e n t d a t a

o f c l a s s t h r e e , B i o m a s s a n d n u t r i e n t c o n t e n t f o r c l a s s e s 4, 5 a n d 6 w e r e

o b t a i n e d b y m u l t i p l y i n g n u m b e r s o f t r e e s i n e a c h . c l a s s b y c o r r e s p o n d i n g

b i o m a s s a n d n u t r i e n t c o n t e n t o f t h e s a m p l e d t r e e f o r e a c h p a r t i c u l a r c l a s s .

C l a s s e s 8, 9, 1 0 a n d 1 1 w e r e c a l c u l a t e d w i t h c l a s s 7, u s i n g b i o m a s s a n d

n u t r i e n t c o n t e n t d a t a o f c l a s s 7.

I t i s o b v i o u s t h a t b i o m a s s a n d n u t r i e n t c o n t e n t w e r e o v e r - -

e s t i m a t e d f o r c l a s s e s 1 a n d 2 a n d w e r e u n d e r - e s t i m a t e d f o r c l a s s e s 8, 9, 1 0

a n d 1 1 , A l l t o g e t h e r , t h e b i o m a s s a n d n u t r i e n t c o n t e n t f o r s m a l l b r a n c h e s

o f m o u n t a i n h e m l o c k a r e p r o b a b l y s l i g h t l y u n d e r - e s t i m a t e d b e c a u s e a l t h o u g h

t h e n u m b e r o f t r e e s w h i c h w e r e u n d e r - e s t i m a t e d a n d o v e r - e s t i m a t e d i s a p -

p r o x i m a t e l y t h e s a me ( 1 0 a n d 9, r e s p e c t i v e l y ) , t h e u n d e r - e s t i m a t i o n w a s f o r

a m u c h , w i d e r c l a s s r a n g e t h a n t h e o v e r - e s t i m a t i o n .

No s i g n i f i c a n t r e g r e s s i o n e q u a t i o n s w e r e f o u n d f o r b i o m a s s o f

y e l l o w c e d a r . T h e n u m b e r o f s a m p l e d t r e e s ( 4 ) w a s n o t s u f f i c i e n t t o f i n d

s i g n i f i c a n t c o r r e l a t i o n s b e t w e e n m e n s u r a t i o n a l p a r a m e t e r s a n d t r e e b i o m a s s .

47

T h e a m o u n t o f b i o m a s s a n d t h e a m o u n t o f i n d i v i d u a l n u t r i e n t s i n p a r t i c u l a r

d i a m e t e r c l a s s e s w e r e , t h e r e f o r e , c a l c u l a t e d i n t h e f o l l o w i n g m a n n e r : f o r

c l a s s 1 ( T a b l e 1 1 ) t h e a v e r a g e a m o u n t o f b i o m a s s a n d n u t r i e n t s f o r p a r t i c u -

l a r t r e e c o m p o n e n t s w a s c a l c u l a t e d f r o m t w o s a m p l e d t r e e s f o r t h i s c l a s s .

A v e r a g e v a l u e s w e r e m u l t i p l i e d b y t h e n u m b e r o f t r e e s i n c l a s s 1. F o r

c l a s s 2 a v e r a g e a m o u n t o f b i o m a s s a n d n u t r i e n t f o r i n d i v i d u a l t r e e c o m p o n e n t s

w e r e c a l c u l a t e d f r o m t h e a v e r a g e o f t h e l a r g e s t t r e e i n s i z e c l a s s 1 a n d t h e

t r e e i n s i z e c l a s s 3. A v e r a g e v a l u e s w e r e a g a i n m u l t i p l i e d b y t h e n u m b e r o f

t r e e s i n c l a s s 2, F o r c l a s s 3 , b i o m a s s a n d n u t r i e n t v a l u e s f o r t h e s a m p l e

t r e e w e r e s i m p l y m u l t i p l i e d b y t h e n u m b e r o f t r e e s i n c l a s s 3, T h e same

wa s d o n e f o r c l a s s 5.

L i m i t a t i o n o n D a t a I n t e r p r e t a t i o n :

T h e n u m b e r o f o b s e r v a t i o n s u s e d f o r d e v e l o p i n g t h e r e g r e s s i o n

e q u a t i o n s w a s v e r y l i m i t e d ( 5 t o 1 2 o b s e r v a t i o n s p e r s p e c i e s ) d u e t o t h e

l a r g e s i z e o f many o f t h e t r e e s s a m p l e d a n d t h e s h o r t p e r i o d o f f i e l d w o r k

i m p o s e d b y s n o w c o n d i t i o n s . T h e r e g r e s s i o n e q u a t i o n s a r e , t h e r e f o r e , n o t

a s p r e c i s e a s t h e y w o u l d h a v e b e e n h a d i t b e e n p o s s i b l e t o s a m p l e a g r e a t e r

n u m b e r o f t r e e s ( 3 0 o r m o r e r e s i d u a l d e g r e e s o f f r e e d o m a r e u s u a l l y s u g -

g e s t e d t o o b t a i n a s t a b l e r e g r e s s i o n m o d e l ) . R e g r e s s i o n e q u a t i o n s w e r e

e x t r a p o l a t e d t o e s t i m a t e t h e b i o m a s s o f t h e f e w t r e e s w h i c h o c c u r r e d o u t s i d e

o f t h e r a n g e o f t r e e s i z e s t h a t w e r e s a m p l e d . I am a w a r e o f t h e d a n g e r s o f

u s i n g r e g r e s s i o n s t o e s t i m a t e v a l u e s o u t s i d e o f t h e r a n g e o f s a m p l i n g , b u t

b e c a u s e o f t h e l i m i t a t i o n s i m p o s e d b y t h e f i e l d p r o b l e m s t h e r e w a s n o o t h e r

4 8

r e c o u r s e . Due t o t h e c o m p l e x i t y o f b i o m a s s c a l c u l a t i o n a n d i t s c o m b i n a t i o n

w i t h n u t r i e n t c o n c e n t r a t i o n d a t a , t h e a m o u n t o f m a c r o n u t r i e n t s i m m o b i l i z e d

i n t r e e b i o m a s s p e r h e c t a r e s h o u l d b e c o n s i d e r e d m o r e a s a n a p p r o x i m a t i o n

t h a n a s a n e x a c t n u m b e r .

M i s s i n g D a t a a n d A d j u s t m e n t o f I n c o r r e c t D a t a :

D u r i n g t r a n s p o r t a t i o n f r o m t h e s a m p l e p l o t s t o t h e l a b o r a t o r y o n e

b i o m a s s s a m p l e a n d t h r e e c h e m i c a l a n a l y s i s ''samples w e r e l o s t . B e c a u s e o f

t h e n a t u r e o f t h e c o m p u t e r p r o g r a m f o r t h e c a l c u l a t i o n a d r y b i o m a s s a n d

n u t r i e n t c o n t e n t ( A p p e n d i c e s 10 a n d 1 1 ) a c o m p l e t e s e t o f s p e c i f i c g r a v i t y

d a t a a n d n u t r i e n t c o n c e n t r a t i o n d a t a w a s r e q u i r e d . T h e l o s t b i o m a s s s a m p l e

w a s d i s c #10, t r e e # 4 1 , S p e c i f i c g r a v i t y o f d i s c #9 o f t h e same t r e e w a s

u s e d t o r e p l a c e t h e m i s s i n g d a t a . T h e l o s t c h e m i c a l s a m p l e s w e r e a d i s c

f r o m a t r e e b o l e , a b r a n c h s a m p l e a n d a f o l i a g e s a m p l e . D a t a w e r e r e p l a c e d

b y c o r r e s p o n d i n g d a t a f r o m t h e s a m p l e t r e e o f t h e same s p e c i e s w i t h t h e m o s t

s i m i l a r b i o m a s s .

A p p r o x i m a t e l y 1 0 p e r c e n t o f t h e s p e c i f i c g r a v i t y d a t a w e r e o u t

o f t h e r a n g e c i t e d i n t h e l i t e r a t u r e f o r t h e s e p a r t i c u l a r s p e c i e s . Wood

s p e c i f i c g r a v i t y d a t a w i t h i n 9 5 p e r c e n t c o n f i d e n c e i n t e r v a l f o r P a c i f i c

w i l v e r f i r i s 0.36 ± 0 . 0 7 2 g / c m 3 , f o r m o u n t a i n h e m l o c k i t i s 0 , 4 1 ± 0 . 0 8 2

g/cm^ ( F o r e s t r y H a n d b o o k ) . D a t a w h i c h w e r e t h r e e t i m e s s t a n d a r d d e v i a t i o n

o u t o f t h i s r a n g e w e r e c o r r e c t e d t o t h e m e a n o f t h e v a l u e s f o r t h e t w o

n e i g h b o u r i n g d i s c s w i t h i n t h e same t r e e . T h i s m e t h o d w a s s e l e c t e d , e v e n

t h o u g h f r o m s t a t i s t i c a l s t a n d p o i n t i t i s n o t t h e m o s t a p p r o p r i a t e o n e

b e c a u s e t h e s p e c i f i c g r a v i t y d a t a d i d n o t s h o w a n y t r e n d a n d b e c a u s e

t h e s i m p l i c i t y o f t h e c o r r e c t i o n .

5 0

V R E S U L T S AND D I S C U S S I O N

B i o m a s s D 1 s t r i b u t i o n : ^

B o t h s t a n d s c a n b e c h a r a c t e r i z e d a s m a t u r e o r o v e r - m a t u r e ( m e a n

a g e : S q u a m i s h - 4 1 7 y e a r s ; H a n e y - 2 4 8 y e a r s ) a n d t h e r e f o r e t h e y a r e b o t h

i n a c o m p a r a b l e p h y s i o l o g i c a l s t a g e . T h e r e l a t i v e d i s t r i b u t i o n o f b i o m a s s

among d i f f e r e n t t r e e c o m p o n e n t s b e t w e e n b o t h s t a n d s s h o u l d t h e r e f o r e b e

m o r e a f u n c t i o n o f s i t e a n d e l e v a t i o n t h a n d i f f e r e n c e i n s t a n d a g e .

P e r h a p s t h e m o s t d r a m a t i c d i f f e r e n c e b e t w e e n t h e t w o p l o t s i s t h e

f a c t t h a t t h e 3 q u a m i s h s a m p l e p l o t h a s a b o u t e i g h t t i m e s a s m u c h a b o v e -

g r o u n d t r e e b i o m a s s p e r h e c t a r e a s t h e H a n e y p l o t . T h e d i s t r i b u t i o n o f

t h i s b i o m a s s b e t w e e n d i f f e r e n t c o m p o n e n t s a l s o v a r i e s b e t w e e n t h e t w o s i t e s

( T a b l e 1 7 ) . T h e p r o p o r t i o n o f b i o m a s s o n t h e H a n e y p l o t t o t h e b i o m a s s o n

t h e S q u a m i s h p l o t e x h i b i t s t h e r a t i o 1 t o 9.3 f o r s t e m w o o d , 1 t o 1 0 . 8 f o r

s t e m b a r k , 1 t o 6.6 f o r b i g b r a n c h e s , 1 t o 1.8 f o r s m a l l b r a n c h e s , 1 t o 6.4

f o r t w i g s a n d f o l i a g e , 1 t o 9.5 f o r t r e e b o l e s , 1 t o 4.7 f o r t r e e c r o w n s ,

a n d 1 t o 8.2 f o r t o t a l a b o v e - g r o u n d t r e e b i o m a s s . T h e g r e a t e s t d i f f e r e n c e

i s i n t h e b i o m a s s o f s t e m b a r k a n d s t e m w o o d , a n d t h e l e a s t i s i n t h e b i o m a s s

o f s m a l l b r a n c h e s . T h e b i o m a s s o f t r e e c r o w n s ( 1 : 4.7) s h o w s c o n s i d e r a b l y

l e s s d i f f e r e n c e t h a n t h e b i o m a s s o f t r e e b o l e s ( 1 : 9 . 5 ) .

T h e d i s t r i b u t i o n o f b i o m a s s among d i f f e r e n t t r e e c o m p o n e n t s i s

1 - Some o f t h e m a t e r i a l i n t h i s s e c t i o n w a s p r e s e n t e d a t t h e I U FRO m e e t i n g i n V a n c o u v e r , B. C. i n A u g u s t , 1 9 7 3 .

T A B L E 1 7 : B i o m a s s d i s t r i b u t i o n f o r f o u r t r e e s p e c i e s i M o u n t a i n h e m l o c k = 7 4 ; P a c i f i c s i l v e r f i r =

T r e e S p e c i e s Wood B a r k B i g

B r a n c h S m a l l B r a n c h

M o u n t a i n h e m l o c k 9 1 3 7 8 2 7 8 2 6 1 0 4 8 6 2 2 7 1

P a c i f i c s i l v e r f i r 2 2 7 5 6 1 5 0 1 6 7 2 5 0 8 4 8 1 7 2

W e s t e r n h e m l o c k 2 7 3 4 8 6 5 9 5 4 6 6 8 5 2 7 3

Y e l l o w c e d a r 6 9 3 4 5 9 8 6 7 5 4 1 1

S q u a m i s h P l o t T o t a l 3 1 8 9 3 9 7 7 9 9 3 3 5 5 7 0 1 0 4 4 3

H a n e y P l o t T o t a l 3 4 2 8 2 7 1 9 3 5 3 4 3 5 6 8 4

n t h e S q u a m i s h a n d H a n e y s a m p l e p l o t s ( i n k g / h a ) . n 1 8 2 ; W e s t e r n h e m l o c k = 6 7 ; Y e l l o w c e d a r = 1 5 .

T w i g F o l i a g e T w i g & F o l .

T r e e B o l e C r o w n

W h o l e T r e e

3 3 2 7 3 3 4 5 5 9 0 5 1 1 9 2 0 3 1 8 6 6 2 1 3 7 8 6 5

— — 1 8 4 5 5 2 7 7 7 2 8 5 1 7 1 1 3 2 9 4 4 0

1 4 5 4 1 9 7 5 3 0 8 4 3 3 9 4 4 1 3 0 2 5 4 6 9 7 0

— — 7 1 3 7 5 2 3 1 7 9 9 9 3 2 2

— — 2 4 3 6 0 3 9 6 9 3 1 7 0 3 7 3 4 6 7 3 0 5

3 7 9 7 4 1 4 6 7 1 4 8 2 4 5 6 2 9 2

5 2

r a t h e r s i m i l a r f o r a l l f o u r s p e c i e s w i t h t h e e x c e p t i o n o f b i o m a s s o f s m a l l

b r a n c h e s ( T a b l e 18)_, T h e p r o p o r t i o n o f s t e m w o o d v a r i e s f r o m 6 0 , 7 p e r c e n t

o f t o t a l a b o v e - g r o u n d t r e e b i o m a s s f o r m o u n t a i n h e m l o c k t o 7 1 . 8 p e r c e n t

f o r y e l l o w c e d a r . T h e p r o p o r t i o n o f b a r k v a r i e s f r o m 6,4 p e r c e n t f o r

y e l l o w c e d a r t o 1 7 . 9 p e r c e n t f o r m o u n t a i n h e m l o c k . T h e p r o p o r t i o n o f

s m a l l b r a n c h e s i s a b o u t s i x t i m e s g r e a t e r f o r w e s t e r n h e m l o c k ( 1 1 . 2 p e r c e n t )

t h a n f o r m o u n t a i n h e m l o c k ( 1 . 9 p e r c e n t ) . T h e r e m a i n i n g t w o s p e c i e s h a v e

i n t e r m e d i a t e p r o p o r t i o n s o f s m a l l b r a n c h e s . T r e e b o l e a c c o u n t s f o r 74 t o

85 p e r c e n t o f t h e w h o l e t r e e b i o m a s s ; t r e e c r o w n f o r 1 5 t o 26 p e r c e n t .

T h e b i o m a s s o f p a r t i c u l a r t r e e c o m p o n e n t s a s w e l l a s t o t a l a b o v e -

g r o u n d t r e e b i o m a s s c a n b e d e s c r i b e d b y s i m p l e l o g a r i t h m i c r e g r e s s i o n e q u a -

t i o n s o r s i m p l e l i n e a r r e g r e s s i o n e q u a t i o n s u s i n g m e n s u r a t i o n a l t r e e p a r a -

m e t e r s a s s i m p l e ( e . g . D) o r c o m b i n e d ( e . g . D 2 . H) i n d e p e n d e n t v a r i a b l e s

( A p p e n d i x 3 ) . I n t h i s s t u d y l o g a r i t h m i c r e g r e s s i o n e q u a t i o n s w i t h t h e

h i g h e s t c o e f f i c i e n t o f d e t e r m i n a t i o n ( r 2 . 1 0 0 ) w e r e u s e d f o r p r e d i c t i o n o f

b i o m a s s f o r a l l t r e e s o n t h e s a m p l e p l o t s . I n t h e f o l l o w i n g d i s c u s s i o n

b o t h l o g a r i t h m i c a n d l i n e a r e q u a t i o n s a r e d i s c u s s e d , b u t t h e y w e r e n o t s t a -

t i s t i c a l l y c o m p a r e d . T h e d e c i s i o n t o u s e l o g a r i t h m i c e q u a t i o n s w a s made

b e c a u s e o f t h e i r b e t t e r f i t f o r v e r y s m a l l a n d v e r y b i g t r e e s .

W h o l e t r e e b i o m a s s a n d w o o d b i o m a s s : T h e b e s t e s t i m a t o r o f t h e

o

t r e e b i o m a s s a n d w o o d b i o m a s s i s t h e c o m b i n e d v a r i a b l e D ,H o r D a l o n e , i n

b o t h t h e l i n e a r a n d l o g a r i t h m i c e q u a t i o n .

T A B L E 1 8 : A v e r a g e b i o m a s s p r o p o r t i o n s o f d i f f e r e n t t r e e c o m p o n e n t s f o r f o u r t r e e s p e c i e s o n S q u a m i s h a n d H a n e y s a m p l e p l o t s ( i n p e r c e n t ) . n ; M o u n t a i n h e m l o c k = 5 ; P a c i f i c s i l v e r f i r = 7; W e s t e r n h e m l o c k = 8; Y e l l o w c e d a r = 4.

S q u a m i s h S q u a m i s h H a n e y H a n e y

T r e e C o m p o n e n t P a c i f i c s i l v e r f i r M o u n t a i n h e m l o c k W e s t e r n h e m l o c k Y e l l o w c e d a r

S t e m w o o d R a n g e 5 9 . 6 - 7 0 . 5 6 4 . 1 - 6 9 . 2 5 4 . 8 - 6 8 . 1 6 3 . 5 - 7 7 . 3 M e a n 6 6 . 3 6 7 . 2 6 0 . 8 7 1 . 7

S t e m b a r k R a n g e 9 . 5 - 1 8 . 8 1 7 . 0 - 1 9 . 1 9 . 8 - 1 6 . 4 5 . 4 - 8 . 2 M e a n 1 3 . 9 1 7 . 9 1 3 . 3 6.4

B i g b r a n c h e s R a n g e 4 . 4 - 1 2 . 4 5 . 2 - 9 . 6 0 . 6 - 1 3 . 2 2 . 3 - 9 . 7 M e a n 8.1 6.9 6.6 5.9

S m a l l b r a n c h e s R a n g e 1 . 6 ^ 6 . 5 1.1-2.4 7 . 0 - 1 5 . 9 3 . 4 - 1 0 . 4 M e a n 3.9 1.9 1 1 . 3 6.2

T w i g s R a n g e 2 . 2 - 3 . 2 1 . 7 - 6 . 5 M e a n 2.5 3.3

F o l i a g e R a n g e 2 . 9 - 4 . 5 2 . 7 - 7 . 0 M e a n 3.6 4.7

T w i g & F o l i a g e R a n g e 4 . 5 - 1 1 . 9 5 . 1 - 7 . 6 5 . 9 - 1 0 . 0 6 . 2 - 1 6 . 4 M e a n 7.8 6.1 8.0 9.8

B o l e s M e a n 8 0 . 2 8 5 . 1 7 4 . 1 7 8 . 1

C r o w n s M e a n 1 9 . 8 1 4 . 9 2 5 . 9 2 1 . 9

54

B a r k b i o m a s s ; B a r k b i o m a s s i s m a i n l y d e p e n d e n t o n t h e b o l e

2

d i a m e t e r a n d t h e r e f o r e B A , D o r D ,H a r e t h e b e s t i n d e p e n d e n t v a r i a b l e s

i n b o t h t h e l i n e a r a n d l o g a r i t h m i c e q u a t i o n s .

B i g b r a n c h e s b i o m a s s : On t h e S q u a m i s h p l o t t h e b i o m a s s o f b i g

b r a n c h e s i s d e p e n d e n t o n t r e e d i a m e t e r , t r e e l e n g t h , c r o w n d i a m e t e r a n d i n

t h e l o g a r i t h m i c e q u a t i o n , o n c r o w n l e n g t h . I n b o t h t h e l i n e a r a n d l o g a r -

i t h m i c e q u a t i o n f o r h e m l o c k , t h e m o s t i m p o r t a n t i n d e p e n d e n t v a r i a b l e i s

D2.H,CW, w h i l e f o r P a c i f i c s i l v e r f i r I t i s D.CL i n t h e l o g a r i t h m i c e q u a -

t i o n , a n d BA o r D i n t h e l i n e a r e q u a t i o n . On t h e H a n e y p l o t , t h e b e s t

i n d e p e n d e n t v a r i a b l e f o r h e m l o c k i n t h e l i n e a r e q u a t i o n i s D.CL, w h i l e i n

t h e l o g a r i t h m i c e q u a t i o n i t i s D a l o n e .

S m a l l b r a n c h e s b i o m a s s : B i o m a s s o f s m a l l b r a n c h e s f o r h e m l o c k o n

t h e S q u a m i s h p l o t d o e s n o t s h o w a n y r e l a t i o n s h i p w i t h t h e i n d e p e n d e n t v a r i -

a b l e s t e s t e d i n e i t h e r t h e l i n e a r o r t h e l o g a r i t h m i c e q u a t i o n s . T h e b i o m a s s

o f s m a l l b r a n c h e s f o r P a c i f i c s i l v e r f i r o n t h e S q u a m i s h p l o t i s b e s t d e -

s c r i b e d i n t h e l i n e a r e q u a t i o n b y e i t h e r t r e e l e n g t h H o r t r e e d i a m e t e r D,

o r b y t h e c o m b i n a t i o n o f t r e e d i a m e t e r a n d c r o w n l e n g t h (D + C L ) . I n t h e

l o g a r i t h m i c e q u a t i o n b i o m a s s o f s m a l l b r a n c h e s i s b e s t d e s c r i b e d b y t h e l o g -

a r i t h m o f t r e e d i a m e t e r a n d c r o w n l e n g t h b y e i t h e r l o g D.CL o r l o g D . C L 2 .

B i o m a s s o f s m a l l b r a n c h e s f o r h e m l o c k o n t h e H a n e y p l o t i s b e s t d e s c r i b e d

i n t h e l i n e a r e q u a t i o n b y t r e e l e n g t h ( H ) a l o n e o r b y t h e c o m b i n a t i o n o f

d . b . h . a n d c r o w n l e n g t h (D + C L ) . I n t h e l o g a r i t h m i c e q u a t i o n s t h e m o s t

s i g n i f i c a n t i n d e p e n d e n t v a r i a b l e i s t h e c o m b i n a t i o n o f d . b . h , , t r e e l e n g t h

5 5

a n d c r o w n w i d t h (D Z .H,CW)>

T w i g s a n d f o l i a g e - b i o m a s s : T h e b i o m a s s o f t w i g s a n d f o l i a g e f o r

b o t h s p e c i e s i n S q u a m i s h i s b e s t d e s c r i b e d i n t h e l i n e a r e q u a t i o n b y t r e e

d i a m e t e r i n c o m b i n a t i o n w i t h c r o w n l e n g t h ( D . C L ) o r b y t r e e d i a m e t e r ( D )

a l o n e . I n t h e l o g a r i t h m i c e q u a t i o n , t h e m o s t s i g n i f i c a n t i n d e p e n d e n t v a r -

i a b l e i s t h e c o m b i n a t i o n o f t r e e d i a m e t e r w i t h t r e e l e n g t h ( D 2 , H ) f o r

m o u n t a i n h e m l o c k , a n d t r e e d i a m e t e r w i t h c r o w n l e n g t h ( D . C L ) f o r P a c i f i c

s i l v e r f i r . F o r h e m l o c k i n H a n e y , t r e e l e n g t h a l o n e ( H ) o r t h e c o m b i n a t i o n

o f t r e e d i a m e t e r a n d c r o w n l e n g t h ( D . C L ) a r e t h e b e s t i n d e p e n d e n t v a r i a b l e s

i n l i n e a r e q u a t i o n s . I n t h e l o g a r i t h m i c e q u a t i o n , t h e c o m b i n a t i o n o f t r e e

d i a m e t e r w i t h c r o w n l e n g t h ( D . C L ) i s t h e m o s t s i g n i f i c a n t i n d e p e n d e n t v a r i -

a b l e .

T w i g b i o m a s s ' • F o r m o u n t a i n h e m l o c k o n t h e S q u a m i s h p l o t , t h e

c o m b i n a t i o n o f t r e e d i a m e t e r a n d c r o w n l e n g t h ( D . C L ) o r t r e e d i a m e t e r

a l o n e (D) i s t h e b e s t i n d e p e n d e n t v a r i a b l e i n t h e l i n e a r e q u a t i o n . I n t h e

l o g a r i t h m i c e q u a t i o n , t r e e d i a m e t e r w i t h c r o w n l e n g t h o r t r e e d i a m e t e r w i t h

c r o w n l e n g t h s q u a r e d ( D . C L o r D . C L 2 ) i s t h e b e s t i n d e p e n d e n t v a r i a b l e . F o r

h e m l o c k o n t h e H a n e y p l o t , c r o w n w i d t h s q u a r e d ( C W 2 ) i s t h e b e s t i n d e p e n -

d e n t v a r i a b l e i n t h e l i n e a r e q u a t i o n , w h i l e t h e c o m b i n a t i o n o f t r e e b a s a l

a r e a a n d t r e e l e n g t h w i t h c r o w n d i a m e t e r a n d l e n g t h ( B A . H ) + (CL.CW)

i s t h e b e s t i n d e p e n d e n t v a r i a b l e i n t h e l o g a r i t h m i c e q u a t i o n .

F o l i a g e b i o m a s s : F o r m o u n t a i n h e m l o c k o n t h e S q u a m i s h p l o t e i t h e r

56

t h e c o m b i n a t i o n o f t r e e d i a m e t e r w i t h c r o w n l e n g t h ( D . C L ) o r t r e e d i a m e t e r

a l o n e i s t h e b e s t i n d e p e n d e n t v a r i a b l e i n t h e l i n e a r e q u a t i o n . I n t h e

l o g a r i t h m i c e q u a t i o n , t r e e d i a m e t e r w i t h t r e e l e n g t h (D ,H) a p p e a r s t o b e

t h e m o s t s i g n i f i c a n t i n d e p e n d e n t v a r i a b l e . F o r h e m l o c k o n t h e H a n e y p l o t ,

t r e e l e n g t h a l o n e (II) o r t r e e d i a m e t e r w i t h c r o w n l e n g t h ( D . C L ) a r e t h e

m o s t s i g n i f i c a n t i n d e p e n d e n t v a r i a b l e s i n t h e l i n e a r r e g r e s s i o n e q u a t i o n .

I n t h e l o g a r t h m i c e q u a t i o n t r e e d i a m e t e r i n c o m b i n a t i o n w i t h c r o w n l e n g t h

( D . C L ) i s t h e b e s t i n d e p e n d e n t v a r i a b l e .

I n s u m m a r y , f r o m t h e a b o v e d i s c u s s i o n , i t w o u l d a p p e a r t h a t t h e

e q u a t i o n

l o g Y = b + b1 l o g D

i s a m o d e l w h i c h c a n b e u s e d f o r p r e d i c t i o n o f t h e w h o l e t r e e b i o m a s s a s

w e l l a s t h e b i o m a s s o f a n y i n d i v i d u a l t r e e c o m p o n e n t . T h i s i s i n a g r e e m e n t

w i t h a n u m b e r o f o t h e r p u b l i c a t i o n s l i s t e d b y M a d g w i c k ( 1 9 6 3 ) . T h e m o d e l

f i t s w e l l f o r b o t h s p e c i e s o n t h e S q u a m i s h p l o t ( w i t h t h e e x c e p t i o n o f

s m a l l b r a n c h e s f o r m o u n t a i n h e m l o c k ) , a n d f o r h e m l o c k o n t h e H a n e y p l o t .

A l o g a r i t h m i c e q u a t i o n i n t h e f o r m

l o g ( w e i g h t ) = a + b ( l o g D )

w a s u s e d b y M o r r i s o n ( 1 9 7 2 ) . f o r j a c k p i n e s t a n d s i n N o r t h e r n O n t a r i o . On

t h e o t h e r h a n d , K u r u c z ( 1 9 6 9 ) . u s e d l i n e a r r e g r e s s i o n e q u a t i o n s f o r b i o m a s s

p r e d i c t i o n o f D o u g l a s - f i r , w e s t e r n h e m l o c k a n d w e s t e r n r e d c e d a r i n U.B.C.

r e s e a r c h f o r e s t i n H a n e y .

57

N u t r i e n t D i s t r l h u t l d i K W i t h i n - t h e - T r e e L a y e r ;

A v e r a g e n u t r i e n t c o n c e n t r a t i o n s f o r p a r t i c u l a r t r e e c o m p o n e n t s

f o r t h e f o u r t r e e s p e c i e s a r e s u m m a r i z e d i n T a b l e s 1 9 , 2 0 , 2 1 a n d 2 2 .

N u t r i e n t c o n c e n t r a t i o n s f o r i n d i v i d u a l s a m p l e t r e e s a r e p r e s e n t e d i n A p p e n -

d i c e s 6 a n d 7. I w o u l d l i k e t o d i s c u s s f i r s t t h e c o n c e n t r a t i o n o f s p e c i f i c

e l e m e n t s i n p a r t i c u l a r t r e e c o m p o n e n t s f o r e a c h t r e e s p e c i e s s e p a r a t e l y

a n d t h e n c o m p a r e n u t r i e n t c o n c e n t r a t i o n s b e t w e e n s p e c i e s .

C o n c e n t r a t i o n s o f n u t r i e n t s w i t h i n d i f f e r e n t b i o m a s s c o m p o n e n t s

o f P a c i f i c s i l v e r f i r w e r e f o u n d t o b e i n t h e s e q u e n c e stemwood<C b i g

b r a n c h e s <C s m a l l b r a n c h e s <C stembark<C t w i g s a n d f o l i a g e f o r a l l n u t r i e n t s

w i t h t h e e x c e p t i o n o f c a l c i u m . C o n c e n t r a t i o n o f c a l c i u m i s h i g h e r i n b a r k

t h a n i n t w i g s a n d f o l i a g e . N u t r i e n t c o n c e n t r a t i o n s i n p a r t i c u l a r t r e e c o m -

p o n e n t s w e r e f o u n d t o b e i n t h e s e q u e n c e :

w o o d C a ^ N £ = K > M g > P

b a r k C a > N > K > P > M g

b i g b r a n c h e s C a > N > K > - Mg £z P

s m a l l b r a n c h e s C a > N > K > M g « r P

t w i g s a n d f o l i a g e N ;> C a >• K >• P >• Mg

c o n e s N > K > P > M g > C a

N u t r i e n t c o n c e n t r a t i o n s w i t h i n m o u n t a i n h e m l o c k w e r e f o u n d t o b e

i n t h e s e q u e n c e : s t e m w o o d < ^ b i g b r a n c h e s <^ s m a l l b r a n c h e s <^ s t e m b a r k < ^

twigs<C f o l i a g e f o r a l l n u t r i e n t s e x c e p t p h o s p h o r o u s a n d c a l c i u m . T h e

T A B L E 1 9 : M e a n c o n c e n t r a t i o n ( p e r c e n t a g e o v e n d r y w e i g h t ) . , o n N, P, K, C a a n d Mg i n c o m p o n e n t s a n d s u b - c o m p o n e n t s o f P a c i f i c s i l v e r f i r s a m p l e t r e e s .

C o m p o n e n t N P K C a Mg

Wood 1 0.05 0 , 0 1 0.06 0,05 0 , 0 1 ( d i s c n o , f r o m 3 0.04 0 . 0 1 0.05 0.07 0 . 0 1 t h e b o t t o m ) 5 0.05 0 . 0 1 0.06 0,06 0 . 0 1

7 0,05 0 . 0 1 0.06 0,06 0 . 0 1 9 0.06 0 . 0 1 0.07 0.06 0 . 0 1

1 0 0.07 0.02 0.07 0.05 0 . 0 1

B a r k 1 0.25 0.04 0 . 1 1 0.49 0.02 ( d i s c n o , f r o m 3 0 . 2 1 0.04 0.12 0.48 0.03 t h e b o t t o m ) 5 0.22 0.04 0.13 0.47 0.03

7 0.24 0.05 0.14 0.46 0.04 9 0 . 3 1 0.06 0.18 , 0.50 0.04

1 0 0.40 0.08 0.24 0.49 0.04

B i g b r a n c h e s 0.15 0.02 0.09 0 . 3 1 0.02

S m a l l b r a n c h e s 0.22 0.03 0.12 0.29 0.03

T w i g s & F o l i a g e 1 0.73 0.10 0.36 0.40 0.08 ( u p p e r , m i d d l e , 2 0.72 0.10 0 . 3 1 0.46 0.07 l o w e r c r o w n ) 3 0.76 0.10 0.32 0.46 0.06

C o n e s 1.35 0.24 0.70 0.04 ' 0.08

D a t a f o r w o o d a n d b a r k b a s e d o n 1 2 s a m p l e d t r e e s . D a t a f o r c r o w n c o m p o n e n t s b a s e d o n 7 s a m p l e d t r e e s . D a t a f o r c o n e s b a s e d o n 6 s a m p l e d t r e e s .

T A B L E 2 0 : M e a n c o n c e n t r a t i o n ( p e r c e n t a g e o v e n d r y w e i g h t ) , o f N, P, K, a n d Mg i n c o m p o n e n t s a n d s u b - c o m p o n e n t s o f m o u n t a i n h e m l o c k s a m p l e t r e e s .

C o m p o n e n t N P K. C a Mg

Wood 1 0,06 0.02 0.07 0,06 0,02 ( d i s c n o , f r o m 3 0.05 0.01 0.06 0.07 0.02 t h e b o t t o m ) 5 0,05 0.01 0,06 0,07 0.02

7 0.05 0,02 0.07 0.08 0.02 9 0.06 0.02 0.07 0.06 0.02

10 0.07 0.02 0.07 o;o6 0.02

B a r k 1 0.22 0.07 0.09 0.40 0.02 ( d i s c n o , f r o m 3 0.15 0.06 0.09 0.31 0.02 t h e b o t t o m ) 5 0.17 0.05 0.09 0.31 0.02

7 0.17 0.05 0.09 0.24 0.02 9 0.22 0.06 0.12 0.25 0.02

1 0 0.28 0.07 0.16 0.27 0.03

B i g b r a n c h e s 0.12 0.03 0.07 0.16 0.02

S m a l l b r a n c h e s 0.17 0.04 0.09 0.18 0.03

T w i g s 1 0.43 0.06 0.16 0.19 0.04 ( u p p e r , m i d d l e , 2 0.39 0.06 0.15 0.18 0.04 l o w e r c r o w n ) 3 0.34 0.06 0.17 0.17 0.05

F o l i a g e 1 0.80 0.11 0.28 0.31 0.08 ( u p p e r , m i d d l e , 2 0.91 0.12 0 . 3 1 0.28 0.08 l o w e r c r o w n ) 3 0.83 0.11 0.31 0.32 0.08

C o n e s 0.92 0.17 0.32 0.03 0.05

D a t a f o r w o o d a n d b a r k b a s e d o n 8 s a m p l e d t r e e s . D a t a f o r c r o w n c o m p o n e n t s b a s e d o n 5 s a m p l e d t r e e s . D a t a f o r c o n e s b a s e d o n 1 s a m p l e d t r e e .

6 0

T A B L E 2 1 : M e a n c o n c e n t r a t i o n ( p e r c e n t a g e o v e n d r y w e i g h t ) , o f N , P, K, C a a n d Mg i n c o m p o n e n t s a n d s u b - c o m p o n e n t s o f w e s t e r n h e m l o c k s a m p l e t r e e s .


Wood 1 0.04 0.02 0.07 0.05 0 . 0 1 ( d i s c n o . f r o m 3 0.04 0 . 0 1 0.07 0.06 0 . 0 1 t h e b o t t o m ) 5 0,04 0 . 0 1 0.06 0.05 0 . 0 1

7 0.05 0 . 0 1 0.06 0.05 0 . 0 1 9 0.06 0.01 0.07 0.04 0 . 0 1

1 0 0.06 0 . 0 1 0.07 0.04 0 . 0 1

B a r k 1 0.15 0.03 0.08 0.40 0.02 ( d i s c n o . f r o m 3 0.16 0.03 0.12 0.44 0.02 t h e b o t t o m ) 5 0.18 0.03 0.14 ,0.38 0.02

7 0.22 0.04 0.16 0.30 0.03 9 0.27 0.05 0.19 0.29 0.03

1 0 0.32 0.06 0.22 0.29 0.04

B i g b r a n c h e s 0.10 0,02 OgOS 0.17 0.02

S m a l l b r a n c h e s 0.14 0.02 0.09 . 0.19 0.02

T w i g s 1 0.53 0.08 0 . 2 1 0.29 0.06 ( u p p e r , m i d d l e , 2 0.45 0.07 0.19 0.29 0.05 l o w e r c r o w n ) 3 0.40 0.06 0.17 0.28 0.05

F o l i a g e 1 0.76 0 . 1 1 0.37 0.39 0.10 ( u p p e r , m i d d l e , 2 0.74 0.12 0.33 0.48 0.12 l o w e r c r o w n ) 3 0.74 0.14 0.34 0.49. 0.12

A l l d a t a b a s e d o n 8 s a m p l e d t r e e s .

61

T A B L E 2 2 : M e a n c o n c e n t r a t i o n ( p e r c e n t a g e o v e n d r y w e i g h t ) , o f N, P, K, C a a n d Mg i n c o m p o n e n t s a n d s u b - c o m p o n e n t s o f y e l l o w c e d a r s a m p l e t r e e s .


Wood 1 0,05 0 . 0 1 0.05 0.12 0 . 0 1 ( d i s c n o . f r o m 3 0,06 0 . 0 1 0,05 0.12 0 . 0 1 t h e b o t t o m ) . 5 0.05 0 . 0 1 0.05 .0.11 0 . 0 1

7 0.07 0 . 0 1 0.05 0.09 0 . 0 1 9 0.07 0 . 0 1 0.06 0 . 1 1 0 . 0 1

1 0 0,07 0 . 0 1 0.06 0.12 0 . 0 1

B a r k 1 0.27 0.05 0.20 0.99 0.04 ( d i s c n o . f r o m 3 0.27 0,05 0.18 0.40 0.04 t h e b o t t o m ) 5 0.27 0.06 0.25 0.49 0.05

7 0.33 0.07 0.27 0 . 1 1 0.05 9 0.37 0.08 0.30 0.87 0.06

1 0 0,44 0.08 0.30 0.62 0.06

B i g b r a n c h e s 0.15 0.02 0.09 0.64 0.02

S m a l l b r a n c h e s 0.23 0.03 0.14 0 . 8 1 0.03

T w i g s & F o l i a g e 1 0.75 0.09 0.37 0.24 0.07 ( u p p e r , m i d d l e , 2 0.70 0.08 0.32 0.33 0.07 l o w e r c r o w n ) 3 0.77 0.08 0.36 0 . 5 3 0.08

A l l d a t a b a s e d o n 4 s a m p l e d t r e e s .

62

c o n c e n t r a t i o n o f P i n b a r k a n d t w i g s i s a p p r o x i m a t e l y t h e same ( t h e m a i n

c o m p o n e n t o f t w i g s i s b a r k ) . , T h e c o n c e n t r a t i o n o f C a i s h i g h e r i n b a r k

t h a n i n t w i g s a n d b a r k c o n c e n t r a t i o n i s a p p r o x i m a t e l y e q u a l t o f o l i a g e c o n -

c e n t r a t i o n . N u t r i e n t c o n c e n t r a t i o n s i n p a r t i c u l a r t r e e c o m p o n e n t s a p p e a r

t o b e i n t h e s e q u e n c e :

w o o d C a « s N « ? K > P * = : M g

b a r k C a > N > K > P > M g

b i g b r a n c h e s C a > N > K > P « M g

s m a l l b r a n c h e s C a < » N > K > P > M g

t w i g s N > C a > K > P > Mg

f o l i a g e N > C a > K > P > M g

c o n e s N > K > P > Mg > C a

N u t r i e n t c o n c e n t r a t i o n s w i t h i n w e s t e r n h e m l o c k w e r e f o u n d t o b e i n

t h e s e q u e n c e : s t e m w o o d < b i g b r a n c h e s <C s m a l l b r a n c h e s < s t e m b a r k < t w i g s <

f o l i a g e f o r a l l n u t r i e n t s w i t h t h e e x c e p t i o n o f c a l c i u m . T h e c o n c e n t r a t i o n

o f c a l c i u m i s h i g h e r i n t h e b a r k t h a n i n t h e t w i g s . N u t r i e n t c o n c e n t r a t i o n s

i n p a r t i c u l a r t r e e c o m p o n e n t s w e r e f o u n d t o b e i n t h e s e q u e n c e :

w o o d K > C a ~ N > P ? » M g

b a r k C a > N > K > P > Mg

b i g b r a n c h e s C a > N > K > P «2Mg

s m a l l b r a n c h e s C a > N > K > P ^ M g

t w i g s N > C a > K > P > Mg

f o l i a g e N > C a > K > P > M g

63

N u t r i e n t c o n c e n t r a t i o n s i n y e l l o w c e d a r w e r e f o u n d t o b e i n t h e

s e q u e n c e : s t e m w o o d < f b i g b r a n c h e s < f s m a l l branches<C b a r k < ^ t w i g s a n d f o l -

i a g e f o r a l l n u t r i e n t s w i t h t h e e x c e p t i o n o f c a l c i u m . T h e c o n c e n t r a t i o n o f

c a l c i u m i s h i g h e r i n b a r k t h a n i n t w i g s a n d f o l i a g e , N u t r i e n t c o n c e n t r a -

t i o n s i n p a r t i c u l a r t r e e c o m p o n e n t s a p p e a r t o b e i n t h e o r d e r :

w o o d C a > N > K > P <srMg

b a r k C a > N > K > P > Mg

b i g b r a n c h e s C a > N > K > P ^ M g

s m a l l b r a n c h e s C a > N > K > P Mg

t w i g s a n d f o l i a g e C a ^> N > K > P 5 » M g

Y e l l o w c e d a r h a s c o n s i d e r a b l y h i g h e r a m o u n t s o f c a l c i u m i n a l l

t r e e c o m p o n e n t s t h a n t h e o t h e r s p e c i e s .

C o n c e n t r a t i o n s o f n u t r i e n t s i n s t e m w o o d do n o t s h o w a n y s i g n i f i -

c a n t t r e n d b e t w e e n d i f f e r e n t d i s c s f r o m t h e b o t t o m t o t h e t o p o f t h e t r e e .

C o n c e n t r a t i o n s o f n u t r i e n t s i n s t e m b a r k , h o w e v e r , d o s h o w a n i n c r e a s i n g t r e n d

f r o m b o t t o m t o t o p o f t h e t r e e f o r a l l n u t r i e n t s e x c e p t c a l c i u m . C o n c e n t r a -

t i o n o f c a l c i u m i n b a r k o f P a c i f i c s i l v e r f i r i s c o n s t a n t , w h i l e f o r t h e

o t h e r t r e e s p e c i e s i t d e c r e a s e s f r o m t h e b o t t o m t o t h e t o p o f a t r e e .

T h e f o u r t r e e s p e c i e s c a n b e a r r a n g e d i n d i f f e r e n t o r d e r a c c o r d i n g

t o t h e n u t r i e n t c o n c e n t r a t i o n s i n v a r i o u s t r e e c o m p o n e n t s . A l l s p e c i e s a r e

a p p r o x i m a t e l y e q u a l i n c o n c e n t r a t i o n o f a l l n u t r i e n t s i n s t e m w o o d , w i t h t h e

e x c e p t i o n o f y e l l o w c e d a r w h i c h e x h i b i t s c o n s i d e r a b l y h i g h e r c a l c i u m c o n c e n -

t r a t i o n . T h e f o u r s p e c i e s a p p e a r i n t h e f o l l o w i n g o r d e r a c c o r d i n g t o c o n -

c e n t r a t i o n s o f d i f f e r e n t e l e m e n t s i n v a r i o u s t r e e c o m p o n e n t s .

64

B a r k N Y . c . > P . s . f . > W.h. > M.h. P • Y , c , « P . s . f . ;= M,h. > W,h. K Y , c , > P . s . f . « = W.h, > M , h .

C a a n d Mg Y . c . > P . s . f , > W.h, > M.h.

B i g N Y . c . ^ P . s . f . > M . h . > W,h, B r a n c h e s C a Y . c , > P . s . f , > W.h. M.h.

P, K, Mg Y , c . s = P . s . f . « W.h. « H , h (

S m a l l N Y . c , » P . s . f . > W . h , M , h , B r a n c h e s P, Mg Y , c . a s - P . s . f . ~ W.h. zz. M.h.

K, C a Y . c , > P , s . f , > W.h, <=sM,h,

T w i g s a n d f o l i a g e c a n n o t b e c o m p a r e d b e c a u s e o f t h e n a t u r e o f

s a m p l i n g a n d a n a l y s e s . F o r b o t h h e m l o c k s p e c i e s t w i g s a n d f o l i a g e w e r e

s a m p l e d a n d a n a l y s e d s e p a r a t e l y , w h i l e f o r P a c i f i c s i l v e r f i r a n d y e l l o w

c e d a r t w i g s a n d f o l i a g e w e r e n o t s e p a r a t e d a n d w e r e a n a l y s e d t o g e t h e r .

I n s u m m a r i z i n g t h e d a t a o n n u t r i e n t d i s t r i b u t i o n i n t h e a b o v e - g r o u n d

t r e e b i o m a s s , t h e f o l l o w i n g p o i n t s c a n b e m a d e : a ) y e l l o w c e d a r e x h i b i t s c o n -

s i d e r a b l y h i g h e r c a l c i u m c o n c e n t r a t i o n s i n a l l t r e e c o m p o n e n t s t h a n o t h e r

s p e c i e s : b ) P a c i f i c s i l v e r f i r e x h i b i t s h i g h e r n u t r i e n t c o n c e n t r a t i o n s f o r a l -

m o s t a l l e l e m e n t s i n a l l t r e e c o m p o n e n t s t h a n b o t h h e m l o c k s p e c i e s ; c ) n u t r i -

e n t c o n c e n t r a t i o n s o f m o u n t a i n h e m l o c k a r e a p p r o x i m a t e l y e q u a l t o t h e c o n c e n -

t r a t i o n s o f w e s t e r n h e m l o c k , One m u s t k e e p i n m i n d t h a t t h e s i t e c o n d i t i o n s

o f t h e t w o s a m p l e p l o t s w e r e c o n s i d e r a b l y d i f f e r e n t . T r e e s p e c i e s m i g h t a p -

p e a r i n a d i f f e r e n t o r d e r i f t h e y w e r e g r o w n u n d e r u n i f o r m s i t e c o n d i t i o n s .

C o n c e n t r a t i o n o f m a c r o n u t r i e n t s i n f o l i a g e o r f o l i a g e a n d t w i g s

w i t h i n t h e u p p e r t h i r d o f t r e e c r o w n s i s s u m m a r i z e d i n T a b l e 2 3 , T h e d a t a

c a n n o t b e c o m p a r e d w i t h f o l i a g e c o n c e n t r a t i o n s f r o m o t h e r p u b l i c a t i o n s

b e c a u s e , a ) , t h e s a m p l e s w e r e o b t a i n e d r a n d o m l y f r o m a n y l o c a t i o n w i t h i n

t h e u p p e r t h i r d o f t h e t r e e c r o w n ; b ) t h e s a m p l e s i n c l u d e d f o l i a g e o f a l l

65

T A B L E 2 3 : C o n c e n t r a t i o n o f m a c r o n u t r i e n t s i n f o l i a g e w i t h i n u p p e r t h i r d o f t r e e c r o w n ( i n p e r c e n t ) , 1

T r e e T r e e S p e c i e s N o . N P C a Mg K

M o u n t a i n h e m l o c k 68 0.80 0,10 0.31 0.10 0.23 74 0.65 0.11 0.39 0.08 0.28 80 0.86 0.12 0.22 0.06 0.31

105 0.83 0.10 0.32 0.07 0.27 119 0.84 0.10 0.32 0.08 0.30

R a n g e 0.66- 0.10- 0.22- 0.06- 0.24-0.87 0.12 0.39 0.10 0.

M e a n 0.80 0.11 0.31 0.08 0.28

P a c i f i c s i l v e r f i r 2 22 0.76 0.11 0.41 0.06 0.31 41 0.80 0.10 0.30 0.07 0.32

110 0.69 0.09 0.34 0.05 0.37 159 0.52 0.07 0.51 0.06 0.44 175 0.73 0.10 0.44 0.08 0.33 213 0.76 0.10 0.40 0.06 0.38 243 0.82 0.11 0.37 0.10 0.35 R a n g e 0.52- 0.07- 0.30- 0.04- 0.31-

0.82 0.11 0.51 0.10 o.. M e a n 0.73 0.10 0.40 0.07 0.36

W e s t e r n h e m l o c k 50 0.73 0.12 0.43 0.11 0.33 54 0.74 0.10 0.49 0.11 0.37 65 0.82 0.10 0.47 0.09 0.47 69 0.76 0.12 0.33 0.11 0.34 72 0.79 0.11 0.32 0.07 0.43

103 0.76 0.12 0.36 0.13 0.34 104 0.62 0.12 0.41 0.13 0.42 108 0.87 0.12 0.28 0.09 0.23

R a n g e 0.62- 0.10- 0.28- 0.07- 0.23-0.87 0.12 0.49 0.13 0,<

M e a n 0.76 0.11 0.39 0.10 0.37

Y e l l o w c e d a r 8 0.70 0,08 1.14 0.06 0.31 63 0.81 0.09 1.22 0.09 0.48 76 0.73 0.09 1.18 0.06 0.30

102 0.77 0.08 1.43 0.08 0.40 R a n g e 0.70- 0.08- 1.14- 0.06- 0.30-

0.81 0.09 1.43 0.09 o.. M e a n 0.75 0.09 1.24 0.07 0.37

1 - D a t a f r o m s a m p l e s i n c l u d i n g s e v e r a l f o l i a r a g e c l a s s e s t a k e n b e t w e e n l a t e M ay a n d t h e e n d o f J u n e f o r y e l l o w c e d a r a n d w e s t e r n h e m l o c k , a n d b e t w e e n e a r l y J u l y a n d l a t e A u g u s t f o r P a c i f i c s i l v e r f i r a n d m o u n t a i n h e m l o c k .

2 - D a t a f o r P a c i f i c s i l v e r f i r a n d y e l l o w c e d a r a r e f o r t w i g s a n d f o l i a g e t o g e t h e r a n d t h e r e f o r e t h e v a l u e s a r e l o w e r .

66

a g e s , i n some c a s e s u p t o 2 0 y e a r s o l d ; c ) . f o l i a g e a n d t w i g s w e r e n o t s e p -

a r a t e d f o r P a c i f i c s i l v e r f i r a n d y e l l o w c e d a r ; d ) . f o l i a g e s a m p l e s f r o m

d i f f e r e n t t r e e s w e r e o b t a i n e d t h r o u g h a t i m e - p e r i o d o f a b o u t t w o m o n t h s .

T h e d i s t r i b u t i o n o f f i v e m a c r o n u t r i e n t s ( i n k g / h a ) i n t h e d i f f e r -

e n t t r e e c o m p o n e n t s i s s u m m a r i z e d f o r t h e S q u a m i s h s a m p l e p l o t i n T a b l e 24

a n d f o r t h e H a n e y s a m p l e p l o t i n T a b l e 2 5 , A s c a n b e s e e n , t h e a b o v e -

g r o u n d t r e e b i o m a s s o n t h e S q u a m i s h s a m p l e p l o t c o n t a i n s a b o u t s e v e n a n d

o n e - h a l f t i m e s a s m u c h m a c r o n u t r i e n t s p e r h e c t a r e a s o n t h e H a n e y s a m p l e

p l o t . T h e r a t i o o f b i o m a s s o n t h e t w o p l o t s i s s i m i l a r t o t h e n u t r i e n t

r a t i o , w i t h t h e S q u a m i s h s a m p l e p l o t h a v i n g a p p r o x i m a t e l y e i g h t t i m e s a s

m u c h b i o m a s s a s t h e H a n e y s a m p l e p l o t . T h i s r e f l e c t s t h e r e l a t i v e l y s m a l l

v a r i a t i o n i n c o n c e n t r a t i o n o f m a c r o n u t r i e n t s r e l a t i v e t o t h e d i f f e r e n c e i n

b i o m a s s .

A v e r a g e r e l a t i v e a m o u n t s o f m a c r o n u t r i e n t s i n d i f f e r e n t a b o v e -

g r o u n d t r e e c o m p o n e n t s a r e l i s t e d i n T a b l e 2 6 . T r e e b o l e s c o n t a i n f r o m

74 t o 85 p e r c e n t o f t h e a b o v e - g r o u n d t r e e b i o m a s s , b u t o n l y 4 1 t o 56 p e r

c e n t o f t h e t r e e n i t r o g e n c o n t e n t , 47 t o 73 p e r c e n t o f t h e t r e e p h o s p h o r o u s

c o n t e n t , 5 1 t o 76 p e r c e n t o f t h e t r e e c a l c i u m c o n t e n t , 4 9 t o 7 4 p e r c e n t

o f t h e t r e e m a g n e s i u m c o n t e n t a n d 5 3 t o 74 p e r c e n t o f t h e t r e e p o t a s s i u m

c o n t e n t .

T h e s i t u a t i o n i n t h e t r e e c r o w n i s r e v e r s e d . T h e t r e e c r o w n s

c o n t a i n f r o m 1 5 t o 26 p e r c e n t o f t h e a b o v e - g r o u n d t r e e b i o m a s s , b u t 44 t o

67

T A B L E 2 4 : D i s t r i b u t i o n o f m a c r o n u t r i e n t s b y p a r t i c u l a r t r e e c o m p o n e n t s o n t h e S q u a m i s h s a m p l e p l o t ( i n k g / h a ) . , n ; M o u n t a i n h e m l o c k = 7 4 ; P a c i f i c s i l v e r f i r = 1 8 2 ,

T r e e T r e e C o m p o n e n t S p . N P C a Mg K

Wood M.h. 46 16 6 1 16 6 1 P . s . f , 1 0 7 1 3 1 4 2 26 1 3 3 Sum 1 5 4 29 2 0 3 4 2 1 9 4

B a r k M.h. 4 9 17 87 5 25 P . s . f . 1 0 9 20 2 1 9 1 3 6 0 Sum 1 5 8 37 3 0 6 18 8 5

B i g b r a n c h e s M.h. 13 3 16 2 7 P . s . f . 3 9 5 87 ' 7 25 Sum 5 2 8 1 0 2 9 3 2

S m a l l b r a n c h e s M.h. 3 1 3 1 2 P . s . f . 1 8 3 25 2 1 0 Sum 2 1 4 2 8 3 1 2

T w i g s M.h. 1 3 2 6 2 6 P . s . f . - - - - -Sum - - — — —

F o l i a g e M.h. 2 8 4 10 3 1 0 P . s . f , - - - - -Sum — - - — —

T w i g s & F o l i a g e M.h. 4 0 6 16 4 1 5 P . s . f . 1 3 2 1 8 86 1 2 .61 Sum 1 7 2 24 1 0 2 16 76

T r e e B o l e M.h. 95 3 3 1 4 8 2 1 86 ( t o 1 - i n c h o f P . s . f . 2 1 7 3 3 3 6 1 39 1 9 2 d i a m e t e r ) Sum 3 1 2 66 5 0 9 6 0 2 7 9

T r e e C r o w n M.h. 56 9 35 7 24 P . s . f . 1 8 9 2 6 1 9 7 2 0 96 Sum 2 4 6 35 2 3 2 27 1 2 0

T o t a l M.h. 1 5 1 42 1 8 3 27 1 1 1 P . s . f . 4 0 6 59 5 5 8 6 0 2 8 8 Sum 5 5 7 1 0 1 7 4 1 87 3 9 9

68

T A B L E 2 5 : D i s t r i b u t i o n o f m a c r o n u t r i e n t s b y p a r t i c u l a r t r e e c o m p o n e n t s o n t h e H a n e y s a m p l e p l o t ( i n k g / h a ) , n ; W e s t e r n h e m l o c k = 6 7 ; Y e l l o w c e d a r = 1 5 .

T r e e T r e e C o m p o n e n t S p . N P C a Mg K

Wood W.h. 1 2 4 14 3 2 1 Y . c , 4 1 8 1 3 Sum 16 5 2 2 4 2 4

B a r k W.h. 1 0 2 2 3 1 6 Y . c . 2 0.4 14 0.3 2 Sum 1 2 2 37 1 8

B i g b r a n c h e s W.h. 5 1 8 1 3 Y . c . 1 0.3 4 0.3 0.5 Sum 6 1 1 2 1 4

S m a l l b r a n c h e s W.h. 7 1 9 1 5 Y . c . 1 0.1 3 0.1 0.6 Sum 8 1 1 2 1 5

T w i g s W.h. 6 1 4 1 3 Y . c . - - - ' — _

Sum - - - - -F o l i a g e W.h. 15 2 9 2 7

Y . c . - - - — —

Sum - - - - -T w i g s & F o l i a g e W.h. 2 1 3 1 3 3 9

Y . c . 6 1 1 1 0.6 3 Sum 27 4 24 3 12

T r e e B o l e W.h. 2 3 6 37 4 27 ( t o 1 - i n c h o f Y . c . 6 1 2 2 1 5 d i a m e t e r ) Sum 29 7 5 9 5 32

T r e e C r o w n W.h. 34 6 3 0 5 17 Y . c . 7 1 19 1 4 Sum. 4 1 7 49 6 2 1

T o t a l W.h. 57 1 1 66 9 44 Y . c . 14 2 4 1 2 9 Sum 70 13 1 0 7 1 1 5 3

69

T A B L E 2 6 : A v e r a g e p r o p o r t i o n s o f f i v e m a c r o n u t r i e n t s i n d i f f e r e n t t r e e c o m p o n e n t s o n t h e S q u a m i s h a n d H a n e y s a m p l e p l o t s ( i n p e r c e n t ) . n ; M o u n t a i n h e m l o c k = 5; P a c i f i c s i l v e r f i r = 7; w e s t e r n hem-l o c k = 8; Y e l l o w c e d a r = 4.

T r e e C o m p o n e n t T r e e N P C a Mg K B i o m a s s S p .

Mg

Wood M.h. 2 9 . 7 3 8 . 2 3 8 , 8 6 0 . 3 5 2 . 7 6 7 . 2 P . s . f . 2 5 . 0 2 0 . 9 2 4 . 9 4 3 . 8 4 5 . 9 6 6 . 3 W.h. 2 2 . 0 3 5 . 4 2 2 . 4 3 6 . 5 4 6 . 5 6 0 . 8 Y . c . 2 8 . 1 2 6 . 0 1 8 . 2 3 6 . 2 3 8 . 2 7 1 . 7

B a r k M.h. 2 6 . 0 3 5 . 1 3 7 . 3 1 3 . 8 2 1 . 1 1 7 . 9 P . s . f . 2 4 . 4 3 1 . 5 3 7 . 6 2 1 . 8 1 9 . 8 1 3 . 9 W.h. 1 8 . 8 1 5 . 7 3 4 . 6 1 2 . 8 1 5 . 3 1 3 . 3 Y . c . 1 3 . 2 2 1 . 3 3 2 . 7 1 3 . 8 1 5 . 2 6.4

B i g b r a n c h e s M.h. 8.2 6.7 9.2 6.9 6.6 6.9 P . s . f . 1 0 . 4 9.8 1 7 . 3 1 0 . 9 9.1 8.1 W.h. 6.9 6.1 1 0 . 0 6.9 6.4 6.6 Y . c . 7.8 9.1 1 2 . 2 9.3 7.1 5.9

S m a l l b r a n c h e s M.h. 2.3 1.8 2.2 1.9 1.7 1.9 P . s . f . 4.8 4.9 4.5 4.0 3.7 3.9 W.h. 1 1 . 3 9.6 1 2 . 5 1 0 . 5 9.0 1 1 . 3 Y . c . 7.6 7.6 8.1 6.9 6.9 6.2

T w i g s M.h. 8.3 4.8 3.6 4.9 5.0 2.5 P . s . f . - - - - - -W.h. 9.8 7.4 5.3 6.7 5.2 3.3 Y . c . — — — - — -

F o l i a g e M.h. 2 5 . 5 1 3 . 4 8.9 1 2 . 2 1 2 . 9 3.6 P . s . f . - - - - - -W.h. 3 1 . 2 2 5 . 8 1 5 . 2 2 6 . 6 1 7 . 6 4.7 Y . c . — — — — — -

T w i g s & F o l i a g e M.h. 3 3 . 8 1 8 . 2 1 2 . 5 1 7 . 1 1 7 . 9 6.1 P . s . f . 3 5 . 4 3 2 . 9 1 5 . 7 1 9 . 5 2 1 . 9 7.8 W.h, 4 1 . 0 3 3 . 2 2 0 . 5 3 3 . 3 2 2 . 8 8.0 Y . c . 4 3 . 3 3 6 . 0 2 8 . 8 3 3 . 8 3 2 . 6 9.8

T r e e B o l e M.h. 5 5 . 7 7 3 . 3 7 6 . 1 7 4 . 1 7 3 . 8 8 5 . 1 P . s . f . 49.. 4 5 2 . 4 6 2 . 5 6 5 . 6 6 4 . 8 8 0 . 2 W.h. 4 0 . 8 5 1 . 1 5 7 . 0 4 9 . 3 6 1 . 8 7 4 . 1 Y . c . 4 1 . 3 4 7 . 3 5 0 . 9 5 0 . 0 5 3 . 4 7 8 . 1

70

T A B L E 26 ( c o n t i n u e d ) . :

T r e e T r e e C o m p o n e n t S p . N P C a Mg K B i o m a s s

T r e e C r o w n M.h. 4 4 . 3 2 6 . 7 2 3 . 9 2 5 . 9 2 6 . 2 1 4 . 9 P . s . f . 5 0 . 6 4 7 . 6 3 7 . 5 3 4 . 4 3 5 . 2 1 9 . 3 W.h. 5 9 . 2 4 8 . 9 4 3 . 0 5 0 . 7 3 8 . 2 2 5 . 9 Y . c . 5 8 . 7 5 2 . 7 4 9 . 1 5 0 . 0 4 6 . 6 2 1 . 9

59 p e r c e n t o f t h e t r e e n i t r o g e n c o n t e n t , 27 t o 5 3 p e r c e n t o f t h e t r e e

p h o s p h o r o u s c o n t e n t , 24 t o 4 9 p e r c e n t o f t h e t r e e c a l c i u m c o n t e n t , 26 t o 5 1

p e r c e n t o f t h e t r e e m a g n e s i u m c o n t e n t a n d 26 t o 47 p e r c e n t o f t h e t r e e

p o t a s s i u m c o n t e n t . T o c o m p a r e w i t h d a t a f o r y o u n g s t a n d s , s e e T a b l e s 7

a n d 9.

C o m p a r i n g d i f f e r e n t t r e e s p e c i e s , we f i n d t h a t m o u n t a i n h e m l o c k

h a s t h e h i g h e s t p r o p o r t i o n o f c a t i o n s a n d p h o s p h o r o u s w i t h i n t h e b o l e , b u t

t h e l o w e s t w i t h i n t h e c r o w n . C o n v e r s e l y , y e l l o w c e d a r h a s t h e l o w e s t p r o p o r -

t i o n o f c a t i o n s a n d p h o s p h o r o u s w i t h i n t h e b o l e a n d t h e h i g h e s t w i t h i n t h e

c r o w n . T h e d i s t r i b u t i o n o f n i t r o g e n i s m o r e p r o p o r t i o n a l t o t h e d i s t r i b u -

t i o n o f b i o m a s s . T h u s , i n s u m m a r y , t h e f o u r s p e c i e s a p p e a r i n t h e f o l l o w i n g

o r d e r a c c o r d i n g t o t h e p r o p o r t i o n a l n u t r i e n t a m o u n t i n t r e e t r u n k s :

m o u n t a i n h e m l o c k ^ P a c i f i c s i l v e r f i r > w e s t e r n

h e m l o c k ^> y e l l o w c e d a r ,

I f we c o m p a r e a m o u n t o f n u t r i e n t s w i t h i n t r e e c r o w n s , t h e s p e c i e s w o u l d

a p p e a r i n t h e r e v e r s e o r d e r .

71

T h e n u t r i e n t c o n t e n t w i t h i n t h e a b o v e - g r o u n d t r e e b i o m a s s o n t h e

S q u a m i s h p l o t i s 5 5 7 k g / h a o f n i t r o g e n , 1 0 1 k g / h a o f p h o s p h o r o u s , 7 4 1 k g / h a

o f c a l c i u m , 8 7 k g / h a o f m a g n e s i u m a n d 3 9 9 k g / h a o f p o t a s s i u m . T h e c o r r e s -

p o n d i n g v a l u e s f o r t h e H a n e y p l o t a r e s i g n i f i c a n t l y l o w e r : 7 0 k g / h a o f

n i t r o g e n , 1 3 k g / h a o f p h o s p h o r o u s , 1 0 7 k g / h a o f c a l c i u m , 1 1 k g / h a o f m a g n e s -

i u m a n d 5 3 k g / h a o f p o t a s s i u m ( T a b l e s 24 a n d 2 5 ) . T h e s e a m o u n t s a r e c o m -

p a r e d w i t h d a t a f o r o t h e r t r e e s p e c i e s f r o m o t h e r g e o g r a p h i c a l a r e a s i n

T a b l e 2 7 .

B e c a u s e o f t h e v a r i a t i o n i n m a c r o n u t r i e n t c o n c e n t r a t i o n d i s c u s s e d

a b o v e t h e d i s t r i b u t i o n o f m a c r o n u t r i e n t s d o e s n o t a c c u r a t e l y r e f l e c t b i o m a s s

d i s t r i b u t i o n ( T a b l e 2 6 ) . Wood o f t r e e b o l e s r e p r e s e n t s f r o m 6 1 t o 72 p e r

c e n t o f t h e w h o l e t r e e b i o m a s s b u t a c c o u n t s f o r o n l y 22 t o 3 0 p e r c e n t o f

t h e n i t r o g e n , 2 1 t o 38 p e r c e n t o f t h e p h o s p h o r o u s , 1 8 t o 39 p e r c e n t o f

t h e c a l c i u m , 3 6 t o 6 0 p e r c e n t o f t h e m a g n e s i u m a n d 38 t o 5 3 p e r c e n t o f

t h e p o t a s s i u m . B a r k o f t r e e b o l e s r e p r e s e n t s f r o m 6 t o 1 8 p e r c e n t o f t h e

w h o l e t r e e b i o m a s s b u t c o n t a i n s 1 3 t o 26 p e r c e n t o f t h e n i t r o g e n , 16 t o 35

p e r c e n t o f t h e p h o s p h o r o u s , 3 3 t o 38 p e r c e n t o f t h e c a l c i u m , 1 3 t o 22

p e r c e n t o f t h e m a g n e s i u m a n d 1 5 t o 2 1 p e r c e n t o f t h e p o t a s s i u m . B i g

b r a n c h e s r e p r e s e n t f r o m 6 t o 8 p e r c e n t o f t h e t o t a l t r e e b i o m a s s a n d r e p -

r e s e n t a p p r o x i m a t e l y t h e same p r o p o r t i o n o f m a c r o n u t r i e n t s : 7 t o 1 0 p e r

c e n t o f t h e n i t r o g e n , 6 t o 1 0 p e r c e n t o f t h e p h o s p h o r o u s , 1 0 t o 17 p e r

c e n t o f t h e c a l c i u m , 7 t o 1 1 p e r c e n t o f t h e m a g n e s i u m a n d 6 t o 9 p e r c e n t

o f t h e p o t a s s i u m . S m a l l b r a n c h e s r e p r e s e n t f r o m 2 t o 1 1 p e r c e n t o f t h e

w h o l e t r e e b i o m a s s a n d a c c o u n t f o r a p p r o x i m a t e l y t h e same a m o u n t o f t h e

T A B L E 2 7 : T h e a m o u n t o f m a c r o n u t r i e n t s i m m o b i l i z e d i n a b o v e - g r o u n d t r e e b i o m a s s b y v a r i o u s s p e c i e s i n d i f f e r e n t c o u n t r i e s .

i n

X

A u t h o r , y e a r D e s c r i p t i o n o f s t a n d A g e 1 0 0 0 k g / h a k g / h a A u t h o r , y e a r B i o m a s s N. P K C a Mg

R e n n i e , 1 9 5 5 P i n e s , a v e r a g e o f many s t a n d s E u r o p e . 1 0 0 - - 1 3 6 3 1 2 9 —

R e n n i e , 1 9 5 5 O t h e r c o n i f e r s , a v e r a g e o f many s t a n d s , E u r o p e . 1 0 0 - - 3 1 1 7 0 3 0 7 —

R e n n i e , 1 9 5 5 H a r d w o o d s , a v e r a g e o f many s t a n d s , E u r o p e , 1 0 0 - - 3 1 1 4 4 5 8 2 —

W r i g h t & W i l l , 1 9 5 8 S c o t s p i n e , E n g l a n d , 64 1 1 9 2 0 5 2 3 1 0 0 2 0 2 3 5 W i l l , 1 9 6 4 R a d i a t a p i n e , New Z e a l a n d . 35 3 6 2 2 3 0 32 2 8 3 1 8 8 —

C r a i g & M i l l e r , 1 9 6 6 C o r s i c a n p i n e , E n g l a n d . 35 2 0 0 5 8 6 75 2 0 7 2 3 0 9 3 S w i t z e r & N e l s o n , 1 9 7 2 L o b l o l l y p i n e , M i s s i s s i p p i ,

U.S.A. 20 9 0 1 7 4 1 9 9 8 9 0 2 3 S m i r n o v a , 1 9 5 1 S p r u c e f o r e s t , U.S.S.R. 9 3 - 4 6 8 85 2 7 8 5 5 2 9 0 C o l e , e t a l . , 1 9 6 7 D o u g l a s - f i r ( i n c l u d i n g r o o t s ) ,

3 3 3 W a s h i n g t o n , U.S.A. 36 2 0 5 3 2 0 66 2 2 0 3 3 3 —

M o r r i s o n , 1 9 7 2 12 j a c k p i n e s t a n d s , O n t a r i o . 65 1 0 6 1 6 1 1 3 8 1 1 1 6 1 8 O v i n g t o n , 1 9 6 8 D o u g l a s - f i r s t a n d , l i t t e r a n d

2 5 7 3 8 8 O v i n g t o n , 1 9 6 8

u n d e r s t o r e y v e g e t a t i o n . 47 2 6 4 6 3 6 6 2 2 5 7 3 8 8 —

O v i n g t o n , 1 9 6 8 S c o t s p i n e s t a n d , l i t t e r a n d 2 2 9

O v i n g t o n , 1 9 6 8 u n d e r s t o r e y v e g e t a t i o n . 47 1 7 6 5 5 6 43 3 2 5 2 2 9 —

O v i n g t o n , 1 9 6 8 N o r w a y s p r u c e s t a n d , l i t t e r & 5 6 1 9 1 2 7 7

O v i n g t o n , 1 9 6 8 u n d e r s t o r e y v e g e t a t i o n . 47 1 6 5 6 9 1 5 6 1 9 1 2 7 7 —

O v i n g t o n , 1 9 6 8 Oak s t a n d , l i t t e r a n d u n d e r -2 9 2 s t o r e y v e g e t a t i o n . 47 1 3 4 4 6 4 4 0 2 5 4 2 9 2 —

M a t e r n a , 1 9 6 4 N o r w a y s p r u c e , C z e c h o s l o v a k i a . 1 0 0 — 8 3 0 1 0 9 3 1 7 4 5 5 3 0

T h e S q u a m i s h s a m p l e p l o t . 4 1 7 4 6 7 5 5 7 1 0 1 3 9 9 7 4 1 87 T h e H a n e y s a m p l e p l o t . 2 4 8 5 6 7 0 1 3 5 3 1 0 7 1 1

N3

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m a c r o n u t r i e n t s : 2 t o 1 1 p e r c e n t o f t h e n i t r o g e n , 2 t p 1 0 p e r c e n t o f

t h e m a g n e s i u m a n d 2 t o 9 p e r c e n t o f t h e p o t a s s i u m . T w i g s a n d f o l i a g e

w e r e s t u d i e d s e p a r a t e l y f o r m o u n t a i n a n d w e s t e r n h e m l o c k o n l y . T w i g s r e p -

r e s e n t 2.5 a n d 3.3 p e r c e n t o f t h e t o t a l t r e e b i o m a s s a n d c o n t a i n 8 a n d 1 0

p e r c e n t o f t h e n i t r o g e n , 5 a n d 7 p e r c e n t o f t h e p h o s p h o r u s , 4 a n d 5 p e r

c e n t o f t h e c a l c i u m , 5 a n d 7 p e r c e n t o f t h e m a g n e s i u m , a n d 5 p e r c e n t o f t h e

p o t a s s i u m f o r m o u n t a i n a n d w e s t e r n h e m l o c k , r e s p e c t i v e l y . F o l i a g e r e p r e -

s e n t 3.7 a n d 4,8 p e r c e n t o f t h e w h o l e t r e e b i o m a s s b u t r e t a i n s a v e r y h i g h

p r o p o r t i o n o f t r e e n u t r i e n t s : 25 a n d 3 1 p e r c e n t o f t h e n i t r o g e n , 1 3 a n d

26 p e r c e n t o f t h e p h o s p h o r u s , 9 a n d 15 p e r c e n t o f t h e c a l c i u m , 1 2 a n d 27

p e r c e n t o f t h e m a g n e s i u m , 1 3 a n d 18 p e r c e n t o f t h e p o t a s s i u m f o r m o u n t a i n

a n d w e s t e r n h e m l o c k , r e s p e c t i v e l y . I f we c o n s i d e r t w i g s a n d f o l i a g e t o -

g e t h e r f o r a l l f o u r s p e c i e s o n t w o s a m p l e p l o t s , i t a p p e a r s t h a t t w i g s a n d

f o l i a g e r e p r e s e n t s 6 t o 10 p e r c e n t o f t h e w h o l e t r e e b i o m a s s a n d c o n t a i n

3 4 t o 4 3 p e r c e n t o f t h e n i t r o g e n , 1 8 t o 36 p e r c e n t o f t h e p h o s p h o r u s , 1 2

t o 29 p e r c e n t o f t h e c a l c i u m , 17 t o 39 p e r c e n t o f t h e m a g n e s i u m a n d 1 8 t o

3 3 p e r c e n t o f t h e p o t a s s i u m .

R e m o v a l o f M a c r o n u t r i e n t s . f r o m t h e E c o s y s t e m Due t o D i f f e r e n t

L e v e l s o f T r e e U t i l i z a t i o n :

T a b l e s 28 a n d 2 9 s u m m a r i z e t h e a m o u n t o f b i o m a s s a n d m a c r o n u t r i e n t s

r e m o v e d f r o m t h e e c o s y s t e m d u e t o d i f f e r e n t l e v e l s o f t r e e u t i l i z a t i o n .

L e a v i n g s m a l l b r a n c h e s , t w i g s a n d f o l i a g e a t s t u m p w o u l d r e s u l t i n a 7 p e r

c e n t d e c r e a s e i n b i o m a s s y i e l d f r o m t h e S q u a m i s h s a m p l e p l o t . I t w o u l d

T A B L E 2 8 : R e m o v a l o f b i o m a s s a n d m a c r o n u t r i e n t s a c c o m p a n y i n g d i f f e r e n t l e v e l s o f u t i l i z a t i o n i n t h e S q u a m i s h s a m p l e p l o t .

B i o m a s s N P K C a Mg m e t r i c k g / k g / k g / k g / k g / t o n s / h a % h a % h a % h a % h a % h a %

W h o l e t r e e l o g g i n g 4 6 7 1 0 0 5 5 7 1 0 0 1 0 1 1 0 0 3 9 9 1 0 0 7 4 1 1 0 0 87 1 0 0

U t i l i z a t i o n o f t r e e b o l e s u p t o 1 - i n c h o f d i a m e t e r a n d b r a n c h e s o v e r 1 - i n c h o f d i a m e t e r . 4 3 3 9 3 3 6 4 65 73 73 3 1 1 78 6 1 2 8 3 69 79

U t i l i z a t i o n o f t r e e b o l e s u p t o 1 - i n c h o f d i a m e t e r . 3 9 7 8 5 3 1 2 5 6 65 65 2 7 9 70 5 0 9 69 6 0 69

U t i l i z a t i o n o f s t e m w o o d o n l y u p t o 1 - i n c h o f d i a m e t e r . 3 1 9 68 1 5 4 28 29 29 1 9 4 49 2 0 3 27 42 48

T A B L E 2 9 : R e m o v a l o f b i o m a s s a n d m a c r o n u t r i e n t s a c c o m p a n y i n g d i f f e r e n t l e v e l s o f u t i l i z a t i o n i n t h e H a n e y s a m p l e p l o t .

B i o m a s s N P K C a Mg m e t r i c k g / k g / k g / k g / k g / t o n s / h a % h a % h a % h a % h a % h a %

W h o l e t r e e l o g g i n g 5 6 1 0 0 70 1 0 0 1 3 1 0 0 5 3 1 0 0 1 0 7 1 0 0 1 1 1 0 0

U t i l i z a t i o n o f t r e e b o l e s u p t o 1 - i n c h o f d i a m e t e r a n d b r a n c h e s o v e r 1 - i n c h d i a m e t e r . 47 84 3 5 4 9 8 57 36 68 71 66 6 5 5

U t i l i z a t i o n o f t r e e b o l e s u p t o 1 - i n c h o f d i a m e t e r . 4 1 7 3 29 4 1 7 5 0 32 6 0 59 5 5 5 4 5

U t i l i z a t i o n o f s t e m w o o d o n l y up t o 1 - i n c h o f d i a m e t e r . 27 4 8 16 23 5 36 24 45 22 2 1 4 36

76

d e c r e a s e t h e r e m o v a l o f n i t r o g e n b y 35 p e r c e n t a n d c a l c i u m b y 17 p e r c e n t .

U t i l i z a t i o n o f t r e e b o l e o n l y u p t o 1 - i n c h o f d i a m e t e r w o u l d d e c r e a s e b i o -

m a s s y i e l d b y 1 5 p e r c e n t b u t i t w o u l d d e c r e a s e t h e r e m o v a l o f m a c r o n u t r i -

e n t s m o r e s u b s t a n t i a l l y . R e m o v a l o f n i t r o g e n f r o m t h e e c o s y s t e m w o u l d b e

r e d u c e d b y 44 p e r c e n t , p h o s p h o r o u s b y 3 5 p e r c e n t a n d c a t i o n s b y 3 0 p e r

c e n t . I n c o m p a r i s o n , h a r v e s t i n g o f s t e m w o o d o n l y u p t o 1 - i n c h o f d i a m e t e r

w o u l d r e d u c e b i o m a s s y i e l d b y 3 2 p e r c e n t , r e m o v a l o f N, P, C a b y a s m u c h

a s 72 p e r c e n t a n d r e m o v a l o f K a n d Mg b y 5 1 p e r c e n t . On t h e H a n e y s a m p l e

p l o t , r e d u c t i o n o f b i o m a s s y i e l d w o u l d b e g r e a t e r a c c o m p a n i e d a l s o b y g r e a t e r

r e d u c t i o n o f n u t r i e n t l o s s e s .

77

V I SUMMARY AND CONCLUSIONS

T h e m e t h o d u s e d f o r t h e d e t e r m i n a t i o n o f t h e n u t r i e n t d e m a n d o n

t h e s i t e o f t w o f o r e s t s t a n d s w a s b a s e d o n a m e a s u r e m e n t o f t h e t o t a l n u t r i -

e n t i m m o b i l i z a t i o n b y t h e s t a n d ( e x c l u d i n g s t u m p a n d r o o t b i o m a s s ) : t h e

s o - c a l l e d s u b s t i t u t i o n p r i n c i p l e (Tamm, 1 9 6 4 ) . A c c o r d i n g t o Tamm, t h i s i s

o n l y a f i r s t a p p r o x i m a t i o n f o r d e t e r m i n i n g t r e e n u t r i e n t r e q u i r e m e n t s .

N e v e r t h e l e s s , t h i s m e t h o d h a s b e e n u s e d b y a n u m b e r o f d i f f e r e n t a u t h o r s

( e . g . E b e r m a y e r 1 8 7 6 , 1 8 8 2 ; R e n n i e 1 9 5 5 ; O v i n g t o n 1 9 6 2 ; W e e t m a n a n d W e b b e r

1 9 7 2 , e t c . ) t o a p p r o x i m a t e t h e n e t n u t r i e n t u p t a k e o f f o r e s t s t a n d o v e r a

r o t a t i o n a n d t h e l o s s o f n u t r i e n t s f r o m a f o r e s t s i t e i n h a r v e s t e d m a t e r i a l .

T h e f o l l o w i n g c o n c l u s i o n s w e r e r e a c h e d o n t h e b a s i s o f t h i s

s t u d y :

1. V a r i o u s a b o v e - g r o u n d t r e e c o m p o n e n t s o f f o u r s t u d i e d s p e c i e s

a p p e a r t o b e i n t h e f o l l o w i n g o r d e r a c c o r d i n g t o t h e i r n u t r i e n t c o n c e n t r a -

t i o n s :

stemwood<C b i g b r a n c h e s < f s m a l l b r a n c h e s <^ s t e m b a r k <^ t w i g s < ^

f o l i a g e ( h e m l o c k spp„») o r t w i g s a n d f o l i a g e ( P a c i f i c s i l v e r f i r a n d y e l l o w

c e d a r ) . ,

T h i s o r d e r i s v a l i d f o r a l l m a c r o n u t r i e n t s w i t h t h e e x c e p t i o n o f c a l c i u m .

T h e c o n c e n t r a t i o n o f c a l c i u m i n s t e m b a r k i s h i g h e r t h a n i n t w i g s a n d a p p r o x -

i m a t e l y e q u a l t o f o l i a g e c o n c e n t r a t i o n ( h e m l o c k s p p . ) o r t w i g s a n d f o l i a g e

78

c o n c e n t r a t i o n ( f i r a n d c e d a r ) . C o n e s , w h e r e s t u d i e d , e x h i b i t e d t h e h i g h e s t

c o n c e n t r a t i o n o f N, P, K; t h e l o w e s t o f C a . T h e i r Mg c o n c e n t r a t i o n w a s

a p p r o x i m a t e l y e q u a l t o t h a t o f t w i g s a n d f o l i a g e ( f i r ) o r t w i g s ( h e m l o c k ) .

2. T h e f o u r t r e e s p e c i e s i n v e s t i g a t e d a p p e a r t o b e i n t h e f o l -

l o w i n g o r d e r a c c o r d i n g t o m a c r o n u t r i e n t c o n c e n t r a t i o n :

W e s t e r n h e m l o c k ^ M o u n t a i n h e m l o c k <^ P a c i f i c s i l v e r f i r <^

Y e l l o w c e d a r .

Y e l l o w c e d a r e x h i b i t s c o n s i d e r a b l y h i g h e r c a l c i u m c o n c e n t r a t i o n i n a l l t r e e

c o m p o n e n t s t h a t d o t h e o t h e r s p e c i e s . P a c i f i c s i l v e r f i r e x h i b i t s h i g h e r

n u t r i e n t c o n c e n t r a t i o n s f o r a l m o s t a l l e l e m e n t s a n d t r e e c o m p o n e n t s t h a n d o

b o t h h e m l o c k s p e c i e s . N u t r i e n t c o n c e n t r a t i o n s i n m o u n t a i n h e m l o c k a r e a p -

p r o x i m a t e l y e q u a l t o c o n c e n t r a t i o n s i n w e s t e r n h e m l o c k f o r a l l t r e e c o m p o n -

e n t s . S i t e c o n d i t i o n s o f t h e t w o s a m p l e p l o t s w e r e c o n s i d e r a b l y d i f f e r e n t

a n d t r e e s p e c i e s might a p p e a r i n a d i f f e r e n t o r d e r i f t h e y w e r e g r o w n u n d e r

u n i f o r m s i t e c o n d i t i o n s .

3. T h e w e i g h t d i s t r i b u t i o n o f n u t r i e n t s w i t h i n t h e a b o v e - g r o u n d

t r e e b i o m a s s i s n o t p r o p o r t i o n a l t o t h e b i o m a s s d i s t r i b u t i o n . T r e e c r o w n s

( b r a n c h e s , t w i g s a n d f o l i a g e ) c o n t a i n a s m u c h n u t r i e n t s a s t r e e b o l e s .

W i t h i n t h e c r o w n , t h e l a r g e s t a m o u n t o f n u t r i e n t s i s b u i l t u p w i t h i n t h e

f o l i a g e a n d t w i g s . T h e p r o p o r t i o n o f n u t r i e n t s i n t h e c r o w n c o m p o n e n t s

v a r i e s f o r v a r i o u s e l e m e n t s a s w e l l a s f o r d i f f e r e n t t r e e s p e c i e s .

79

4. B a r k c o n t a i n s a p p r o x i m a t e l y h a l f o f t h e n u t r i e n t s o f t h e

w h o l e t r e e b o l e . T h e p r o p o r t i o n s o f e l e m e n t s w i t h i n b a r k a n d w o o d v a r y

f o r d i f f e r e n t e l e m e n t s a n d t r e e s p e c i e s ,

5. T h e u p t a k e o f n u t r i e n t s i m m o b i l i z e d i n t h e a b o v e - g r o u n d t r e e

b i o m a s s o v e r a t i m e p e r i o d o f 4 2 0 y e a r s i n t h e S q u a m i s h s a m p l e p l o t l o c a t e d

i n t h e S u b a l p i n e M o u n t a i n H e m l o c k Z o n e - p l a n t a s s o c i a t i o n A b i e t o ( a m a b i l i s )

- T s u g e t u m m e r t e n s i a n a e v a r i a n t a b i e t o - t s u g o s u m m e r t e n s i a n a e ( B r o o k e , 1 9 7 0 )

w a s c a l c u l a t e d t o b e 5 5 7 k g o f n i t r o g e n , 1 0 1 k g o f p h o s p h o r u s , 3 9 9 k g o f

p o t a s s i u m , 7 4 1 k g o f c a l c i u m a n d 87 k g o f m a g n e s i u m p e r h e c t a r e f o r o v e n

d r y b i o m a s s o f 4 6 7 m e t r i c t o n s .

6. T h e u p t a k e o f n u t r i e n t s i m m o b i l i z e d i n t h e a b o v e - g r o u n d t r e e

b i o m a s s o v e r a t i m e p e r i o d o f 2 5 0 y e a r s i n t h e H a n e y s a m p l e p l o t l o c a t e d i n

t h e W e t S u b z o n e o f t h e C o a s t a l W e s t e r n H e m l o c k Z o n e - p l a n t a s s o c i a t i o n

A b i e t o - T s u g e t u m h e t e r o p h y l l a e ( O r l o c i , 1 9 6 5 ) w a s c a l c u l a t e d t o b e 70 k g o f

n i t r o g e n , 1 3 k g o f p h o s p h o r u s , 5 3 k g o f p o t a s s i u m , 1 0 7 k g o f c a l c i u m a n d 1 1

k g o f m a g n e s i u m p e r h e c t a r e f o r o v e n d r y b i o m a s s o f 56 m e t r i c t o n s .

7. I n c o m p a r i s o n w i t h w h o l e s - t r e e l o g g i n g , t h e a m o u n t o f n u t r i e n t s

r e m o v e d f r o m t h e s t u d y s i t e s c a n b e r e d u c e d b y 5 0 - 7 0 p e r c e n t i f o n l y s t e m -

w o o d w e r e h a r v e s t e d a n d o t h e r t r e e c o m p o n e n t s w e r e l e f t o n t h e s i t e .

8 0

R E F E R E N C E S

A l l a n , J . E. 1 9 6 9 . T h e p r e p a r a t i o n o f a g r i c u l t u r a l s a m p l e s f o r a n a l y s i s b y a t o m i c a b s o r p t i o n s p e c t r o s c o p y , V a r i a n A e r o g r a p h , C a l i f o r n i a , U.S.A.

B e a t o n , J . D., e t a l . 1 9 6 5 . O b s e r v a t i o n s o n f o l i a g e n u t r i e n t c o n t e n t o f s e v e r a l c o n i f e r o u s t r e e s p e c i e s i n B r i t i s h C o l u m b i a . F o r . C h r o n . 4 1 ( 2 ) : 2 2 2 - 3 6 .

B r o o k e , R. C. 1 9 6 6 . V e g e t a t i o n - e n v i r o n m e n t r e l a t i o n s h i p s o f s u b a l p i n e m o u n t a i n h e m l o c k z o n e e c o s y s t e m s , P h . D . T h e s i s , D e p a r t m e n t o f B o t a n y , U n i v e r s i t y o f B r i t i s h C o l u m b i a , V a n c o u v e r , 2 2 5 p . , A p p . 1 1 0 p .

B r o o k e , R. C , E . B. P e t e r s o n , V. J . K r a j i n a . 1 9 7 0 . T h e s u b a l p i n e m o u n t a i n h e m l o c k z o n e . E c o l . o f W e s t . N. A m e r . 2 ( 2 ) : 1 4 8 - 3 4 9 .

C o l e , D. W., G e s s e l , S. P., D i c e , S. F. 1 9 6 7 . D i s t r i b u t i o n a n d c y c l i n g o f N, P, K, C a i n a s e c o n d - g r o x ^ t h D o u g l a s - f i r e c o s y s t e m . S y m p o s i u m o n p r i m a r y p r o d u c t i v i t y a n d m i n e r a l c y c l i n g i n n a t u r a l e c o s y s t e m s . A m e r i c a n A s s o c i a t i o n f o r t h e A d v a n c e s o f S c i e n c e 1 3 t h A n n u a l M e e t i n g , New Y o r k C i t y , D e c e m b e r 2 7 , 1 9 6 7 . P u b l i s h e d b y t h e U n i v e r s i t y o f M a i n e P r e s s .

C u r r y , R. R. 1 9 7 3 . G e o l o g i c a n d h y d r o l o g i c e f f e c t s o f e v e n - a g e m a n a g e m e n t o n p r o d u c t i v i t y o f f o r e s t s o i l s , p a r t i c u l a r l y i n t h e D o u g l a s - f i r r e g i o n . E v e n - A g e M a n a g e m e n t S y m p o s i u m , A u g u s t 1 9 7 2 . E d . R. K. H e r m a n n , D. P. L a v e n d e r . O r e g o n S t a t e U n i v e r s i t y .

D a v i d , J . D. 1 9 6 0 . T h e d e t e r m i n a t i o n o f e x c h a n g e a b l e N a , K, C a , Mg i n s o i l s b y a t o m i c - a b s o r p t i o n s p e c t r o p h o t o m e t r y . A n a l y s t 8 5 : 4 9 5 - 5 0 3 .

D u k e U n i v e r s i t y S c h o o l o f F o r e s t r y B u l l e t i n 1 5 , 1 9 5 9 . M i n e r a l n u t r i t i o n o f t r e e s . A S y m p o s i u m . D u r h a m , N o r t h C a r o l i n a , M a y 1 9 5 9 .

D u v i g n e a u d , P., D e n a e y e r - D e S m e t , S. 1 9 6 4 . L e c y c l e d e s e l e m e n t s b i o g e n e s d a n s l ' e c o s y s t e m e f o r e t . L ' U n i v e r s i t e L i b r e d e B r u x e l l e s , P u b l . 5 8 .

81

D u v i g n e a u d , P., D e n a e y e r - D e S m e t , S. 1 9 6 7 , B i o m a s s , P r o d u c t i v i t y a n d M i n e r a l C y c l i n g i n D e c i d u o u s F o r e s t s i n B e l g i u m . S y m p o s i u m o n P r i m a r y P r o d u c t i v i t y a n d M i n e r a l C y c l i n g i n N a t u r a l E c o s y s t e m s . A m e r i c a n A s s o c i a t i o n f o r t h e A d v a n c e m e n t o f S c i e n c e , 1 3 t h A n n u a l M e e t i n g , New Y o r k C i t y , D e c e m b e r 2 7 , 1 9 6 7 . P u b l i s h e d b y t h e U n i v e r s i t y o f M a i n e P r e s s .

D y e r , R. F . 1 9 6 7 . F r e s h a n d d r y w e i g h t , n u t r i e n t e l e m e n t s a n d p u l p i n g c h a r a c t e r i s t i c s o f n o r t h e r n w h i t e c e d a r , T h u j a o c c i d e r i t a l i s . T e c h . B u l l . Me. A g r i c . E x p . S t a , , No. 2 7 , p . 4 0 .

E b e r m a y e r , E . 1 8 7 6 . D i e g e s a m t e L e h r e d e r W a l d s t r e u m i t R i i c k s i c h t a u f d i e c h e m i s c h e S t a t i k d e s W a l d b a u e s , S p r i n g e r , B e r l i n .

E b e r m a y e r , E . 1 8 8 2 . P h y s i o l o g i s c h e C h e m i e d e r P f l a n z e n . V o l . 1, 8 6 1 p p . , S p r i n g e r , B e r l i n .

F r e e s e , F . 1 9 6 4 . L i n e a r r e g r e s s i o n m e t h o d s f o r f o r e s t r e s e a r c h . U. S. D e p a r t m e n t o f A g r i c u l t u r e , F o r e s t S e r v i c e F o r e s t P r o d u c t L a b . , M a d i s o n , W i s .

F o r e s t r y H a n d b o o k f o r B r i t i s h C o l u m b i a , T h i r d E d i t i o n , 1 9 7 1 . T h e F o r e s t C l u b , U n i v e r s i t y o f B r i t i s h C o l u m b i a .

G e s s e l , S. P. 1 9 6 2 , P r o g r e s s a n d p r o b l e m s i n m i n e r a l n u t r i t i o n o f f o r e s t t r e e s , T r e e G r o w t h . E d . T. T. K o z l o w s k i . T h e R o n a l d P r e s s C o m p a n y , New Y o r k .

G e s s e l , S. P., C o l e , D. W. 1 9 6 5 . I n f l u e n c e o f r e m o v a l o f f o r e s t c o v e r o n m o v e m e n t o f w a t e r a n d a s s o c i a t e d e l e m e n t s t h r o u g h s o i l . J o u r n a l o f A m e r i c a n W a t e r W o r k s A s s o c i a t i o n , New Y o r k , 5 7 , 1 9 6 5 ( 1 3 0 1 - 1 0 ) .

G e s s e l , S. P., D. W, C o l e , J , T u r n e r , 1 9 7 3 . E l e m e n t a l c y c l i n g a n d e v e n - a g e f o r e s t m a n a g e m e n t . E v e n - A g e M a n a g e m e n t , S y m p o s i u m , A u g u s t 1 9 7 2 . E d . R, K, H e r m a n n , D. D. L a v e n d e r . S c h o o l o f F o r e s t r y , O r e g o n S t a t e U n i v e r s i t y .

H a l l i d a y , W. E . D. 1 9 3 7 . A f o r e s t c l a s s i f i c a t i o n o f C a n a d a . C a n a d a D e p t . M i n e s a n d R e s o u r c e s , F o r e s t S e r v i c e B u l l e t i n 8 9 , K i n g ' s P r i n t e r O t t a w a .

H e i n r i c h s , F. J . 1 9 5 4 . R a p i d s p e c i f i c g r a v i t y d e t e r m i n a t i o n . J . F o r . P r o d . R e s . S o c . 4 ( 1 ) : 6 8 .

82

H o l m e n , H, 1 9 6 4 , F o r e s t e c o l o g i c a l s t u d i e s o n d r a i n e d p e a t l a n d i n t h e p r o v i n c e o f U p p l a n d , S w e d e n , P a r t s I - I I I . S t u d , F o r . S v e c i a 1 6 .

I n g e s t a d , T, 1 9 6 2 , M a c r o E l e m e n t N u t r i t i o n o f P i n e , S p r u c e a n d B i r c h S e e d l i n g s i n N u t r i e n t S o l u t i o n . 1 2 1 p p . M e d d e l a n d e n F r a n S t a t e n s S k o g s f o r s k n i n g s i n s t i t u t 5 1 ( 7 ) .

I n g e s t a d , T. 1 9 7 0 . A d e f i n i t i o n o f o p t i m u m n u t r i e n t r e q u i r e m e n t s i n b i r c h s e e d l i n g s , P h y s i o l . P l a n t . 2 3 ( 6 ) : 1 1 2 7 - 3 8 ,

K a u l , 0. N., S r i v o s h a n a , P. B. L . 1 9 6 4 , M i n e r a l c o n t e n t o f f i r ( A b i e s p i n d r o w ) . a n d K a i l ( P i n u s w a l l i c h i a n a ) . J o u r n a l o f t h e S o c i e t y o f I n d i a n F o r e s t e r s 4 ( 2 1 3 ) : 1 1 4 - 2 0 .

K e a y , J . , A. G, T u r t o n . 1 9 6 9 . D i s t r i b u t i o n o f b i o m a s s a n d m a j o r n u t r i e n t s i n a m a r i t i m e p i n e p l a n t a t i o n , A u s t r a l i a n F o r e s t r y 3 5 : 3 9 - 4 8 ,

K e l l e y , J . D., S h i e r , R. W. 1 9 6 5 , S e a s o n a l c h a n g e s i n t h e m a c r o n u t r i e n t c o m p o s i t i o n o f l e a v e s a n d s t e m s o f T a x u s m e d i a . P r o c . A m e r . S o c . H o r t . S c i . 8 6 : 8 0 9 - 1 4 .

K r a j i n a , V. H. 1 9 6 5 . B i o g e o c l i m a t i c z o n e s a n d c l a s s i f i c a t i o n o f B r i t i s h C o l u m b i a , E c o l , o f W e s t e r n N. Am. 1 : 1 - 1 7 .

K r a j i n a , V. J . 1 9 6 9 , E c o l o g y o f f o r e s t t r e e s i n B r i t i s h C o l u m b i a . E c o l . o f W e s t e r n N. Am. 2 : 1 - 1 4 6 .

K u r u c z , J . 1 9 6 9 . C o m p o n e n t w e i g h t s o f D o u g l a s - f i r , w e s t e r n h e m l o c k a n d w e s t e r n r e d c e d a r b i o m a s s f o r s i m u l a t i o n o f a m o u n t a n d d i s t r i -b u t i o n o f f o r e s t f u e l s , M.F. T h e s i s , F a c u l t y o f F o r e s t r y , U n i v e r s i t y o f B r i t i s h C o l u m b i a ,

L a f o n d , A., L a f l o m m e , Y. 1 9 7 1 . R e l a t i v e c o n c e n t r a t i o n s o f i r o n a n d m a n g a -n e s e : a f a c t o r a f f e c t i n g j a c k p i n e r e g e n e r a t i o n a n d j a c k p i n e -b l a c k s p r u c e s u c c e s s i o n . I n T r e e g r o w t h a n d f o r e s t s o i l s . E d , C h . T. Y o u n g b e r g , C h . B. D a v e y . O r e g o n S t a t e U n i v e r s i t y P r e s s , C o r v a l l i s 1 9 7 1 , p p . 3 0 5 - 1 2 .

L a m b e r t s , D. 1 9 6 2 . U p t a k e a n d r e m o v a l o f m i n e r a l n u t r i e n t s b y P i n u s n i g r a v a r . c a l a b r i c a P e d o l o g i e 1 2 ( 1 ) : 1 0 1 - 1 4 .

L e y t o n , L . 1 9 5 7 . T h e m i n e r a l r e q u i r e m e n t s o f f o r e s t p l a n t s . T h e O h i o J o u r n a l o f S c i e n c e 5 7 ( 6 ) : 3 3 7 .

8 3

L i k e n s , G, E . , B o r m a n n , F, H., J o h n s o n , N. M., P i e r c e , R, S, 1 9 6 7 . T h e c a l c i u m , m a g n e s i u m , p o t a s s i u m a n d s o d i u m b u d g e t s f o r a s m a l l f o r e s t e d e c o s y s t e m . E c o l o g y , 4 8 : 7 7 2 - 7 8 5 .

L j a m e b o r s a j , S. H. 1 9 6 8 . £ C a l c u l a t i o n o f t h e c o e f f i c i e n t s o f r e m o v a l o f N a n d a s h e l e m e n t s b y d i f f e r e n t t r e e s p e c i e s ] , D o k l a d y , M o s k o v s k a j a S e l s k c h o z j a j s t v e n n a j a A k a d e m i j a i m . K, A. T i m i r j a z e v a , M o s k v a , 1 3 : 4 1 1 - 5 ,

M a d e r , D, L . , H o w a r t h , J . A. 1 9 6 8 , R e l a t i o n s h i p s b e t w e e n f o l i a r n u t r i e n t l e v e l s a n d t r e e g r o w t h i n r e d p i n e s t a n d s i n M a s s a c h u s e t t s . T r e e g r o w t h a n d F o r e s t S o i l s , E d , C h , T. Y o u n g b e r g , C h . B, D a v e y , 1 9 6 8 . C o r v a l l i s , O r e g o n S t a t e U n i v e r s i t y P r e s s .

M a d g w i c k , H. A. I . 1 9 6 3 . N u t r i e n t r e s e a r c h , some p r o b l e m s o f t h e t o t a l t r e e a p p r o a c h . P r o c . S o i l S c i , S o c . A m e r . 27 ( 5 ) : 5 9 8 - 6 0 0 .

M a t e r n a , J . 1 9 6 4 . £ D a t a f o r e v a l u a t i n g t h e n u t r i t i o n o f N o r w a y s p r u c e t r e e s a n d s t a n d s J . P r a c e V y z k u m . U s t . L e s n . C S S R 2 9 : 1 0 3 - 3 3 .

M a t e r n a , J . 1 9 6 7 . B o h e m i a .

S t u d i e s o n t h e n u t r i t i o n o f P i n u s s i l v e s t r i s s t a n d s i n

P r a c e V y z k u m . U s t , L e s n . H o s p . M y s l . Z b r a s l a v - S t r n a d y C S S R 3 4 : 8 1 - 1 0 6 .

M e t z , L . J . , a n d W e l l s , C. G. 1 9 6 5 , W e i g h t a n d n u t r i e n t c o n t e n t o f t h e a b o v e - g r o u n d p a r t s o f some l o b l o l l y p i n e s . U.S. F o r . S e r v . R e s . P o p . S t h e a s p . F o r . E x p . S t a . N o . S E - 1 7 , 1 9 6 5 , P. 2 0 .

M o r r i s o n , I . K. 1 9 7 2 . V a r i a t i o n w i t h c r o w n p o s i t i o n a n d l e a f a g e i n c o n t e n t o f s e v e n e l e m e n t s i n l e a v e s o f P i n u s b a n k s i a n a L a m b . C a n . J , F o r . R e s . 2 : 8 9 - 9 4 ,

M o r r i s o n , I , K, 19.73, D i s t r i b u t i o n o f e l e m e n t s i n a e r i a l c o m p o n e n t s o f s e v e r a l n a t u r a l j a c k p i n e s t a n d s i n N o r t h e r n O n t a r i o . C a n , J . F o r . R e s . 3 : 1 7 0 - 7 9 .

O r l o c i , L . 1 9 6 1 . F o r e s t t y p e s o f t h e C o a s t a l W e s t e r n H e m l o c k Z o n e , M, S c . T h e s i s , D e p t . B i o l o g y a n d B o t a n y , U n i v e r s i t y o f B r i t i s h C o l u m b i a , V a n c o u v e r . 2 0 6 p .

O r l o c i , L . 1 9 6 4 . V e g e t a t i o n a l a n d e n v i r o n m e n t a l v a r i a t i o n i n t h e e c o s y s t e m s o f t h e C o a s t a l W e s t e r n H e m l o c k Z o n e . P h . D . T h e s i s , D e p t . B o t a n y , U n i v e r s i t y o f B r i t i s h C o l u m b i a , V a n c o u v e r , 1 9 9 p .

84

O v i n g t o n , J , D. 1 9 5 9 . M i n e r a l c o n t e n t o f p l a n t a t i o n s of- P i n u s s i l v e s t r i s L . A n n a l s o f B o t a n y , N , S. 2 3 ( 8 9 ) : 75-88 .

O v i n g t o n , J , D. 1 9 5 9 , T h e c i r c u l a t i o n o f m i n e r a l s i n p l a n t a t i o n s o f P i n u s ' - ' s i l v e s t r i s L .

A n n a l s o f B o t a n y , N, S. 2 3 ( 9 0 ) : 2 2 9 - 2 3 9 .

O v i n g t o n , J . D. 1 9 6 0 . T h e n u t r i e n t c y c l e a n d i t s m o d i f i c a t i o n t h r o u g h s i l v i c u l t u r a l p r a c t i c e . 5 t h W o r l d F o r e s t r y C o n g r e s s , S e a t t l e , W a s h i n g t o n ,

O v i n g t o n , J . D, 1 9 6 2 , Q u a n t i t a t i v e e c o l o g y a n d t h e w o o d l a n d e c o s y s t e m c o n c e p t . A d v a n c e s i n E c o l o g i c a l R e s e a r c h , V o l . 1, A c a d e m i c P r e s s , L o n d o n , 1 9 6 2 , p p . 1 0 3 - 9 2 .

O v i n g t o n , J . D., M a d g w i c k , A. A. I . 1 9 5 9 . D i s t r i b u t i o n o f o r g a n i c m a t t e r a n d p l a n t n u t r i e n t s i n a p l a n t a t i o n o f S c o t s p i n e . F o r e s t S c i . 5 ( 4 ) : 3 4 4 - 3 5 5 .

O v i n g t o n , J . D., W, G. F o r r e s t , J , E . A r m s t r o n g . 1 9 6 7 . T r e e b i o m a s s e s t i -m a t i o n . S y m p o s i u m o n P r i m a r y P r o d u c t i v i t y a n d M i n e r a l C y c l i n g i n N a t u r a l E c o s y s t e m s . A m e r i c a n A s s o c i a t i o n f o r t h e A d v a n c e m e n t o f S c i e n c e 1 3 t h A n n u a l M e e t i n g , New Y o r k C i t y , D e c e m b e r 1 9 6 7 . P u b l i s h e d b y t h e U n i v e r s i t y o f M a i n e P r e s s .

O r m a n , H, R., G. M. W i l l , 1 9 6 0 . T h e n u t r i e n t c o n t e n t o f P i n u s r a d i a t a t r e e s . N. Z. J , S c i . 3 : 5 1 0 - 2 2 .

P h i r i , M. A. R. 1 9 7 3 . T h e n u t r i e n t s t a t u s o f t w o h i g h e l e v a t i o n f o r e s t s o i l s a n d i m p l i c a t i o n s f o r some f o r e s t m a n a g e m e n t p r a c t i c e s . B . S . F . T h e s i s , F a c u l t y o f F o r e s t r y , U n i v e r s i t y o f B r i t i s h C o l u m b i a , V a n c o u v e r , B . C.

R e n n i e , P. J . 1 9 5 5 . T h e u p t a k e o f n u t r i e n t s b y m a t u r e f o r e s t g r o w t h . P l a n t a n d S o i l 7 ( 1 ) : 4 9 - 9 1 .

R e n n i e , P. J . 1 9 6 6 . A f o r e s t s a m p l i n g p r o c e d u r e f o r n u t r i e n t u p t a k e s t u d i e s . C o m m o n w e a l t h F r , R e v , 4 5 ( 2 ) : 1 1 9 - 2 8 .

R o d i n , L . E . , B a z i l e v i c h , N. I . 1967. P r o d u c t i o n a n d m i n e r a l c y c l i n g i n t e r r e s t r i a l v e g e t a t i o n . O l i v e r a n d B o y d , E d i n b u r g , 1 9 6 7 . ( 2 8 8 p p . ) .

85

R o m e l l , L . G, 1 9 5 7 , Man-made " n a t u r e 1 ' o f n o r t h e r n l a n d s , I n " P r o c e e d i n g s a n d P a p e r s o f t h e S i x t h T e c h n i c a l M e e t i n g , I n t e r n a t i o n a l U n i o n o f C o n s e r v a t i o n o f N a t u r e a n d N a t u r a l R e s o u r c e s , E d i n b u r g , 1 9 5 6 " , L o n d o n , p p . 5 1 - 5 3 ,

R o w e , J , S, 1 9 5 9 . F o r e s t r e g i o n s o f C a n a d a . C a n a d a D e p t . N o r t h e r n A f f a i r s a n d N a t u r a l R e s o u r c e s , F o r e s t r y B r a n c h , B u l l e t i n 1 2 3 , Q u e e n ' s P r i n t e r , O t t a w a , 7 1 p p ,

S m i t h , W. H., L . E . N e l s o n , G. L , S w i t z e r , 1 9 6 3 . T h e c h a r a c t e r i z a t i o n o f d r y m a t t e r a n d n i t r o g e n a c c u m u l a t i o n b y l o b l o l l y p i n e ( P i n u s

-- t a e d a L . ) o n p o o r s i t e s . S o i l S c , S o c . o f Am. P r o c , 2 7 : 4 6 5 - 4 6 8 .

S m i t h , J . H. G. 1 9 7 4 . P e r s o n a l c o m m u n i c a t i o n ,

S m i r n o v a , K. M. 1 9 5 1 . K r u g o v o r o t a z o t a i z o l n y c h e l e m e n t o v v j e l n i k e z e l e n o m o s n i k e . V e s t n i k m o s k o y s k o g o U n i v e r s i t e t a 6 : 6 9 - 8 3 .

S w i t z e r , G. L . , N e l s o n , L . E . , S m i t h , W. I I . 1 9 6 4 . T h e m i n e r a l c y c l e i n f o r e s t s t a n d s . F o r e s t F e r t i l i z a t i o n , T h e o r y a n d P r a c t i c e . T.V.A. T e n n . 1 9 6 4 , p p . 1 -9.

S w i t z e r , G. L . , N e l s o n , L . E . 1 9 7 2 , N u t r i e n t a c c u m u l a t i o n a n d c y c l i n g i n l o b l o l l y p i n e ( P i n u s t a e d a L . ) p l a n t a t i o n e c o s y s t e m s ; t h e f i r s t t x r e n t y y e a r s . P r o c . S o i l S c i . S o c . A m e r . 36 ( l ) : 1 4 3 - 7 .

Tamm, C, 0. 1 9 6 4 . D e t e r m i n a t i o n o f n u t r i e n t r e q u i r e m e n t s o f f o r e s t s t a n d s . I n t e r n a t i o n a l R e v i e w o f F o r e s t r y R e s e a r c h A c a d e m i c P r e s s (New Y o r k ) 1 : 1 1 5 - 1 7 0 .

Tamm, C. 0. 1 9 6 9 , S i t e d a m a g e s b y t h i n n i n g d u e t o r e m o v a l o f o r g a n i c m a t t e r a n d p l a n t n u t r i e n t s , P r o c , I U F R 0 M e e t . , S t o c k h o l m , S e p t e m b e r 1 9 6 9 .

T e c h n i c o n A u t o - A n a l y s e r M e t h o d o l o g y , T e c h n i c o n C o r p o r a t i o n , T a r r y t o w n , New Y o r k ,

T s u t s u m i , T,, K a w a h a r a , T,, S h i d e i , T. 1 9 6 8 . T h e c i r c u l a t i o n o f n u t r i e n t s i n f o r e s t e c o s y s t e m I , On t h e a m o u n t o f n u t r i e n t s c o n t a i n e d i n t h e a b o v e - g r o u n d p a r t s o f s i n g l e t r e e a n d o f s t a n d . J . J a p . F o r . S o c . 5 0 ( 3 ) : 1 6 6 - 7 4 .

86

T u r t o n , A. G,, K e a y , J . 1 9 7 0 . D i s t r i b u t i o n o f b i o m a s s a n d m a j o r n u t r i e n t s i n a m a r i t i m e p i n e p l a n t a t i o n , A u s t . F o r . 34 ( 1 ) : 3 9 ^ 4 8 .

T w i n e , J . R., C, H, W i l l i a m s . 1 9 7 1 , T h e d e t e r m i n a t i o n o f p h o s p h o r o u s i n K j e l d a h l d i g e s t o f p l a n t m a t e r i a l b y a u t o m a t i c a n a l y s i s . Comm. i n S o i l S c i e n c e a n d P l a n t A n a l y s i s , 2 ( 6 ) : 4 8 5 - 4 8 9 .

V a r l e y , J . A. 1 9 6 6 , A u t o m a t i c m e t h o d s f o r t h e d e t e r m i n a t i o n o f N, P, a n d K i n p l a n t m a t e r i a l . A n a l y s t 9 1 : 1 1 9 - 1 2 6 .

V i r o , J . P, 1 9 5 6 , I n v e s t i g a t i o n s o n f o r e s t l i t t e r . Comm. I n s t . F o r e s t . F e n n i a e , 4 5 : 1 - 6 5 ,

V i r o , J , P. 1 9 6 9 . P r e s c r i b e d b u r n i n g i n f o r e s t r y . Comm. I n s t . F o r . F e n n . 67 ( 7 ) .

V o i g t , G. H. 1 9 6 3 . V a r i a t i o n i n n u t r i e n t u p t a k e o f t r e e s . F o r e s t f e r t i l i z a -t i o n , t h e o r y a n d p r a c t i c e . P a p e r s p r e s e n t e d a t t h e s y m p o s i u m o n f o r e s t f e r t i l i z a t i o n , A p r i l 1 9 6 7 , a t G a i n e s v i l l e , F l o r i d a . T e n n e s s e e V a l l e y A u t h o r i t y N a t i o n a l F e r t i l i z e r D e v e l o p m e n t C e n t r e , M u s c l e S h o a l s , A l a . , 1 9 6 8 . 3 0 6 p p .

W e e t m a n , G. F. 1 9 7 1 . E f f e c t o f t h i n n i n g a n d f e r t i l i z a t i o n o n t h e n u t r i e n t u p t a k e , g r o w t h a n d w o o d q u a l i t y o f u p l a n d b l a c k s p r u c e . W o o d l . P o p . P u l p P o p . R e s . I n s t . C o n . 2 8 : 1 8 .

W e e t m a n , G, F . , W e b b e r , B. 1 9 7 2 . T h e i n f l u e n c e o f w o o d h a r v e s t i n g o n t h e n u t r i e n t s t a t u s o f t w o s p r u c e s t a n d s . C a n . J . F o r e s t R e s . 2 : 3 5 1 - 3 6 9 .

W e l l s , C. G, 1 9 6 5 . N u t r i e n t r e l a t i o n s h i p s b e t w e e n s o i l s a n d n e e d l e s o f l o b l o l l y p i n e ( P i n u s t a e d a ) . P r o c . S o i l S c i . S o c . A m e r . 2 9 ( 5 ) : 6 2 1 - 4 .

W i l d e , S, A., V o i g t , G, H., T y e r , J , G. 1 9 6 4 . S o i l a n d p l a n t a n a l y s i s f o r t r e e c u l t u r e . O x f o r d P u b l i s h i n g H o u s e , New D e l h i ,

W i l l , G, M. 1 9 5 7 . V a r i a t i o n s i n t h e m i n e r a l c o n t e n t o f r a d i a t a p i n e n e e d l e s w i t h a g e a n d p o s i t i o n i n t r e e c r o w n . New Z e a l a n d F o r e s t R e s . N o t e 1 1 , R e p t . New Z e a l a n d J o u r . S c i . a n d T e c h n o l . 3 8 ( 7 ) : 1 3 - 2 5 .

W i l l , G. M, 1 9 5 9 , N u t r i e n t r e t u r n i n l i t t e r a n d r a i n f a l l u n d e r some e x o t i c c o n i f e r s t a n d s i n New Z e a l a n d , New Z e a l a n d F o r e s t R e s . N o t e 1 8 , R e p t . New Z e a l a n d J o u r , o f A g , 2 ( 4 ) : 7 1 9 - 7 3 4 .

87

W i l l , G, M, 1 9 6 4 , D r y m a t t e r p r o d u c t i o n a n d n u t r i e n t u p t a k e b y P i n u s r a d i a t a i n New Z e l a n d , Commonw, F o r , R e v , 4 3 (l).:57<-70.

W i l l , G, M, 1 9 6 8 . T h e u p t a k e , c y c l i n g a n d r e m o v a l o f m i n e r a l n u t r i e n t s b y c r o p s o f P i n u s r a d i a t a , P r o c . N. A. E c o l , S o c . 1 5 : 2 0 - 2 4 ,

W i l l i a m s , C. H., J . R, T w i n e . 1 9 6 7 . D e t e r m i n a t i o n o f N, S, P, K, N a , C a a n d Mg i n p l a n t m a t e r i a l b y a u t o m a t i c a n a l y s i s . D i v i s i o n o f P l a n t I n d u s t r y T e c h n i c a l P a p e r N o . 2 4 . C o m m o n w e a l t h S c i e n t i f i c a n d I n d u s t r i a l R e s e a r c h O r g a n i z a t i o n , A u s t r a l i a , M e l b o u r n e , 1 9 6 7 ,

W r i g h t , T. W., G. M. W i l l , 1 9 5 8 * T h e n u t r i e n t c o n t e n t o f S c o t s a n d C o r s i c a n p i n e g r o w i n g o n s a n d d u n e s . F o r e s t r y 3 1 : 1 3 - 2 5 .

Y o u n g , H. E . , L . S t r a n d , R. A l t e n b e r g e r . 1 9 6 4 . P r e l i m i n a r y f r e s h a n d d r y w e i g h t t a b l e s f o r s e v e n t r e e s p e c i e s i n M a i n e . M a i n e A g r i c u l t u r a l E x p e r i m e n t a l S t a t i o n T e c h . B u l l . 1 2 , U n i v e r s i t y o f M a i n e , N o v e m b e r 1 9 6 4 .

Y o u n g , H. E . , P. N. C a r p e n t e r , R. A, A l t e n b e r g e r . 1 9 6 4 . P r e l i m i n a r y t a b l e s o f some c h e m i c a l e l e m e n t s i n s e v e n s p e c i e s i n M a i n e , M a i n e A g r i c u l t u r a l E x p e r i m e n t a l S t a t i o n T e c h . B u l l e t i n 2 0 , U n i v e r s i t y o f M a i n e , O c t o b e r 1 9 6 5 .

Y o u n g , H, E . , G u i n n , V. P, 1 9 6 6 . C h e m i c a l e l e m e n t s i n c o m p l e t e m a t u r e t r e e s o f s e v e n s p e c i e s i n M a i n e . T a p p i 4 9 ( 5 ) . .

Y a n g , H. E , , D y e r , R. F , , D u b e , G. F , 1 9 6 7 . N u t r i e n t d i s t r i b u t i o n i n t h e c r o w n o f p o l e s i z e r e d s p r u c e a n d w h i t e p i n e , M a i n e F a r m R e s e a r c h , O r o n o 1 9 6 7 , ( A p r i l ) , p . 5 .

A P P E N D I X 1

S C I E N T I F I C AND COMMON NAMES OF T R E E S P E C I E S

A b i e s - a m a b i l i s ( D o u g l . ) F o r b e s P a c i f i c s i l v e r f i r

A b i e s l a s i o c a r p a ( H o o k . ) N u t t , A l p i n e f i r

C h a m a e c y p a r i s ribotkatensis ( D . Don). S p a c h Y e l l o w c e d a r

L a r i x l e p t o l e p i s ( S i e b . e t Z u c c ) . G o r d . J a p a n e s e l a r c h

P i c e a a b i e s ( L . ) K a r s t N o r w a y s p r u c e

P i c e a e n g e l m a n i i i i P a r r y E n g e l m a n s p r u c e

P i c e a s i t c h e n s i s ( B o n g . ) . C a r r . S i t k a s p r u c e

P i n u s b a n k s i a n a L a m b . J a c k p i n e

P i n u s c o n t o r t a D o u g l * . L o d g e p o l e p i n e

P i n u s r a d i a t a D. D o n R a d i a t a p i n e

P i n u s s i l v e s t r i s L , S c o t s p i n e

P i n u s t a e d a L . L o b l o l l y p i n e

P s e u d o t s u g a m e n z i e s i i ( M i r b e l ) F r a n c o D o u g l a s - f i r

T h u j a p l i c a t a D. D o n W e s t e r n r e d c e d a r

T s u g a h e t e r o p h y l l a ( R a f . ) S a r g . W e s t e r n h e m l o c k

T s u g a m e r t e n s i a n a ( B o n g . ) C a r r . M o u n t a i n h e m l o c k

A P P E N D I X 2

B A S I C S T A T I S T I C S FOR THE INDEPENDENT V A R I A B L E S

91

P a c i f i c s i l v e r f i r = 7 ; w e s t e r n h e m l o c k =8.

I n d e p e n d e n t V a r i a b l e

T r e e S p e c . U n i t

M e a n V a l u e

S t a n d a r d D e v i a t i o n

M i n i m u m V a l u e

M a x i m u m V a l u e

D 2 . H M.h. P . s . f . W.h.

2 nr-.rrt

12.59 13.83 2,55

6.64 11.81 2.11

4.79 1.71 0.44

19.86 33.67 6.65

BA.H M.h. P . s . f , W.h.

2 irr-.m

9.89 10.86 2.00

5.21 9.28 1.65

3.76 1.34 0.34

15.60 26.44 5.22

D.CL M.h. P . s . f . W.h.

m.m 14.55 14.25 5.13

4.01 8.21 3.15

9.00 3.56 1.88

19.20 25.67 11.37

D . C L 2 M.h. P . s . f . W.h.

m.m2 353.51 361.81 89.57

116.62 262.08 80.37

185.44 41.00 19.41

483.90 729.13 263.74

D.CW2 M.h. P . s . f . W.h.

2 m.m

30.98 33.09 9.76

24.54 20.70 8.51

9.65 9.38 1.54

67.33 63.79 26.11

D2.H.CW M.h. P . s . f . W.h.

2 m .m.m

93.11 107.86 15.12

71.89 101.02 15.93

22.54 9.41 1.51

186.66 282.82 48.55

(BA.H)+CCW.CL) M.h. P . s . f . W.h.

j

m .rn+m.m 169.42 174.85 82.15

62.96 74.37 46.88

100.58 64.59 29.90

252.48 265.00 169.55

BA.H.CL M.h. P . s . f . W.h.

2 m^.m.m

242.09 285.52 35.93

135.21 273.24 38.97

77.55 15.45 4.20

393.03 751.00 121.18

BA M.h. P . s . f . W.h.

™2 m 0.296 0.304 0.087

0.127 0.210 0.058

0.150 0.075 0.020

0.456 0.642 0.189

cw2' M.h. P . s . f . W.h.

m.m 46.68 52.10 27.10

28.56 18.44 15.98

22.59 30.25 8.41

88.36 72.25 53.29

CR.VOL M.h. P . s . f . W.h.

m 3 299.58 321.99 123.47

194.54 172.49 102.84

119.13 91.07 22,68

582.94 546.64 323.67

CR.SURF M.h. P . s . f . W.h.

m 2 253.39 260.98 130.67

93.46 104.28 74.59

153.07 102,15 47.38

378.51 390.01 269.30

A P P E N D I X 3

S I G N I F I C A N T INDEPENDENT V A R I A B L E S I N S I M P L E L I N E A R

AND L O G A R I T H M I C R E G R E S S I O N EQUATIONS FOR BIOMASS

P R E D I C T I O N OF P A R T I C U L A R T R E E COMPONENTS

S i g n i f i c a n t i n d e p e n d e n t v a r i a b l e s i n s i m p l e l i n e a r r e g r e s s i o n e q u a t i o n s f o b i o m a s s p r e d i c t i o n o f p a r t i c u l a r c o m p o n e n t s ( v a r i a b l e s a p p e a r i n o r d e r o f d e c r e a s i n g r 2 v a l u e s ) , S q u a m i s h s a m p l e p l o t , n ; M o u n t a i n h e m l o c k = 5; P a c i f i c s i l v e r f i r = 7,

T r e e S E E C o m p o n e n t T r e e I n d e p e n d e n t R e g r e s s , i n

( v a r i a b l e Y ) S p . V a r i a b l e X. I n t e r c e p t C o e f f , r 2 k g

w h o l e t r e e M.h. D 2 . H 3 6 6 . 4 8 4 1 3 1 . 6 6 7 9 9 . 7 4 5 2 b i o m a s s D - 1 9 0 4 . 5 5 0 6 5 2 6 , 1 3 0 9 8 . 0 1 1 4 3

P . s . f . D Z . H 1 7 6 . 5 6 7 1 2 8 . 1 3 9 9 9 . 2 6 1 4 3 D - 2 0 2 9 . 0 5 0 6 7 7 5 . 6 4 0 9 7 . 5 3 2 6 1

w o o d M.h. D 2 . H 2 6 6 . 1 6 0 8 6 . 2 9 2 9 9 . 0 5 65 b i o m a s s D - 1 1 9 4 . 9 5 0 4 2 3 1 . 7 8 0 9 5 . 2 9 1 4 4

B A ' 4 2 . 5 3 8 4 4 3 0 . 2 2 0 9 5 . 1 2 1 4 7 P . s . f . D 2 . H 8 0 . 8 2 4 9 0 . 4 7 8 9 9 . 5 8 76

B A - 2 1 3 . 4 9 7 5 0 8 9 . 5 2 0 9 9 . 4 6 86 D - 1 4 6 7 . 7 2 0 4 7 6 9 . 2 1 0 9 7 . 2 3 1 9 5

b a r k M.h. B A - 1 5 . 1 9 3 1 2 8 2 . 7 4 0 9 8 . 7 0 2 2 b i o m a s s D - 3 6 9 . 5 1 9 1 2 1 8 . 6 7 0 9 7 . 8 0 28

D 2 . H 5 8 . 3 5 9 2 4 . 2 8 6 9 7 . 1 0 32 P . s . f . D 2 . H 1 7 . 7 5 3 1 8 . 7 6 3 9 6 . 1 2 4 9

B A - 4 2 . 3 2 4 1 0 5 2 . 2 8 0 9 5 . 4 3 5 3

b i o m a s s o f M.h. D2.11.CW 2 2 . 0 4 8 1 . 3 7 8 9 9 . 2 3 1 0 b i g b r a n c h e s D.CW2 2 5 . 6 3 2 4 . 0 2 7 9 8 . 7 0 13

B A - 7 9 . 4 0 2 7 7 6 . 9 9 3 9 8 . 0 0 16 CR.VOL - 0 . 5 8 6 0 . 5 0 4 9 7 . 1 4 19 D - 2 9 0 . 5 1 6 7 3 2 . 3 5 3 9 5 . 5 8 2 4 D 2 . H - 3 2 . 1 7 6 1 4 . 4 9 8 9 3 . 6 5 29

P . s . f . B A - 3 9 . 7 7 0 6 6 3 . 7 7 8 8 1 . 9 3 72 D - 2 0 2 . 4 1 3 6 2 0 . 4 1 1 7 9 . 6 9 76 D 2 . l i 1 . 0 4 1 1 1 . 6 2 5 7 9 . 6 1 76 B A . H . C L 2 2 . 0 2 5 0 . 4 9 0 7 5 . 6 3 83

b i o m a s s o f M.h. n o o n e i n d e p , v a r i a b l e i s s i g n i f i c a n t , s m a l l b r a n c h e s P . s . f . L - 2 8 . 9 4 4 2 . 6 6 0 8 8 . 0 5 9

D - 7 . 5 5 8 1 0 3 . 6 7 5 8 5 . 9 8 1 0 D.CL 1 3 . 6 3 2 2 . 7 8 4 3 5 . 3 3 1 0 ( B A . H ) . + (CL.CW) 0 . 9 5 0 0 . 2 9 9 8 0 . 9 3 1 2 C L - 2 4 . 6 0 9 3.472 8 0 . 4 9 12 CR.SURF - 1 . 8 3 3 0 . 2 1 1 7 9 . 2 4 12

94

T r e e S E E C o m p o n e n t T r e e I n d e p e n d e n t R e g r e s s , i n

( v a r i a b l e Y ) S p . V a r i a b l e X. I n t e r c e p t C o e f f . r 2 k g

b i o m a s s o f M.h, D.CL - 2 0 . 6 3 7 9.679 9 7 . 0 3 8 t w i g s a n d D - 5 1 . 4 4 8 2 8 5 . 1 9 5 9 2 . 1 2 1 3 f o l i a g e B A.H.CL * 5 3 . 0 7 0 0 . 2 7 7 9 0 . 4 2 14

BA 3 3 . 8 9 7 2 9 1 . 9 8 8 8 7 . 9 5 16 D 2 . H 5 0 . 1 8 5 5 . 5 6 4 8 7 . 1 6 16 (BA.H). + (CL.CW) 2 2 . 9 9 5 0 . 5 7 4 8 3 . 9 2 18

P . s . f . D.CL - 2 . 9 9 5 8 . 6 9 6 9 4 . 6 5 19 D - 6 4 . 8 4 9 3 1 6 . 4 1 0 9 1 . 0 9 24 3 A 2 1 . 9 4 7 3 2 5 . 8 5 9 8 6 . 7 8 29 (BA.H): + (CL.CW) - 3 9 . 6 6 1 0 . 9 1 8 8 6 . 5 8 29 C L - 1 1 8 . 1 0 7 1 0 . 6 5 1 8 6 . 0 8 3 0 BA.H.CL 5 0 . 4 5 0 0 . 247 8 4 . 9 3 32

b i o m a s s o f M.h. D.CL - 1 8 . 7 8 0 4 . 7 0 0 9 9 . 2 8 2 t w i g s D - 3 4 . 1 5 2 1 3 9 . 1 5 7 9 5 . 1 8 5

BA.H.CL 1 6 . 6 6 2 0 . 1 3 6 9 4 . 4 8 5 ( B A . H ) + (CL.CW) 0 . 1 4 3 0 . 2 9 2 9 4 . 2 9 5 CR.SURF 0.027 0 . 1 9 6 9 3 . 3 1 6 D 2 . H * 1 5 . 2 8 6 2 . 7 2 7 9 1 . 3 9 6

b i o m a s s o f M.h. D.CL * - 1 . 8 5 7 4 . 9 8 0 8 8 . 1 6 8 f o l i a g e D * - 1 7 . 2 9 6 1 4 6 . 0 4 0 8 2 . 9 1 1 0

BA.H.CL * 3 6 . 4 0 8 0 . 1 4 1 8 0 . 5 2 1 1 D 2 . H * 3 4 . 8 9 9 2 . 8 3 7 7 8 . 2 4 1 1

* s i g n i f i c a n t o n l y a t 5% l e v e l , o t h e r f i g u r e s s i g n i f i c a n t a t 1 % l e v e l .

95

S i g n i f i c a n t i n d e p e n d e n t v a r i a b l e s i n s i m p l e l o g a r i t h m i c r e g r e s s i o n e q u a t i o n s f o r b i o m a s s p r e d i c t i o n o f p a r t i c u l a r t r e e c o m p o n e n t s ( v a r i a b l e s a p p e a r i n o r d e r o f d e c r e a s i n g r 2 v a l u e s ) , S q u a m i s h s a m p l e p l o t , n ; M o u n t a i n hem-l o c k = 5; P a c i f i c s i l v e r f i r = 7,

T r e e C o m p o n e n t T r e e I n d e p e n d e n t R e g r e s s , S E E

( v a r i a b l e &) S p . V a r i a b l e X. I n t e r c e p t C o e f f . r 2 ( l o g )

l o g w h o l e M.h. l o g D 2 . H 2 . 4 6 9 0.769 9 9 . 1 7 0 . 0 2 0 8 3 t r e e l o g D 3 . 7 3 3 2 . 0 1 6 9 8 . 6 6 0 . 0 2 6 4 1 b i o m a s s P . s . f . l o g D 2 , H 2 . 2 7 9 0 . 8 9 9 9 9 . 6 5 0 . 0 2 8 2 8

l o g D 3 . 7 7 9 2 . 4 7 3 9 8 . 9 5 0 . 0 4 8 6 0

l o g w o o d M.h. l o g D 2 . H + 2 . 3 1 9 0. 7 4 6 9 8 . 4 3 0 . 0 2 7 8 7 b i o m a s s l o g D 3 . 5 4 4 1.947 9 7 . 1 3 0 . 0 3 7 6 5

P . s . f . l o g D 2 . H + 2. 0 4 7 0 . 9 5 3 9 9 . 5 3 0 . 0 3 4 4 1 l o g D 3 . 6 3 6 2 . 6 1 8 9 8 . 6 1 0 . 0 5 9 4 3

b a r k M.h. l o g BA + 3.109 1.039 9 8 . 1 8 0 . 0 3 1 8 9 b i o m a s s l o g D 3. 0 0 0 2 . 0 7 8 9 8 . 1 7 0 . 0 3 1 9 0

l o g D S H 1 . 7 0 1 0 . 7 8 8 9 7 . 5 9 0 . 0 3 6 6 2 P . s . f . l o g D 2 . H 1.346 0 . 967 9 5 . 8 3 0 . 1 0 6 4 4

l o g BA + 3 . 0 9 6 1.327 9 4 . 6 5 0 . 1 2 0 5 9 l o g D 2.95 7 2 . 6 5 4 9 4 . 6 5 0 . 1 2 0 5 9

b i g M.h. l o g D2.H.CW + 0.647 0 . 7 8 5 9 9 . 3 8 0 . 0 2 7 2 9 b r a n c h e s l o g D 2 . 8 0 0 3 . 0 7 4 9 9 . 1 0 0 . 0 3 2 9 1 b i o m a s s D.CL 1.017 0 . 0 7 4 9 7 . 8 3 0 . 0 5 1 2 4

D 2 . H 1.537 0 . 0 4 5 9 7 . 7 4 0 . 0 5 2 2 7 l o g D2.H 0.89 4 1 . 1 5 1 9 5 . 9 8 0 . 0 6 9 6 8 ( B A . H ) + (CL.CW) 1 . 2 9 1 0 . 0 0 5 9 4 . 7 1 0 . 0 7 9 9 8 CR.SURF 1.302 0 . 0 0 3 9 4 . 6 9 0 . 0 8 0 1 5

P . s . f . l o g D 2.6 6 5 2 . 4 9 3 9 1 . 9 0 0 . 1 4 1 5 3 D.CL + 1.269 0 . 0 5 3 9 0 . 3 0 0 . 1 5 4 8 5

l o g D 2 , H 1.169 0 . 8 9 0 8 9 . 3 3 0 . 1 6 2 4 3 D . C L 2 1 . 4 3 3 0 . 0 0 2 8 6 , 8 4 0 . 1 8 0 3 7

l o g D2.H.CW 0 . 6 3 1 0 . 7 7 1 8 6 , 1 3 0 . 1 8 5 1 7

l o g s m a l l b r a n c h e s b i o m a s s

M.h. n o o n e i n d e p e n d e n t v a r i a b l e i s s i g n i f i c a n t

P . s . f , l o g D.CL + 0 . 8 6 1 0 . 7 6 0 9 5 . 0 6 0 . 0 5 8 7 5 l o g D . C L 2 0 . 4 2 3 0 . 5 2 1 9 4 . 5 2 0 . 0 6 1 8 7 l o g D 2 , H 1.218 0 . 4 8 1 9 2 . 3 8 0 . 0 7 2 9 9 l o g D2.H.CW 0.9 1 8 0 . 4 2 1 9 1 . 1 0 0 . 0 7 8 8 9 l o g D 2. 0 1 9 1.317 9 0 . 6 9 0 . 0 8 0 6 6 l o g CR.SURF - 1 . 0 2 2 1 . 1 3 3 9 0 . 0 0 0 . 0 8 3 6 2

96

T r e e C o m p o n e n t T r e e I n d e p e n d e n t R e g r e s s . S E E

( v a r i a b l e Y ) . S p . V a r i a b l e X, I n t e r c e p t C o e f f . r 2 ( l o g )

l o g t w i g a n d M.h. l o g D 2 . H + D 2 H + 0 . 8 8 5 0 . 1 5 6 9 9 . 3 9 0 . 0 1 8 8 9 f o l i a g e l o g D.CL - 0 . 5 4 2 1.029 9 7 . 1 6 0 . 0 3 3 3 3 b i o m a s s l o g D.CL 0,567 1.297 9 5 . 9 3 0 . 0 3 9 9 2

l o g D * 2 . 4 3 4 1 . 6 5 1 8 8 . 3 1 0 . 0 6 7 6 5 l o g D2.H.CW * 1.290 0 . 4 1 5 8 5 . 7 1 0 . 0 7 4 7 8

P . s . f . l o g D.CL + 0.879 1 . 0 3 8 9 7 . 4 1 0 . 0 5 7 3 9 l o g D . C L 2 0 . 2 7 6 0 . 7 1 3 9 7 . 2 4 0 . 0 5 9 2 8 l o g D 2 . H 1.374 0 . 6 4 8 9 2 . 0 2 0 . 1 0 0 8 0 l o g D 2.457 1.789 9 1 . 9 8 0 . 1 0 1 0 1 l o g D2.H,CW 0 , 9 8 3 0 . 5 6 1 8 8 . 6 2 0 . 1 2 0 3 4

l o g t w i g M.h. l o g D . C L 2 + - 1 . 2 7 2 1 . 1 6 3 9 9 . 4 4 0 . 0 1 6 5 9 b i o m a s s l o g D.CL - 2 , 7 8 0 1.474 9 9 . 2 1 0 . 0 1 9 6 1

l o g D 2.1I.CW+D2,H.CW 0.12 2 - 0 . 0 0 3 9 9 . 0 5 0 . 0 2 6 4 2 l o g D 2 . H 0.909 0.724 9 3 . 9 5 0 . 0 5 4 4 0 l o g D 2. 0 9 9 1 . 8 9 5 9 3 . 0 4 0 . 0 5 8 3 6

l o g f o l i a g e M.h. l o g D 2 . H + 0 . 5 9 8 - 0 . 0 4 5 9 8 . 4 6 0 . 0 2 8 3 9 b i o m a s s l o g D . C L 2 - 0 . 5 3 7 0 . 9 3 6 9 0 . 4 5 0 . 0 5 7 6 8

l o g D.CL 0 . 4 7 8 1. 1 7 6 8 8 . 5 4 0 . 0 6 3 1 9 l o g D 2 . 1 6 8 1 . 4 8 6 8 0 . 2 7 0 . 0 3 2 9 1

+ e q u a t i o n s u s e d f o r c a l c u l a t i n g b i o m a s s o n t h e w h o l e s a m p l e p l o t . * s i g n i f i c a n t o n l y a t 5% l e v e l , o t h e r v a r i a b l e s a r e s i g n i f i c a n t

a t 1 % l e v e l .

97

S i g n i f i c a n t i n d p e n d e n t v a r i a b l e s i n s i m p l e l i n e a r r e g r e s s i o n e q u a t i o n s f o r b i o m a s s ^ p r e d i c t i o n o f p a r t i c u l a r t r e e c o m p o n e n t s ( v a r i a b l e s a p p e a r i n o r d e r o f d e c r e a s i n g r 2 v a l u e s ) . , H a n e y s a m p l e p l o t , n ; W e s t e r n h e m l o c k = 8,

T r e e C o m p o n e n t

( v a r i a b l e Y ) . T r e e S p .

I n d e p e n d e n t V a r i a b l e X. I n t e r c e p t

R e g r e s s . C o e f f , r 2

S E E i n k g

t r e e b i o m a s s W.h. D 2 . H 7 1 . 6 7 8 1 5 5 . 5 2 4 9 6 . 2 2 70

w o o d b i o m a s s W.h. D 2 . H BA

4 8 . 8 5 6 - 9 . 2 3 6

9 0 , 9 5 9 3 3 4 2 . 3 8 0

9 6 . 3 3 9 8 . 1 1

4 0 29

b a r k b i o m a s s W.h. BA - 1 0 . 0 5 4 8 7 9 . 1 4 0 9 3 . 6 9 14

b i o m a s s o f W.h. D.CL - 2 7 . 1 4 7 1 3 . 1 6 1 9 1 . 5 5 14 b i g b r a n c h e s D2.H.CW 1.759 2 . 5 5 4 8 7 . 9 7 16

BA.H.CL 2 . 9 1 9 1.042 8 7 . 6 4 16 CR.VOL - 8 . 7 1 7 0 . 3 9 8 8 8 . 7 8 16 CR.SURF - 3 0 . 6 4 6 0 . 5 4 4 8 7 . 2 9 17 CL - 9 9 . 3 1 0 9.100 8 0 . 7 8 21

b i o m a s s o f W.h. L - 7 2 . 4 9 3 5 . 6 4 7 8 7 . 3 9 1 0 s m a l l D.CL 8.504 7.119 7 5 . 9 7 14 b r a n c h e s D 2 . H + CL - 1 8 . 7 8 0 3 . 5 6 5 7 2 . 6 9 15

CR.SURF 6.726 0 . 2 9 8 7 2 . 0 0 15 BA.H 1 8 . 8 0 7 1 3 . 1 0 1 7 0 . 7 2 15

b i o m a s s o f W.h. L - 8 2 . 7 5 9 5 . 6 9 5 9 3 . 0 0 7 t w i g s a n d D.CL - 2 . 6 0 1 7 . 4 8 0 8 7 . 7 4 1 0 f o l i a g e CW 2 - 3 . 7 5 8 1 . 459 8 5 . 5 8 1 0

BA.H 7.680 1 4 . 0 3 9 8 4 . 9 6 1 1 CR.VOL 8. 3 8 5 0 . 2 2 2 8 2 . 0 1 12 CR.SURF - 4 . 1 9 7 0 . 3 0 6 8 2 . 0 4 12

b i o m a s s o f W.h. CW 2 - 0 . 6 2 1 0 . 4 8 6 9 0 . 8 7 3 t w i g s ( B A . H ) + (CL.CW) - 0 . 6 1 8 0 . 1 6 0 8 5 . 1 2 3

CR.VOL 3 . 4 6 8 0.074 8 6 . 2 3 3 CR,SURF - 0 . 6 3 3 0 . 1 0 1 8 5 . 3 6 3 L - 2 3 . 7 5 1 1 . 7 4 4 8 3 . 4 3 4 D.CL 0 . 7 7 2 2 . 2 9 6 7 9 . 0 5 4

b i o m a s s o f W.h. L - 5 8 . 9 0 7 3 . 9 4 6 9 3 . 9 0 5 f o l i a g e D.CL - 3 . 3 4 0 5 . 1 7 6 8 8 . 3 6 6

BA.H 3 . 3 4 0 5 . 1 7 6 8 7 . 1 3 7 B A - 1 . 0 0 2 2 7 9 . 2 1 8 8 6 . 4 4 7 CR.SURF - 3 . 5 3 0 0 . 2 0 5 7 7 . 2 4 9 ( B A . H ) + (CL.CW) - 3 . 5 1 5 0 . 3 2 5 7 7 . 1 3 9

98

S i g n i f i c a n t i n d e p e n d e n t v a r i a b l e s i n s i m p l e l o g a r i t h m i c r e g r e s s i o n e q u a t i o n s f o r b i o m a s s p r e d i c t i o n o f p a r t i c u l a r t r e e c o m p o n e n t s ( v a r i a b l e s a p p e a r i n o r d e r o f d e c r e a s i n g r 2 v a l u e s ) . , H a n e y s a m p l e p l o t , n ; W e s t e r n h e m l o c k = 8,

T r e e C o m p o n e n t T r e e I n d e p e n d e n t R e g r e s s , S E E

( v a r i a b l e Y ) S p . V a r i a b l e X I n t e r c e p t C o e f f . r 2 ( l o g )

l o g w h o l e W.h. l o g D 2 . H 2.32 7 0 . 8 8 2 9 8 . 9 1 0 . 0 4 3 0 9 t r e e l o g D 3. 6 8 0 2 . 1 3 5 9 6 . 7 3 0 . 0 7 4 4 4 b i o m a s s

l o g w o o d W.h. l o g D 2 , H + 2 , 1 1 2 0 . 8 7 0 9 9 . 2 5 0 . 0 3 5 1 5 b i o m a s s l o g D 3. 4 5 5 2 . 1 2 0 9 8 . 3 2 0 . 0 5 2 5 7

l o g b a r k W.h. l o g D 2 , H + 1 . 4 1 5 0 . 9 9 6 9 7 . 2 1 0 . 0 7 8 4 2 b i o m a s s l o g D 2. 9 3 4 2 . 3 9 4 9 3 . 7 3 0 . 1 1 7 4 8

l o g BA 3. 0 6 0 1.197 9 3 . 7 3 0 . 1 1 7 4 8

l o g b i g W.h. l o g D + 3 . 2 7 0 3 . 8 6 8 7 7 . 8 3 0 . 3 9 1 7 1 b r a n c h e s l o g D 2 . H 0. 8 2 5 1 . 5 7 0 7 6 . 9 4 0 . 3 9 9 5 3 b i o m a s s l o g D2.H.CW 0,07 2 1 . 2 3 1 7 5 . 4 0 0 . 4 1 2 6 6

l o g s m a l l W.h. l o g D2.H.CW + 1.177 0 . 4 9 9 9 2 . 7 2 0 . 0 8 2 1 4 b r a n c h e s l o g D.CL 0. 9 3 6 1.007 9 1 . 1 9 0 . 0 9 1 3 2 b i o m a s s l o g D 2 . H 1.427 0 . 6 2 0 8 9 . 6 0 0 . 0 9 8 1 4

l o g D . C L 2 0 . 3 1 7 0 . 697 8 6 . 2 4 0 . 1 1 2 8 8 l o g D 2 . 3 6 6 1.477 8 4 . 8 3 0 . 1 1 8 5 3 l o g CR.VOL 0.406 0 . 6 0 2 6 9 . 4 2 0 . 1 6 8 3 0

l o g t w i g a n d W.h. l o g D.CL + 0 . 6 2 8 1 . 2 7 2 9 5 . 5 8 0 . 0 7 8 9 2 f o l i a g e l o g D2.H.CW 0 . 8 6 3 0 . 6 2 6 9 5 . 5 1 0 . 0 7 9 6 1 b i o m a s s l o g D . C L 2 - 0 . 1 6 5 0 . 8 8 8 9 1 . 7 3 0 . 1 0 8 0 1

l o g D 2 . H 0. 1 2 5 0 . 7 7 4 9 1 . 8 0 0 . 1 0 7 5 5 l o g D 2 . 4 2 0 1.837 8 6 . 1 3 0 . 1 3 9 9 1 l o g CR.VOL - 0 . 0 5 3 0 . 7 6 7 7 4 . 0 0 0 . 1 9 1 5 4

l o g t w i g W.h. ( B A . H ) . + (CL.CW). + 0.564 0. 0 0 6 8 2 . 8 1 0 . 1 2 7 3 0 b i o m a s s l o g C R.SURF 0 . 5 6 3 0 . 0 0 3 8 3 . 0 2 0 . 1 2 6 5 0

CR.VOL 0 . 7 0 8 0 . 0 0 3 . 8 2 . 1 3 0 . 1 2 9 7 9 l o g D . C L 2 - 0 . 2 1 3 0 . 6 7 9 8 0 . 3 9 0 . 1 3 5 9 7 l o g D.CL 0.42 2 0 . 9 3 1 7 6 . 6 0 0 . 1 4 8 5 1 l o g D2.H.CW 0 . 5 6 0 0 . 4 5 1 7 4 . 2 1 0 . 1 5 5 9 1

l o g f o l i a g e W.h. l o g D.CL+D.CL + - 0 . 1 5 0 - 0 . 1 4 4 9 7 . 4 5 0 . 0 8 6 3 3 b i o m a s s l o g D 2 . H 0 . 9 5 1 1 . 0 2 2 9 2 . 4 3 0 . 1 3 5 2 8

l o g D2,H.CW 0.455 0 . 807 9 1 . 9 0 0 . 1 4 0 4 9 l o g D 2. 5 0 8 2 . 4 5 4 8 8 . 9 8 0 . 1 6 3 9 4 l o g D . C L 2 - 0 . 8 1 6 1.114 8 3 . 6 5 0 . 1 9 9 6 7

+ e q u a t i o n s u s e d f o r c a l c u l a t i n g b i o m a s s f o r t h e w h o l e s a m p l e p l o t . A l l v a r i a b l e s s i g n i f i c a n t a t 1 % l e v e l .

A P P E N D I X 4

BIOMASS D I S T R I B U T I O N FOR 2 0 SAMPLED T R E E S ON THE

SQUAMISH SAMPLE PLOT

S P E C I E S 1 - MOUNTAIN HEMLOCK

S P E C I E S 2 - P A C I F I C S I L V E R F I R

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TREE SPECIE AGE LENGTH LENGTH LENGTH DBH CROWN CROWN BUTT BRANCH BRANCH CONES TWIGS FOLIAGE _TWIG*FOL NO TO 1 TO 6 OIAM LENG LENG OVER I IT01/4

(H) (M) (M) (M) (M) (M) (M) (KG) (KG) (KG) (KG) (KG) (KG)

22 440 37,3 37,0 33.6 0,780 7,5 25,9 11,4 411,4 66,4 0,6 0,0 0,0

TOP —1_TWIGS ANO FOLIAGE _ MID —2 ARE NOT SEPARATED LOW — 3 FOR ABIES AMAB1LIS

197,6

56,6 " 5 8 . 1 '

83.0

SECTION N0| 1 2 J 4 5 6 7 a 9 10

WOOD 446,4 383,4 339,0 290,3 233,2 186,0 164,5 125,1 31,6 6.1 SUM WOODS 2205,8 KG

BARK 75,4 47,1 42,2 41,0 35,1 49,8 53.1 41.5 18,2 » » * SUM BARK* 405,S KG TREE BIOMASS" 73 3;03f3 KG

TREE SPECIE AGE LENGTH LENGTH LENGTH DBH CROWN CROWN BUTT BRANCH BRANCH CONES NO TO 1 TO 6 DIAM LENG LENG OVF.R 1 1T01/4

. ( M j

TWIGS FOLIAGE TWIG*FOL

(M)

J i l l . ITO rsy 14,7 0,356

TMT

6.2 41 300

(M)

t«».2

(M)

2.7

(KG)

33,2

(KG)

31.4

(KG)

0,2

(KG)

0,0

TKG)

0,0

(KGT

57.7

TOP--1 TWIGS AND FOLIAGE MID—2 ARE NOT SEPARATED LOW —3 FOR ABIES AMABILIS

9,6 29,3 18,8

SECTION" NO I 1 - 2 - — j j j - 5 - 6 r fl 9 _

WOOD 63,6 57,5 50,8 43.6 29,4 24,7 13.7 8,1 3,7

" 10"

1.1 SUM WOOOs 296,2 KG

BARK 6,6 9,4 9,7 5,9 4,1 3,9 4,2 1,7 0,5 0,2 SUM BARKs 46,3 KG TREE BIOMASSs ' *t&S$Q KG

TREE SPECIE ACE LENGTH LENGTH LENGTH OBH CROWN CROWN BUTT BRANCH BRANCH CONES _TWIGS_ FOLIAGE TWIG*FOL NO TO 1

(M) (H) TO 6 (H) (M)

OIAM LENG LENG (M) CM) CM)

OVER I" 1T01/4 (KG) (KG) (KG) (KG) (KG) (KG)

45 1 SOO 35,2 23,9 30,3 0.754 0,0 29,2 6,0 0,0 0,0 0,0

TOP--1

0,0

0,0

0.0

0,0

0,0

0,0 MID--2 LOW—.3

0,0 0,0

0,0 0,0

0,0 0.0

SECTION NOl 1 2 3 4 5 6 7 8 9 10

MOOD 544,1 443,7 362,5 288,8 221,0 169,3 120,3 63,0 41,4 8,6 SUM WOODS 2282,7 KG

BARK 65,7 102,6 81.0 67.5 42.6 30.0 30,9 16,2 7,8 1,7 SUM BARKs 465,9 KG TREE BIOMASS8 2748,6 KG

' f t * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * *

TREE SPECIE AGE LENGTH LENGTH NO TO I

LENGTH OBH TO 6

CROMN CROWN BUTT DIAM LENG LENG

BRANCH BRANCH OVER 1 1T01/4

CONES TWIGS FOLIAGE TWIG»FOL

(H) CM) CM) (M) CM) (M) CM) (KG) (KG) (KG) (KG) (KG) (KG)

55 1 460 34,A 33,6 2B,9 0',681 6,0 28,2 6,2 0,0 0,0 0,0 0,0 0,0 0,0

TOP—-1 MID--2

0,0 0,0

0,0 0,0

0,0 0,0

LOW--3 0,0 0,0 0,0

SECTION NOl 1 2 3 4 5 6 7 8 9 "to

NOOO 415,4 357,5 269,6 227,4 198,3 135,9 88,3 60,1 22,3 5,2 SUM WOOD» 1779,9 KG

BARK 94,0 68,1 74,6 62,3 37,7 34,2 23,9 15,4 4,7 1,1 SUM BARKa 436,0 KG TREE BIOMASSs 2216.0 KG

•»«**«»«*»«***»*»****»*»»*»***»«**««»»»»**********«»****»**»*******«^

e TREE SPECIE AGE LENGTH LENGTH LENGTH OBH CROWN CROWN BUTT BRANCH BRANCH CONES TWIGS FOLIAGE TWIG+FOL

9

NO TO 1 CM) (H)

TO 6 (M) (M)

DIAM " LENG LENG CM) CM) (M)

OVER 1 IT01/4 (KG) CKG) (KG) (KG) (KG) (KG)

69 1 325 21,3 20,7 14,7 0,320 0,0 12,4 9,0 0,0 0,0 0,0 0,0 0,0 0.0

© TOP--1 0,0 0,0 0,0 MI0--2 LOW—3

0,0 0,0

0,0 0,0

0,0 0,0

© SECTION NOl 1 2 3 4 5 6 7 8 9 10

MOOD 66,4 53,7 41,8 36,7 29,8 22,1 17,8 11,0 5,B 1,6 SUM WOODa 287,0 KG

0 BARK 9,2 13.7 10.0 8.3 6.1 5.0 3,2 1,6 0,6 0,3 SUM BARKa 58,0 KG TREE BIOMASSa 345,0 KG

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o TREE SPECIE AGE LENGTH LENGTH NO TO 1

LENGTH DBH TO 6

CROWN CROWN BUTT DIAM LENG LENG


CONES TWIGS FOLIAGE TWIG+FOL

(M) (M) CM) (M) CM) CM) CM) (KG) (KG) (KG) (KG) (KG) (KG)

o 66 1 530 38,0 36,9 33,4 0,711 7,9 24,5 13,5 217,5 31,3 0,0_ 64,2 84,0 148,2

0 TOP —1 MID — 2

14,3 11,8

16,6 15,5

30,8 27.3

o

LOW — 3 38,1 51.9 90,1

SECTION NOl 1 2 3 4 5 6 7 8 9 10

•J MOOD 380,9 361,2 316,0 260,6 227,4 185,7 129,5 83,7 39,4 9,3 SUM WOODS 1993.7 KG

BARK 64,2 76,8 86,3 92,8 60,5 36,7 30,4 22,9 15.4 5,3 SUM BARKs 491,4 KG TREE BIOMASSS KG

•»*»»»»»*»*»**•*»««•«***»«»«»*»**«»***»»«******«****«*********•**

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<* TREE SPECIE AGE LENGTH LENGTH LENGTH OBH CROWN CROWN BUTT BRANCH BRANCH CONES TWIGS ; FOLIAGE TWIG*FOL NO TO 1

(M) (M) TO 6 DIAM

(M) (M) (M) LENG

(M) LENG OVER 1 "1T01/4

CM) CKG) CKG) (KG) CKG) (KG) • CKG)

7a I 410 34,2 33,4 28,6 0,762 9,4 25.2 9,0 289,6 53,8 0,0 70.1 90,5 160,6

TOP—1 32.9 30,9 63.8 MID—2 LOW — 3

17.9 19.3

23.7 35,9

41,6 55.2

SECTION NOI 1 2 3 4 5 6 7 8 9 10

WOOD 449,5 359,0 292,6 256,6 221.9 143,6 107,5 70,7 22 ,7 3,6 SUM WOOOn 1927,7 KG

BARK 93,4 117,5 108,7 80,2 39,2 65,1 44,9 19,3 5 ,5 1,2 SUM BARKs 575,0 KG TREE BIOMASSs -SOOb,£ KG

ft**********,.,*,*.****,.*,**!)******,*•**«***,•*•**«»*** A********.******************,*********************,,*,,•****•*•••*•**•<••*<*

TREE SPECIE AGE LENGTH LENGTH NO TO 1

LENGTH OBH CROWN TO 6 DIAM

CROWN LENG

BUTT BRANCH BRANCH LENG OVER 1 1T01/4

CONES TWIGS FOLIAGE TWIG*FOL

(M) CM) (M) (M) (H) CM) CM) CKG) CKG) CKG) CKG) CKG) (KG)

80 1 480 31,2 30,5 2 5 , 1 0 . 5 6 4 6.2 25,8 .5,5 1 0 7 , 1 3 5 , 5 0,0 51,6 70,1 121.7

TOP —1 MID—2

16,4 25,2

17,2 35,4

33.6 60,6

LOW — 3 10,1 17.S 27,5

SECTION NO 1 1 2 3 4 s ^ ? * 8 9 10

• » HOOD 260,5 216,2 174,5 135,4 103.2 75,4 55,8 26,3 10 ,5 2,1 SUM WOODs 1059,9 KG

BARK 55,2 62,9 38,9 37,4 21,4 19,8 25,6 11,3 4 ,0 1,2 SUM BARKs 277,5 KG TREE BIOMASSs

**»******»******»***«**»************************»***********^

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TREE SPECIE AGE LENGTH LENGTH LENGTH_ OBH CROWN CROWN BUTT BRANCH_ BRANCH CONES TWIGS FOLIAGE TWIG*FOL_ NO TO 1 TO 6 "' ' ~ D I A M LENG LENG OVER 1 1T01/4

(M) (M) (M) (M) (M) (M) (M) (KG) (KG) (KG) (KG) (KG) (KG)

I 19 1 410 25.1 2«t» 21,5 0,437 4,7 20,6 4,5 53.1 23,8 0,0

TOP — 1 MID. LOW-

23.6

_S.6_ 10,5 7,4

35.5

10.9 1 7 , 6 "

7.0

59,1

16,5 "28,2" 1«,5

SECTION NOI 1 10

WOOD 147,1 123,9 107,4 91,4 77,3 60,2 45,9 27,8 17,3 3,9 SUM WOOD" 702,2 KG

BARK 32.2 31,5 29.8 26.5 18,1 iq,8 U . 7 7.2 5.1 1,2 SUM BARK" 178,1 KG TREE BIOMASS" fO/C',3 KG

***************************************************

TREE SPECIE AGE LENGTH LENGTH LENGTH OBH CROWN CROWN BUTT BRANCH BRANCH CONES NO TO 1 TO 6 DIAM LENG LENG OVER 1 ITO 1/4


159 018

(M)

29,9

(M)

29,0

(M) (M)

26,1 0,485

(M)

8,1

(M) (M)

18,1 11,8

(KG)

53,3

(KG)

41,9

(KG)

TOP-MID"

(KG)

0,0

(KG)

0,0

•1 TWIG3 AND FOLIAGE •2 ARE NOT SEPARATED

LOW—-3 FOR ABIES AMABILIS

(KG)

59.2

29.2 20.8

9.2

SECTION NO I 1 "2 J 4 5 T " " 8 ~ ~ " 9 " 1 0 " "

WOOD 182,7 151,7 132,4 103.0 87.2 65.6 50.2 34.2 19.7 5,6 SUM WOOQ" 832.2 KG

BARK 37,9 38,3 34,6 28,7 30,8 23,4 16,2 11,0 5,7 2,6 SUM BARK" 229,0 KG TREE BIOMASS" 7-2.767° K<5

**««»««»««»««**««««»»«*«*«***.»**»****«»******««**********************«***«^

TREE SPECIE ACE LENGTH LENGTH LENGTH OBH CROWN_ CROWN BUTT BRANCH_ BRANCH CONES TWIGS NO TO 1 TO 6 DIAM LENG LENG OVER 1 1T01/4

(M) CM) (M) CM) CM) CM) CM) CKG) CKG) CKG) CKG)

FOLIAGE TWIG*F0L_

CKG) CKG)

33,9 0,711 8,5 28,9 9,6 174,0

ci

175 530 38,4 38,0 73,6 0.3 0,0 0,0

TOP—1 TWIGS AND FOLIAGE MID — 2 ARE NOT "SEPARATEO"~ L0W--3 FOR ABIES AMABILI3

SECTION NOt 1 10

191,0

68,0 ~78.5~ 44,5

WOOD 409,3 336,3 286,9 243,5 192,4 148,9 106,4 76,7 37,9 10,2 SUM WOODa 1848,6 KG

BARK 88.5 69,6 59.2 31.1 35.6 19.3 14.5 9.7 6,4 2.4 SUM BARK" 336.4 KG TREE BIOMASS" £6 £ 3 , 9 KG

**************************************************************************

TREE SPECIE AGE LENGTH LENGTH LENGTH OBH CROWN CROWN BUTT BRANCH BRANCH CONES NO TO 1 TO 6 DIAM LENG LENG OVER 1 1T01/4

CM)

TWIGS FOLIAGE TWIGtPOL

176 335

CM)

26,3 25.5

CM) ( M l -

lS,8 0,361

CM)

0,0

CM)

19,6

(M)

6,6

CKG)

0,0

(KG)

0,0

w

0,0

"(KG)

0,0

(KG)

0,0

TOP—1 TWIGS AND FOLIAGE MID—2 ARE NOT SEPARATED LOW — 5 FOR ABIES AMABILIS

(KG 5

0,0

0,0 0,0 0,0

SECTION NO I 8 "10

WOOD 91,9 66.7 58.4 49,8 37.2 30,3 21.7 14.3 7.0 1.6 SUM WOODa 378.9 KG

BARK 14,0 14,1 8,8 7,1 5,7 5,0 2,9 1,9 1,2 0,4 SUM BARKs 61,1 KG TREE BIOMASS" 4 - ; KG

**********************************************************************************************************************************

TREE SPECIE _AGE _ LENGTH LENGTH_LENGTH DBH_ CROWN CROWN _BUTT_ BRANCH BRANCH CONES TWIGS FOLIAGE TWIG*FOL NO TO 1 TO 6 DIAM LENG LENG OVER 1 1T01/4 ~ "" ""


TTB 2 STo 38,9 38,5 32,3 o,88n b~7o 28,7 roTz o76 oTo b~7o : oTo oTo 'oTo

, . T0P--1_TWIGS AND FOLIAGE 0,0 MID — 2" ARE" NOT "SEPARATED 0,0 LOW —3 FOR ABIES AMABILIS 0.0

SECTION NOl 1 2 3 0 5 6 7 8 9 10

S0OO~fj2,6 487,5~449,0 383,8 318,7 25T,TT6lu8HfTr,9 2~3.T if,l WTWoblyi 2931~6 KG :

BARK 136.3 106.2 95.0 89.1 110.7 90.1 65.2 36.4 9,6 1.8 SUM BARK" 740.4 KG TREE BIOMASS" 3672,0 KG

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TREE SPECIE AGE LENGTH LENGTH LENGTH DBH CROWN CROWN BUTT BRANCH BRANCH CONES TWIGS FOLIAGE TWIG+FOL NO , TO 1 TO 6 DIAM LENG LENG OVER 1 IT01/4

(M) (M) (M) fMl CM) (M) (M) (KG) (KG) (KG) (KG) CKGl (KG)

213 2 365 30 «. 0 29,5 _2 4 , l \ , O.S64 5,6 25,1 4,9 92,1 51,3 0,5 0.0 0,0 116,3

TOP—1 TWIGS AND FOLIAGE 32.6 : : ; MID—2 ARE NOT SEPARATED 52,1

LOW —3 FOR ABIES AMABILIS 31,6

" SECTION NOi 1 2 " 3 4 5 6 7 " " 8 9 10 " " " """

WOOD 218,2 174,2 142,3 118,5 88,2 69,9 49.5 27,4 12,7 2,5 SUM WOOD" 903.4 KG ;

BARK 53,7 38,3 25,8 22,7 23,3 15,4 11,7 3,8 3,9 1,4 SUM BARK" 200,0 KG TREE BIOMASS" I3&3t7. . KG

^ : : : : : : )

TREE SPECIE AGE _ LENGTH LENGTH LENGTH OBH CROWN__CROWN BUTT_ BRANCH BRANCH CONES NO TO i TO 6 DIAM LENG LENG OVER i 1T01/4

(M) (M) (M) CM) CM) CM) CM) CKG) (KG) CKG)

TWIG»FOL

CKG)

215 440 36,2 37.7 33.0 0,805 0,0 25,7 12.5 0,0 0.0 0,0 0,0 0,0

TOP-^1 TWIGS AND FOLIAGE "MI6 — 2 ARENOT SEPARATED" L0W--3 FOR ABIES AMABILIS

0,0

0.0 "0,0" 0,0

SECTION NOI 1 2 3 a 5 6 7 8 9 10

WOOD 480,2 299,9 306,5 266,3 216,5 171,0 112,6 87,0 33,9 5.2 SUM WOOD" 1979,0 KG

BARK 125.2 81.0 80.8 77.1 89.6 72,« an.2 34,7 13.0 1.2 SUM BARKs 619,2 KG TREE BIOMASSs 2598,2 KG 0

•»*•••««**»«»»«««»•»*••»•««*««*****«»»•**«*****•»**»***•»***

©

TREE SPECIE AGE LENGTH LENGTH LENGTH DBH CROWN CROWN BUTT BRANCH BRANCH CONES NO TO 1 TO 6 DIAM LENG LENG OVER 1 1T01/4

" CM)"

TWIGS FOLIAGE TWIG*F0L

235 355

CM)

15.1

"W"" 14,6

CM)

12,5 0.343

CM)

0,0

CM)

8.4

CM)

6,6

(KG)

0,0

(KG)

0,0

"(KG)

0,0

(KG)

0,0

TKG")

0,0

T K G F "

0,0

TOP-M TWIGS AND FOLIAGE _MID«-2 ARENOT SEPARATED LOW—-3 t : o f ) ABIES AMABILIS

0,0 0,0 6,0

"SECTION NO! 1 ~r~ "3 W 5 6" 7 8 9"

WOOD 48.2 41.4 36,6 29.8 25.7 22.7 16,2 10.9 8,1 1,5 SUM WOODS 241,0 KG

BARK 8,1 8,1 7,8 7,4 5,2 5,2 3,9 3,8 1,4 0,5 SUM BARKs 51,3 KG TREiE BIOMASSs 292,4 KG

****************

TREE SPECIE AGE LENGTH LENGTH_ LENGTH OBH CROWN_ CROWN BUTT_ BRANCH BRANCH CONES TWIGS FOLIAGE TWJG + FOL NO to i to 6 " O I A M " XENG "CENG" OVER i ¥Toi/a "

(M) (M) (M) (M) (M) CM) (M) CKG) (KG) CKG) CKG) CKG) CKG)

243 380 J7.8 17.1 11,9 0,310 5,5 11,5 6,3 33,8 17,9 0,0 0,0 0,0

TOP —1_TWIGS AND FOLIAGE MID—2 ARE NOT SEPARATED LOW--3 FOR ABIES AMABILIS

30,0

6,B_ 17,8 5.4

SECTION NOI 1 8 10

WOOD 40,0 36,3 27,7 22,6 16,5 12,7 9,3 5,6 3,5 1,1 SUM WOOD* 175,4 KG

BARK 5.5 7.0 6.8 5.8 4,8 3.1 2,1 1,4 0,7 0.3 SUM BARK" 37,5 KG TREE BIOMASS" KG

**********************************************************************************************************************************

TREE SPECIE AGE LENGTH LENGTH LENGTH OBH CROWN CROWN BUTT BRANCH BRANCH CONES NO TO 1 TO 6 DIAM LENG LENG OVER I IT01/4

256 403

CM) CM)

27,9 27,5_

TM) CMT^

19,5 0,475

CM)

0i°__

(M)

20,3

CM)

7.6

CKG)

0,0

(KG)

0,0

(KG)

0,0

TWIGS

TKGT

0,0

FOLIAGE TWIG*FOL

CKG)

0,0

TOP--1 TWIGS AND FOLIAGE MID—-2 ARE NOT SEPARATED

CKG)

0.0

0.0 0,0 LOW--3 FOR ABIES AMABILIS 0,0

SECTION Nb i t '"""" 2 3 "a" s -6 — 7 - 8 9 1 0

WOOD 167,5 125,8 103,9 93.6 77.2 54,0 43,0 28,6 10,8 2,7 SUM WOOD* 707,3 KG

BARK 27,6 29,2 29,5 20,2 17,4 10,4 10,1 8,4 1,6 0,7 SUM BARK" 155,1 KG TREE BIOMASS" 862,4 KG

****************************************************************************************************************** ****************

A P P E N D I X 5

BIOMASS D I S T R I B U T I O N FOR 1 3 SAMPLED T R E E S ON THE

HANEY SAMPLE PLOT

S P E C I E S 1 - WESTERN HEMLOCK

S P E C I E S 2 - WESTERN RED CEDAR

S P E C I E S 3 - YELLOW CEDAR

o

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0

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TREE SPECIE ACE LENGTH LENGTH LENGTH DBH CROWN CROWN BUTT BRANCH BRANCH CONES _MO : 10-1 TO 6 DIAM LENG_ LENG„_O.V.ER_l IT017.4

(M) (M) (M) (M) CM) CM) CM) CKG) (KG) CKG)

TWIGS

_S0_ -17 Q 1J..L -15... 2_ 2,7 0.1611 *US 1A.3 2V9- _li*2_ _uuuu_

CKG)

T...Q.-

FOLIAGE

TKT) 3 . 9 -

TWIG*FOL

CKG)

TOP—1 M ID—2 LOW — 3

1.5

1.8

0,8 .2.1. 1.0

2,4 5.8 2.7

SECTION NOl 1 2 3 4 5

w. 0 Q D 15 , A 11 , 2_ 9 . 6 I*7_

BARK 2,5 1,9 1,7 2,2 1,3 1,2

6 7 8 9 10

_4,1 2,4 1,5 -0,5 3UM_-.W_Q.0DB .66,.3___.KG__

0,7 0,6 0,3 0,3 SUM BARK* 12,7 KG TREE BIOMASSs jO%,±' KG

•»«•«•»•«»»»»«*«««*•«,•»•««*»** »*»«***»**».»•*****»*«*«* * * * ** • * »*_*«_**** * *.****.**.•* * * «««»«««»«««««»«««»«*««««»»«« * ********* *_*_**.**._

TREE SPECIE AGE LENGTH LENGTH LENGTH DBH CROWN CROWN BUTT BRANCH BRANCH CONES TWIGS FOLIAGE TWIGtPOL NO TO 1 TO 6 DIAM LENG LENG OVER 1 1T01/4

(M) CM) CM) CM) (M) CM) (M) (KG) (KG) (KG) (KG) (KG) (KG) e 54 1 410 22,4 20,9 14,8 0,394 1.7 14,8 7,6 35,4 52,3 UUUU U . l 28,5 39,6 o

TOP—1 M10—2 LOW — 3

2.1 5,6 3,5

6,2 15,2 7.1

8,2 20,8 10.6

. aECi icm NOJ. i I _1_ . 4 ? 6 _7. 8 9 10

WOOD 77,a 69,7 65,4 60,7 51,6 52,5 36,3 8,9 3,0 0,9 SUM WOOD* 427,0 KG

BARK 16,7 14,6 14,3 11,8 12,4 10,6 7,1 2.1 2.5 0,4 SUM BARKS 92,5 KG TREE BIOMASSs

•»•«•».«.»««•».**.««»»»*»»»««•««..«*****•»•«**«**•**»****»**«»*****«*•*«**«********»*««**«*««**•***«**«««****••«»•««•*.•***«»•***»

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TREE SPECIE AGE LENGTH LENGTH LENGTH _H0 . 1.0-4. _tXL.6.

_6JJ_ aoo

(N>

_2.a,u_

(M) CM)

_44-*6-0l5. J 3 4

OBH CROHN CROWN BUTT BRANCH BRANCH CONES DIAM LENG LENG OVER-1—U.O.I/.«

(M) (M) (M) (M) (KG) (KG) (KG)

TWIGS FOLIAGE TWIG»FOL

_ruu_ -.D.,.0_ _uuuu_

(KG)

O..JL

(KG)

0,0,

T O P " ! TWIGS ANO FOLIAGE _MID»-2._.ARE.NUT...SEP.ARA TED-

LOW—3 FOR CEDAR

(KG)

0...0-

0,0 .0,0

0,0

e>

&

SECTION NOI 1 2 3 4 5 6 7 8 9 10

_K0QD_44.T.*L_J).3.,.9....^ A.8 JUK-WOOD* 612,8 KG ,

BARK 12,9 14,5 11,1 7,0 9,5 0,0 0,0 0,0 0,0 0,0 SUM BARK" 55,0 KG TREE BIOMASS" 667,9 KG

A * * * * * * * * * * * * * * * * » * * * « « * ***_»«_****_***_*_* **Jl*JL* ** i i i ************* ***** ****** *** *.* ****** *.* * +.***±**lL**L*±^**^^^±t-±^^J!^** **••*• ***** **~

TREE SPECIE AGE LENGTH LENGTH LENGTH DBH CROWN CROWN BUTT BRANCH BRANCH H 0 TO I TO 6 DIAM LENG LENG OVER 1 IT01/4

• (M) (M) (M) (M) (M) (M) JMJ OXGJ £J<GJ

CONES

(KG)

TWIGS

(KG)

65 1 340 23,5 22,3 15,7 0^343 6,1 18,5 4,9 80,7 73,4 UUUU

TOP —1 MID—2 LOW—3

17,3

0.3 7.3 5.7

FOLIAGE

(KG)

30,9

7.4 12.7 10.8

TWIG+POL

(KG)

48,2

11.7 20,0

SECIX.QJLJiO.1 l_ MOOD 65,2 72,9 61,6 42,8 34.1 25,6 16.3 10,0 S.tt 1.8 SUM WOOD" 335,7 KG

BARK 15,4 16,5 11.5 10.9 T7o 5^5 W 271 T72 074 SUM BARK" 74,6 KG TREE BIOMASS" KG


LENGTH TO 6

DBH CROWN DIAM

CROWN BUTT LENG LENG

BRANCH BRANCH OVER t 1T01/0

CONES TWIGS FOLIAGE TWIG+POL


tyo | >\h 18,9 t 7, o 10,2 0,221 10,0 0,9 1,? 25,0 mum S 1 13,2 18,1 c

TOP—-1 MIO — 2

2.1 ..2.4

6,2 .5.8...

8.3 8,2 c

LOW--3 0,7 1.2 1.8

c SECTION NOl 1 2 5 0 5 6 • 7 8 9 10

WOOD ? 2 . « 19.0 16.9 10.6 12.5 4-l+0_

2.«

_J3,6 5,2 2,6 0,8 SUM WOODS 1.1-4,1 KG c

BARK 5.9 5,5 0,6 3,1 3.3

4-l+0_

2.« 1.9 1.1 0,7 0,3 SUM BARKS 28,8 KG TREE BIOMASSs o

******************************** c.

c

c TREE SPECIE AGE LENGTH LENGTH NO TO 1

(M) (M)

LENGTH TO 6

(M)

DBH

(M)

CROWN DIAM

(M)

CROWN BUTT LENG LENG

(M) (M)


(KG) (KG)

CONES

(KG)

TWIGS

(KG)

FOLIAGE

(KG)

TWIG+FOL

(KG) e

72 1 290 20,2 23,1 17,0 0,396 6,6 17,3 6,9 56,0 74,2 UUUU 24,0 44,4 68,0 e

T0P--1 MID — 2 LOW —3

5,7 11.1 7,2

8,8 21.3 14.3

10,5 32,0 21,5

SECTION NOl 1 2 3 0 5 6 _J 8 9 10

WOOD 101,8 70,2 60,0 56,1 07,0 33,4 22,2 13,1 6,3 1,3 SUM WOODS 411,4 KG

BARK 28,3 23,8 18,3 10,2 12,8 8,7 7,6 3,6 1,8 0,5 SUM BARKS 119,5 KG TREE BIOMASSs 7.2.% If

• •*»*«***»»****•••»*••*»•**•*••<<**•****•**

u

->

TREE SPECIE AGE LENGTH LENGTH LENGTH DBH TO 6

CROHN OT AM

CROWN BUTT BRANCH BRANCH .LENG. LENG OVER 1 1T01/4

CONES TWIGS FOLIAGE TWIG+FOL

(M) (M) CH) CM) CM) CM) CM) CKG) CKG) CKG) CKG) CKG) CKG)

10* I I f n,o • 0,3 S,*. S,4 16,5 l l l l l l l l 6,4 3,1 9,5

TOP--1 MID — 2

1.5 2.1

0,7 1,0

2.2 3,2

L0W*-3 2.8 1.3 «.2

SECTION NOl I 2 3 0 5 6 7 8 9 10

. HOOD 13.3 12,3. 10,7 9,2_ 7,4 5,5. _4 ,6 .3 , <L 1., 7. 0 . 5 SUM_.WOODS .._ 66,6 ..KG.

BARK 2,2 2,1 1,7 1,0 1.2 1,1 0,7 0,4 0,4 0,4 SUM BARK* 11,2 KG TREE BIOMASS* KG

**«.»•*»*.*#•*«»«»»•.«..«**•**»•**»•««****»******»******•*«**»»**************«»*««*»««*****.•«•«»*•»**««*•«•»••«*««»«•»*«»»*««»«*»


CM) fMl

LENGTH DBH TO 6

CM) CM)

CROWN DIAM ,

CM)

CROWN BUTT BRANCH BRANCH LENG LENG OVER 1 1T01/4

CM) CM) CKG) CKG)

CONES

CKG)

TWIGS

CKG)

FOLIAGE

CKG)

THIG»POL

CKG)

105 1 360 17,2 16,5 7,3 0,325 4,0 10,4 . 6,9 17,7 29,8 UUUU 5.4 13,8 19.2

' TOP--1 MID—2 LOW--3

0,8 1.8 2,6

1.7 5.6 6.5

2.5 7.4 9.3

SECTION NO 1 1 2 3 4 5 6 7 8 9 10

HOOD 52,8 47,6 48,0 43,4 11.2 6,8 5,3 3,9 1,3 0,5 SUM WOOD* 220,6 KG

BARK 6,9 6,9 7,3 6,8 «,3 1.9 1.2 0,7 0,4 0,4 SUM BARK* 36,8 KG TREE BIOMASS*

• • A * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * *

V ; : . : : : : : ;

SECTION NOI t 2 3 4 5 6 7 8 9 10

WOOD 150,6 130.6__ _M.iL _7.5-.5_ 6-4,5 39,7 26,1 _ i _ . , . _ _ 5.8 1...2. - SUM-WOOD* 601,3. . KG

BARK 25,a 33,9 29,5 22.3 15,8 12,4 8,5 3,5 1.6 0,4 SUM BARKs 153,2 KG TREE BIOMASSs

TREE SPECIE AGE LENGTH LENGTH LENGTH OBH CROWN CROWN BUTT BRANCH BRANCH CONES MO T.Q_1 IO_6 DIAM LENG LENG.-_ftV.ER_l 1X01741

(M) (M) (M) CM) (M) CM) CM) CKG) CKG) (KG)


108 -4_ 365 27,7 26.(1 19.6 0,490 7-+_ 23,2 4.6 122.2- J _ U _ - uuuu

(KG)

2".l

(KG)

«L,.9_

(KG)

-X2,_-

TOP--1 -MID--2.. L0W--3

4.5 -10,0-

9.6

11,0 .20,8... 16,1

15,5 30,7 25,8

**********************************************************************************************************************************

http://_M.iL

http://_7.5-.5_

http://LENG.-_ftV.ER_l

1 o

©

o

o

o

o

o

o

0

TREE SPECIE AGE NO

LENGTH LENGTH TO 1

LENGTH TO 6

OBH CROHN DIAM

CROHN BUTT BRANCH LENG LENG OVER 1

BRANCH 1T01/4

CONES TWIGS FOLIAGE S

TWIG*FOL

CHJ CH) CM! - (H)— CM) (Ml CM) CKG) CKG) (KG) CKG) (KG) (KG)

8 J 115 0 17,3 9,9 0,231 5.8 13.5 5,5 4.4 13,7 UUUU 0,0 0,0 15.3

T0P--1 TWIGS AND FOLIAGE MID--2 ARE NOT SEPARATED

3,5 7,8

L0W--3 FOR CEDAR « . l

SECTION NOI 1 i 3 4 5 6 7 6 9 10

WOOD 28,9 24,5 23, 1 18,0 15,5 14,0 9,2 5,0 2.2 0,4 SUM WOOD" 140,9 KG

BARK 2,5 2.« 2. 5 2.1 1.6 1.3 1.3 0,9 0,6 0,3 SUM BARK" 15,6 KG ' TREE BIOMASS"

—OBH"

CM)

""BRANCH" 1T01/4

CKG) '

T K t t a r t e I t » e t LENGTH NO

( H )

LENGTH LENGTH TO 1 TO 6

CM) CM)

—OBH"

CM)

CKUWfT CKUWN B U T T 8 K A N t n DIAM LENG LENG OVER 1

CM) CM) CM) CKG)

""BRANCH" 1T01/4

CKG) '

CONES TWIGS FOLIAGE

(KG) (KG) (KG)

TwlG*FOL

(KG)

63 3 22) 2T, 2 25,8 20,0 0,401 8.6 15,4 11,8 48,9 23.7 UUUU 0,0 0,0 42,9

- X0P*-1" "TWIGS AND-FOLIAGE MID—-2 ARE NOT SEPARATED L0W--3 FOR CEDAR

5,4 22,3 15.1

SECTION NOt 1 2 3 4 5 6 7 8 9 10

NOOO 114,0 95,1 TT, 4 66,9 59,8 55,9 37,7 16,4 9,3 2,6 SUM WOOD" 539,0 KG

BARK 5,6 « . l •>. 5 6,3 4,8 1.5 4,5 3,9 1.0 0,8 SUM BARKS 42,4 KG TREE BIOMASS"

o

o

o

o o o

o o

TREE SPECIE AGE LENGTH LENGTH LENGTH DBH CROWN CROWN BUTT BRANCH BRANCH CONES NO TO 1 TO 6 DIAM LENG LENG OVER I 1T01/4

(H) (M J CM ) P fT (Ml"

76 195 14,8

- (H)—

13.3 5,3 0,180 6,1

- (rO—

9.1

TMT~

5.7

(KG)

1.7

(KG)

11.1

(KG)

UUUU

TWIGS

"(KG)

0,0

FOLIAGE

"(KG)

0,0

TWIG»FOL

(KG)

17,5

C

TOP--1 TWIGS AND FOLIAGE MID--2 ARE NOT SEPARATED

-C0W--3-"-F0R~"CEDAR

2.4 5,2

"9,-9-

SECTION NOl 1

WOOD 16,9

2

11,9

3

9,9

4

6.2

5

7.3

6

5.7

7

3,9

B

2.2

9

0.9

10

0,5 SUM WOODs 67,6 KG

BARK 0,6 1.1 0.5 0,8 0,6 0,6 0,6 0,6 0.3 0,0 SUM BARKs 5.7 KG TREE BIOMASSs f'P^/S KG

a************************************************** ************************** *******

"TREE—SPECIE AGE—LENGTH LENGTH—LENGTH—OBH—tROITN—CROTN BUTT—BR1TOCH—BR7SNCH—CONES TWIGS FOLIAGE TWTG+FOtT NO TO 1 TO 6 DIAM LENG LENG OVER 1 1T01/4


102 185 25,5 24,3 19,7 0,602 9,1 15,0 10,5 126,2 52,6 UUUU 0,0 0,0 109,6

T O P " - ! TWTGS-"AND-POtTAGE-

MID—-2 ARE NOT SEPARATED LOW--3 FOR CEDAR

-16-rr-49,2 43,8

SECTION NO I I 8 10

WOOD 245,7 167,1 140,5 111,5 99,5 75,5 52,5 31,2 11,5 2,7 SUM WOOD"

BARK 12.7 I3T3 12,5 13,2 9y9 Ty4 4.8 2.0 2,5 0.7 SUM BARKs

937,8 KG

79,0 KB TREE BIOMASSs / 30

**********************************************************************************************************************************

A P P E N D I X 6

NUTRIENT CONCENTRATION AND N U T R I E N T D I S T R I B U T I O N

W I T H I N 20 SAMPLED T R E E S ON THE SQUAMISH SAMPLE PLOT

S P E C I E S 1 - MOUNTAIN HEMLOCK

S P E C I E S 2 - P A C I F I C S I L V E R F I R

TREE NO.

22

SPECIES NO. 2

LENGTH M l

37.3

OBH (Ml

0.780

CROWN DIA(M) 7.5

CROWN BUTT L.(M) L.(M)

25.9 11.*

• ••PERCENT NUTRIENT CONTENT BY SAMPLE SPEC IF ICAT I 0 N * » * * * ********************************************************<•********# * * * » * < • * • » W000**SECTION— * * BARK**S ECT I ON *-

I 3 5 7 J) 10 1 3 5 7 9 10

NITROGEN -j> HO S PHORU S_-_ "C'ATCIUM -MAGNESIUM -POTASSIUM -

0.053 _0, OA 7_ 0.051 0.009 0. 03*

0.0*5 0.005_ 0. 07* 0.011 0.058

0.0*7 0.003

0. 056 0.005

0.063 0.008

0. 069 0. 010

NITROGEN -PHOSPHORUS-

0.277 Oo 037

0. 063 0.012 0.05 9

0. 082 0. 017 0. 068

0.071 0.016 0.10*

0. 067 0. 016 0. 078

CALCIUM MAGNESIUM POTASSIUM

0.335 0. 019 Oc 090

0. 207 0.0 3J_ 0.126 0.028 0. 102

0.208 0.032_

' O.*6"0 0.027 0.093

o.i e*

"b."350 0.031 0.117

0.203 0, 03 *_

' 0,'*26 0.030 0.096

0.297 0.0*7

"6". 6 88" 0.040 0.172

BRANCH BRANCH -TWIGS- -FOL I AGE- CONES >1 1-1/* (TOPI ( MIOI

NITROGEN - 0. 169 0.227 0.000 0.000

(LOW)

0.000

(TOP)

0. OOP

(MID)

0.000

(LOW)

0.000

*—-TWJGS + FOL I AGE

( TOPI " " ( MI D)~ flOW) 2.133 0.789 0 . 6 8 8 0.76*

PHOSPHORUS-CALCIUM MAGNES IUM -

0. 02* 0. 311 0.02*

0. 033 0. 356 0.031

0.000 0. 00 0 0.000

0.000 0.000 0. 000

POTASSIUM 0. 107 0.11* 0.000 0.000

0.000 0.000 0.000 0.000

0.000 0. 000 0. 00 0

0. 000 0. 000 0. 000

0.000 0.000 0„ 000

0. 000 0.000 0.000

0. 368 0. 067 J?„10JL 1.260

0.11* 0.*13 0.060 0.311

0. 102 0.)07 0.612 0.563 0.070 0.073_

0.306 0. 261

***NJTR!ENT AMOUNTS IN DECAGRAMS BY SAMPLE SPEC IFI CAT I ON*****************••*•***••«****•**•*»****»****•***•»***»*•**•***»***• »•***

-WO00«*SECTICN- -BARK**SECT!ON-1C2 3 £ * 5 £6 7£8 10 TOTAL l t2 3£*

NITROGEN - * * .97 29.12 21.69 17.26 1.99 0.*2 11,5. *5 NITROGEN - 29.6* 17.26 PHOSPHORUS- 9.71 3.05 1.66 1.81 0.25 CALCIUM - 51.96 *3.15 30.3* 22.13 2.25 MAGNESIUM - 8. 17 7. 1* 6. 1 3 *. 83 0.50

0.06 16.55 0.*1 150.25 0.09 26.85

PHOSPHORUS-CALC1UM MAGNESIUM -

*.17 2.62 *0i*6 33.51 2. 83 2.26

5t6

16.61 '2.7* 35.20 2.*6

7C8

18. 29 " '3V'1'3 36.71

2.91

9 1 0

3, 69 0. 56 "0", 61 " " OYOS 7,76 1.31 0. 55 0.08

TOTAL

8 6 . 06_ "" 13."3 6 1 5 * . 0 6

11 . 09

POTASSIUM - 59.03 36.89 26.5* 2*.80 3.27

_BRANCH_ BRANCH " >i" ~ "i-i>* (TOP)

---TWIGS (MID) "(LOW)'

0.*8 151.00

* * TOTAL (TOPI

POTASSIUM

--FOLIAGE-(MlUI ' (lOW)

1 1. 77 8.11 8.90 10.06 1,7* 0.33

* CONES * TWIGS»FOLIAGE — "TOTAL " "(TOP) (MIDI ~ (LOW)

TOTAL

NITROGEN - 69. 5* 19. 57 0. 00 0.00 0. 00 0.00 0. 00 0. 00 0. 00 0.00 0. 00 PHOSPHORUS- 9. 77 2. 89 0. 00 0.00 . 0. 00 0.00 0. 00 0. 00 0. 00 0.00 0.00 CALCIUM 128. 1* 3 0. 7* 0. 00 0.00 0. 00 0.00 0. 00 0. 00 0. 00 0. 00 0. 00 MAGNESIUM - 9. 96 2. 66 0. 00 0. 00 0. 00 0.00 0. 00 0. 00 0. 00 0. 00 0.00 POTASSIUM - * 3 . 96 9. 85" "0. bo ~ d dd " " b." 00 0.00 " o. "00" d. 00 0. 00 " "r b.oo" " "0.00

**.65 39.95 63. 39 1*7.99

*0. 92

TP. EE TOTAL"

428,61 6.*3

23.37 3.38

17.61

5.95" 35„5*

* t 05 15. 16

8. H<5~ 2TT27" *6,77 105.68 6.06 1 3 . 4 9

25.36 " 58.13

6i.7T~ 569.7B 6 4 , OS

303. 87"

r

TREE SPECIES LENGTH OBH CROWN CROWN BUTT NO. NO. (Hi (Hi DIAIMI L.(M) L.(M)

41 2 21.9 0.356 6.2 19.2 2 .7

NITROGEN - 0. 051 0. 350 0.05 3 0.057 0.067 0. 090 NITROGEN - 0.276 0.273 0. 307 0. 325 Oc 431 0. 587 PHCSPHOBUS- 0.005 o.ooa 0. 01 0 0. 003 0.021 0.023 PHOSPHORUS- 0. 040 0.056 0.058 0.069 Oo 095 0.134 CAICIUM 0.06 9 0.056 0.051 0. 048 0.052 0. 037 CALCIUM 0.4 54 0.519 0.472 " 0.3"3 6 0"o"318" 0.327 MAGNESIUM - 0.015 0. 010 0.01 0 0.011 0.011 0.013 MAGNESIUM - 0. 018 0.042 0. 043 0. 042 0. 044 . 0.060 POTASSIUM - 0.06 7 0.060 0. 052 0. 060 0. 061 0. 078 POTASSIUM - 0.123 0.167 0.198 0.227 0. 312 0.395

• ••PERCENT NUTRIENT CONTENT BY SAMPLE SPECIF 1 * . W00D^*SECTION-

10 -BARK**SECT ION-

5 7 10

BRANCH BRANCH • TWIGS * _ _* — " " f PL I AGE — — — r _ * CONES * —TWI GS*FOL I AGE — - * >1 1-1/4 (TOP) (MID) (LOW) (TOP) (MID) (LOW) (TOP) (MID) (LOW)

NITROGEN - 0.143 0.214 0.000 0.000 0.000 0.000 0.000 0.000 0.990 0.836 0.735 0. 712 PHOSPHORUS- 0.021. 0.031 0.000 0.000 0.000 0.000 0.000 0.000 0.181 0.098 0.087 0.078 CALCIUM - 0.254 0.239 0.000 0.000 0.000 0. 000 0. 000 0.000 0. 025 0.301 0.346 0.380 MAGNESIUM - 0. 02 5 0. 030 0. 000 0. OOP 0.000 0.000 0.000 0.000 OÔbl 0.073 0. 069 0. 053 POTASSIUM - 0.073 0.114 0.000~~ 0.000 0.000 0. 000 0, 000 0.000 0.486 0.324 0.297 6.267

•••NUTRIENT AMOUNTS IN DECAGRAMS 8Y SAMPLE SPECIF]

-WOOO^SECTION- -BARK**SECTION-TOTAL

_NI TROGEN -PHOSPHORUS-CALCIUM HAGNESIUH -

1C2

6.05

3C4

4. 32

5 66

2.96

7£8

1.35 0.25

10

0.10

TOTAL

15. 55 NITROGEN

1C2

4.39

3C4

4.53

5G6

2.53

7£8 10

0.73 0.32 7.54 5.05 1.51 0.93

0 .48 2.63 0.57

0.32 0.08 1.09 0.19 0.25 0.04

0.03 0.04 0.01

2.50 16. 60

3. 32

PHOSPHORUS-CALCIUM MAGNESIUM -

0.76 0.88 7.78 7.73 0.48 0.66

0.51 3.23 0.34

2.2 3 "0.4 8"

0.22 '0."0'5~

1.93 0. 16 0. 25 0, 02 1. 59 0. 16

0.12 14.01 "0703 2771 ' 0.07 20. 89 0.01 1.77

POTASSIUM - 7.63 5.25 3.00 1.31 0.22 0.09 17.55 POTASSIUM 2. 32 2.85 1.70 0. 08

BRANCH BRANCH •- -TWIGS->1 1-1/4 (TOP) (MID) (LOW) TOTAL (TOP)

^ F 0 LI AG E - - * CON ES_ (Mib) (LOW] TOTAL

TWIGS*FOLI AGE--( TOP) "(MID) "(LOW)"

NITROGEN - 4.93 6.71 0.00 0.00 0. 00 0.00 0.00 . 0.00 0. 00 0.00 0.00 8.03 21.53 13.39 2.56 n w

10.14 7.15 2.01 1.00 "8.70 ~ 5 . 0 3

"TOTAL"

42.95

8. 69

TP EE "TOTAL"

e4.14 PHOSPHORUS-CALCIUM fAGNES HIM -

0.70 0.98 8. 42 7. 49 0.82 0.94

0. 00 0. 00 0. 00

0.00 0.00 0.00 0.00 0.00 0.00

POTASSIUM - 2.59 3.59 0.00 0.00 0. 00

0.00 0.00 0.00 0.00 0.00 0.00

"0.00 oVoo

0. 00 0. 00 0.00 0.00 0.00 0^0 0_ 0. 00 0. 00

^JTOT5orrnr 0.00 0.00 0.00 0. 00_

"d.'O'O 0.00

0.94 2.89 0.70

"3.11"

4.96 20. 18

3.71 16.84"

I I.s; 73. 59 10.57

-49726""

TREE SPECIES LENGTH 08H CROWN NO. NO. (HI (Ml OIAIMI 45 1 35.2 0.754 0.0

CROWN BUTT L.(M) L.(H)

29.2 6.0

•••PERCENT NUTRIENT CONTENT BY SAMPLE SPEC IF ICATION************************************************************** -WOOD**SECTION-- -BARK**SECTION-

1 3 5 7 9 10 1 3 5 7 <J 10

NITROGEN - 0. 057 0. 052 0.053 0. 051 0. 059 0. 067 NITROGEN - 0.259 0.171 0.208 0. 198 0. 247 0. 280 PHOSPHORUS- 0. 017 0. 007 0.009 0.012 0. 020 0.021 PHOSPHORUS- 0. 061 0. 054 0. 052 0. 049 0. 055 0.062 CALCIUM 0. 053 0. 057 0. 070 0. 078 0. 064 0. 081 CALCIUM 0.693 0.615 0.854 0. 374 0. 435 0. 3 59 MAGNESIUM - 3. 012 0. 012 0.016 0. 014 0. 015 0. 016 MAGNESIUM - 0. 016 0.016 0.017 0. 017 0. 020 C.024 . POTASSIUM - 0, 094 0. 057 0.050 0. 054 0. 080 0. 062 POTASSIUM - 0. 084 0.094 0.094 0. 101 0. 109 0.161

BRANCH BRANCH -TWIGS-- -FOL IAGE- CONES * TWIGS+ FOLIAGE---* >1 1- 1/4 (TOP) (M10) (LOW 1 (TOP) (MID) (LOW) ( TOP) (MID) ( LOW)

MTROGEN - 0. 000 0. 000 0.000 0. 000 0. 000 0. 000 0. 000 0.000 0.000 0.000 0.000 0. 000 PHOSPHORUS- 0. 000 0. 000 0.000 0.000 0. 000 0. 00 0 0. 000 0.000 0. 000 0. 000 0. 000 0. 000 CALCIUM - 0. 000 0. 000 0. 000 0.000 0. 000 0. 00 0 0. 000 0.000 0.000 0.000 0.000 0. 000 MAGNESIUM - 0. 000 0. 000 0.000 0. 000 0. 000 0. 000 0.000 0.000 0.000 0.000 0.000 0. 000 POTASSIUM - 0. 00 J 0. 000 0.000 0.000 0. 000 0.000 0. 000 0. 000 0.000 0. 000 0. 000 0. 000

•••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPECIF1

» . WOOD*»SECTION -BARK**SECTION-1C2 3C4 5C6 7E8 10 TOTAL

NITROGEN PHOSPHORUS-CALCIUM MAGNESIUM -

54.03 34. 20 20.2 5 11 17 11.86 5.33 4.16 3.30 54.42 41.30 23.83 14.43 11.35 9. 11 5. 39 3.02

2.44 0.58 122.72 NITROGEN

1&2

40. 50 0.83 2.65 0.63

0.18 0.70 0.13

25.66 142.34 30.62


3 £4

28^16 ,8 6" 10.84 7.

123.16 109.06 3.00 2.43

5 £6

J 4 . 7 6 "3.65 4 4 . 5 6

1.23

7C8

10.48 2";v4

19.04 0.87

9 10

1.92 0.48

"0.43 " o;n 3.39 0.61 0.16 0.04

TOTAL

96. 31 " "2'5. 32"" 299.83

7. 72 POTASSIUM - 74.56 34.86 20.38 13.65 3.33 0.54 147. 32 POTASSIUM- 16.82 13.98 7.09 4 .96 "0785 0.27

_BRANCH_8RANCH_*-- - -

"~>l" ""1-1/4 (TOP I

---TWIGS (MID)" (LOW)

• *. TOTAL ( TOP)

— FOL I AGE-(MIDI (LOW) TOTAL

CONES TWIGS«-FOLIAGE--(TOP)" (MID) " (LOW) TOTAL

43. 98

TREE TOTAL"

NITROGEN - 0. 00 0.00 0. 00 0. 00 0. 00 0. 00 0. 00 0. 00 0. 00 0.00 0. 00 0.00 0. 00 0. 00 0. 00 219.03 PHOSPHORUS- 0. OJ 0.00 0. 00 0. 00 0. 00 0. 00 0. 00 0. 00 0. 00 0.00 0. 00 0.00 0. 00 0. 00 0.00 50.98 CALCIUM 0. 00 0.00 0. 00 0. 00 0. 00 0. 00 0. 00 0. 00 0. 00 0. 00 0. 00 0.00 Oo 00 0. 00 0.00 442.16 MAGNESIUM - 0. 00 0. 00 0. 00 0. 00 0. 00 0. 00 0. 00 0. 00 0. 00 0.00 0. 00 0.00 0. 00 0. 00 0.00 38.35 POTASSIUM - 0. do o.oo" 6. 00 0. 00" """"0. ob" 0. 00 "0. 00"" 0. 00 ' 0. do"""" ~ "b". oo'" "b. 00" " 0.00 0. 00 0. 00 " 0.00 ' " 191.29

TREE NO.

55

SPECIES NO.

1

LENGTH ( H I

34. 4

OBH (Ml

0.631

CROWN OIA( Ml

0.0


28. 2 6.2

•••PERCENT NUTRIENT CONTENT BY SAMPLE S P E C I F I C A T I 0 N » * * * * ^ * * * * ^ * * * » * * * * * * * * * * * * * * * * * * * * • * • * * * • • • * • • • * • * • « • • • • * • • • * • • • • * • * • * • * • « » * » * • WCOD+*SECTION * * BARK**SEC Tl ON *

1 . 3 5 7 9 10 1 3 5 7 9 10

NITROGEN -PHOSPHORUS-CALCIUM MAGNESIUM -POTASSIUM -

0. 055 .0 -.021_ 6. 057 0. 016 0.073

0. 051 __.020_ 0.1)76 0. 022 0.083

0. 04 6 0. 01 2__ 6.079 0. 022 0.048

0. 049 _0. 015_ 0. 079 0. 021 0. 060

0.054 _0.014_ 0.0 51 0.013 0.066

0. 06 0 _0._013_ 0. 05 0 0. O i l 0. 057


0.285 _0i08_3_

0. 2 26 0.017 0.099

0. 162 0.063 0.286 0.016 0.073

0. 177 0.070_ 0. 386 0.018 0.106

0. 188 0.064

"0. 1 92" 0.020 0.101

0. 236 0. 308 0. 075 0.081 0.292 "0.313" 0.026 0. 037 0.120 0. 166

NITROGEN

BRANCH >i

0.000

JBRAil_CH_ 1-1/4

0. 000

( TO P I -TWI_ S-( MIDI ( L O W ) (TOP I

-FOLI AGE-(MI D I ( L O W I

0.000 0.000 0.000 0.000 0.000 0.000

CONES

0.000

* TWIGS* FOL I AGE » (TOPI (MID) " (LOW)

0.000 0.000 0.000 PHOSPHORUS-CALCIUM MAGNESIUM -

0. 000 0.000 0. 000

0.000 0.000 0.000

POTASSIUM - 0.000 0.000

0.000 0.000 0. 000 o."doo

0.000 0.000 0.000

0.000 0.000 0.000

0. 000 0. 000 0. 000

0. 000 0. 000 0.000

0. 000 0.000 0.000

0.000 0.000 0. 000 0. 000 0.000

0.000 0.000 0.000_

"0". 000

0.000 0.000 0.000 0.000 0.000

0.000 0.000 0.000 0.000 0.000 0^000

o". bob"

• ••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPEC I F I C A T I O N * * * * * * * * * * ^ * * * * * * * * * ^ * * * * * * * * * • * • • * • * • * * * « * * * * • * • * * » * * * « • • • « * * * * • * * * * » * » * •

-WOOO^«SECTION- -BARK**SECTION-1C2 3£4 5£6 7£8 9 10 TOTAL U 2 3C4 5£6 7C8 9 10 TOTAL

NITROGEN - 41.02 24. 23 15.95 7.70 1.21 0.31 90.42 NITROGEN - 40. 74 23.22 13.13 8.34 1.11 0. 34 86. 88 PHOSPHORUS- 15.74 7.97 4.58 2.17 0.32 0.07 30. 85 PHOSPHORUS- 13.32 9.11 4.84 — 2 7 7 3 "07'3"S~ "0.09" 30.44 CALCIUM 51.21 38. 38 26.43 9.65 1.13 0.26 127.06 CALCIUM 46.60 46.01 20.77 9.51 1.37 0. 34 124.60 MAGNESIUM - 14.54 10.92 7.11 2.52 0.30 0.06 35.44 MAGNESIUM - 3. 00 2.30 1.34 0.89 0.12 0.04 7.69 POTASSIUM - 62.16 32.59 18.14 9.3 5 1.47 0.30 124.00 POTASSIUM - 15.67 12.28 7.43 4.33 0. 56 0. 18 40.45

BRANCH BRANCH •- -TWIGS :* »->1 1-1/4 (TOPI ( M I D I (LOWI TOTAL

NITROGEN - 0. 00 0. 00 0.00 0. 00 0. 00 0. 00 0. 00 PHOSPHORUS- 0. 00 0. 00 0.00 0. 00 0. 00 0. 00 0. 00 CALCIUM - 0. 00 0. 00 0. 00 0. 00 0. 00 0. 00 0. 00 MAGNESIUM - 0. 00 0. 00 0.00 0. 00 0. 00 0. 00 0. 00 POTASSIUM - 0. 00 0. 00 0. 00 "o"; 00 0. 00 0. 00 0. bo

FQ L IAGE----T T C J P V ( M I D I " (LOW!

0. 00

TOTAL

0.00

CONES * TW I GS + FOL IAGE--< T O P ) ( M I D I " " ( L O W ' l '

0.00 0.00 0.00 0.00

"TOTAL"

0.00

TREE "TOTAL"

177.30 0. 00 0.00 0. 00 0.00

0.00 0.00 0. 00 0.00

0.00 0.00 0.00 b.oo"

0. 00 0.00 0. 00 0.00

0.00 0.00

"0.00"

0.013 5701) 0.00 0.00 0.00 _ 0.00 "0. 00 "0. 00

"DT30" 0. 00 0.00

"0.00

61.29 251.66

43. 13 164.45"'

TREE NO.

64

SPECIES NO. 1

LENGTH 1 M I

21.3

OBH (M)

0.320

CROWN DIA(Ml

0.0

CROWN L.(M)

12.4

BUTT L. (Ml

9.0

»*»OERCENT NUTRIENT CONTENT BY SAMPLE SPECIF ICATION******************************************************** * WOOO**SECTION * * BARK**S ECT ION «

1 3_ 5 7 9 10 1 3 5 7 9 10

NITROGEN -PHOSPHCRUS-CALCIUM MAGNESIUM POTASS!UM

0.062 0.017

0.053 0.027

0.061 0.095 0.016 0.025 0.066 0.081

0.051 0.01 7 0. 06 9 0.018 0.072

0. 055 0±0_ZQ_ 6. 053 0.017 0. 067

0.066 0.014

0. 075 0. 012 0.047 0. 042

0.013 0.013 0.055 0.065


0.251 0. 059

0.178 0.075

0.704 0.345 0.018 0.016 0.072 0.075

0. 198 _0._04J _ 0.308 0.018 0. 084

0.208 0^08 0__ 6.216 0.018 0.093

0.248 0.095

0.339 0.103

0. 194 0.226 0.030 0.045 0. 106 0.159

BRANCH BRANCH >1 1-1/4

-TWIGS- -FOLIAGE-(TOP) (MID) ( LOW) (TOP) (MID) (LOWl

CONES * TWIGS + FOlI AGE * ( TOP) (M ID ) ( LOW )

NITROGEN 0.030 0.000 0.000 0.000 0.000 0. 000 0.000 0.000 0.000 0.000 0.000 0.000 PHOSPHORUS-CALCl 'JM MAGNESIUM -POTASSIUM -

0. 000 0.003 0.03J

0.003 0. 000 0.000

0.003 3.000

0.03 0 0.300 0.00 0

0. 000 0.000 0. 000

0.000 0.000 0.000

0. 000 0. 000 0. 000

0. 000 0. 000 0. 000

0.000 0.000 0. 000

0.000 0.000 0.000 0.000 0.000 0.000

0.000 0.000 0. 000 0. 000

0. 000 0.000 0.000 0.000

0.000 0.000 0. 000 0.000 0.000 0.000 0.000 0.000

•••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPEC IF ICAT ION**** * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * **

-WOOD**SECTI0N- -BARK**SECTION-

NITROGEN

162

6.92

3 64

4. 09 PHOSPHORUS-CALCIUM MAGNESIUM -

2.67 3.76 2.46

1.73 6.04 1.69

566

2.76 0.95 3.13 0. 92

7£8

1.75 0.39

10

0.13

TOTAL

16.03 NITROGEN -

162

4.92

364

3.45 0.49 1.45 0.43

0.08 0.28 0.07

0.02 0.08 0.02

5. 95 19.77

5.60


1.53 12. 01

0.40

1.06 5.97 0.32

5 £6

2. 25 "6.67 2.91 0.20

7£8

U 09 0.42" 0.98 0.11

10 TOTAL

0. 15_ "0. 06 0. 12 0. 02

11. 96 ,77"

0. 10 "OTOT 3. 0.07 22.05 0.01 1.06

POTASSIUM 8.82 6.02 3.62 1.76 0.32 0.12 20.65 POTASSIUM 1.68 1.46 0.98 0.48 0.06 0.05

3RANCH BRANCH _*----------TWIGS :

>1 1-1/4 1 TOP) (MID I (LOW) * * - - - -FOL I AGEj^----

f b TA L " ( TUP) (MIDI ( L 0 W) »

"TOTAL

CONES * TWIGS+FOLIAGE--" " " " " ( T O P ) , .' (MID) " (LOW) T3TAL

4.71

TREE TOTAL"

NITROGEN 3. 30 0.00 0.00 0.00 ' 0. 00 0.00 0.00 0.00 0.00 0. 00 0.00 0.00 0. 00 0. 30 0. 00 28.00 PHOSPHORUS" 0.33 6.00 6.00 0.00 3.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 9.73 CALCIUM - 0.03 0.30 0.30 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.60 0.00 0.00 41.63

_MAGNF SI UM _- 3. 33 0.00 0. 00 0.00 0. 00 _ 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 6.65 POTASSIUM -~ 3.66 0.00" 0.00 "6 . 00 Of. 00' "6 .00 "6.00 0 . 0 0 0 . 0 0 6.00 ' 0 . 0 0 0 . 0 0 " 0.00 0.00 0.00 '25.35

r

TREE NO. 63

SPECIES NO. 1

LENGTH ( M)

33.0

DBH ( .11

0. 711

CROWN DlAIM) 7.9

CROWN BUTT L.IMI L.(M) 24.5 13.5

• •••PERCENT NUTRIENT CONTENT BY SAMPLE SPECIFICATI ON********************************* ********•***•*•***•••*•••••»••***••»*•**•**** * • WOOD**SECT ION * * BARK**SECTI ON *

1 3. 5 7 9 10 1 3 5 7 9_ 10

NITROGEN -JLHO.SiHOjyj.Sj; CALCIUM MAGNESIUM POTASSIUM

3. 053 _0._Q32 0.073 0. 026 0.070

0.054 _0,_P13_ 6.067 0. 320 0.063

0.04 8 0. 012 0.08 8 0.02 9 0. 056

0.055 _P„P16_ 0.078 0.024 0. 071

0.068 J>.026_ 6.090 0.029 0.076

0.075 __0^029_ 0. 076 0. 025 0. 08 5


0. 190 0.070 0.307 0.019 0. 075

0.137 _0.061 6.229"" 0.017 0.084

0. 148 0.053_

" 6. i"96 0.016 0.087

0.155 3.047 '~6".2 45"~ 0.01 7 0.092

0.2 26 0.057

0. 185 0, 047 "0.177 0.282 0.019 0.022 0. 100 0.110

_B?ANCH_ >1

BRANCH 1-1/4

• TWIGS-ITOPT (MID) ( LOW) (TOP)

-FOLIAGE-(MID) (LOW)

CONES * TWIGS*FOLI AGE * "(TOP) (MID) (LOW I

NITROGEN 0. 146 0. 202 0.475 0.449 0.243 0. 840 1.404 0.805 0.000 TQ> 57 0.927 0.1>^* PHDSPHORUS-CALCIUM MAGNESIUM -POTASSIUM

0. 02 7 0.110 0. 022

0. 046 0.125 0.032

0.072 0.178 0.045

0. 067 0. 170 0.050

0.0 34 0.125 0.045

0. 108 0. 313 0. 100

0. 118 0.282 0.094

0. 103 0.279 0.089

0.000 0. 000 0.000

0.074 0.091 0.155 0.156 0.169 0.235 0.250 0.273 0.000

• ••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE s PEC IF ICAT I ON**************************** *************************************"*** ***** *

• WOOD** SECT ION * * BARK** SECT I ON 1C2 3C4 5 £6 7£8 10 TOTAL 1(12 3£4 5£6 7&B TO" TOTAL

NITROGEN - 41.52 29.24 21.16 13.15 2.70 0.70 108.47 NlTROGEN PHOSPHORUS-CALCIUM MAGNESIUM -

16. 86 53.59 16.92

7. 30 44.66 13. 13

5.34 34.45 11.00

4.54 17.94 5.66

1.03 3.54 1.13

0.27 0.71 0.23

35. 84 154.90 48. 88


23.02 25.44 14.69 9.26

37.79 2.53

5.11 T7T3

1.20 76.25

o;"30 27rer 1.49 112.75 0.12 8.48 TJ7F8 42. 57 POTASSIUM 49.06 34.09 26.23 1 5.67 2.99 0.79 128. 83 POTASSIUM - 11.27 15.36 BT71

NITROGEN

BRANCH 9RANCH *->1 1-1/4

31.79 6.32

(TOPI

6. 79

•—TWIGS (MIDI (LOW"!

5.30 9. 24

# *. "TO TAT

-FOLIAGE- CONES (TOP) (MID) (COW) TOTAL

21.34 13.94 21.77 41.78 1.80 5.20 1. 66 "3^9"0"

77.49 8. 97

24.05 7.72

21.92

TWIGS* FOLIAGE--"'( TOP )""""< MI Or" (LOWI

0.00 0.00 0.00 0.00 o. Ob"

20.73 2.83 7.74 2.31 6.12

27.07 2.61 6.37 2.05

'"""5.71"

"TOTAL""

98.82 51.03 6.63

19.23 33.34 6.33 ' 10.69

20.61 " 32.44

TREE "TOTAL"

321.64 85.03

328.77 73.79

"2 22." 69"

PHOSPHORUS- 5.84 CALCIUM - 23.86 MAGNFSIUM - 4.73 POTASSIUM - 16. 02

1. 43 3. 92 W 01 2.8 3"

1. 03 2. 54 0.65 2.22

0.79 2. 01 0^59 1". 85 "

1.28 4. 75 1. 73 6.46

3.10 9.29 2.97

16152'

1.82 4.37 1.45 3.87

5.35 14.48 4. 60

14.15

http://JLHO.SiHOjyj.Sj

TREE NO. 74

SPECIES NO. 1

LENGTH (Ml

34.2

CBH (Ml

0.762

CROWN DIA(M) 9.4

CROWN BUTT L.IM) L.(M)

25.2 9.0

• ••PERCENT NUTRIENT CONTENT BY SAMPLE SPEC IF I CAT I ON*** * • * * * * * * • * * • * • • • • * • • • • * * • • * • • • » * • • • • • « » • • • * • • « • • * » " » *M~j+^*&Vi"rr*«î~M#**** • WOOD^SECTION * * BARK**S ECT I ON »

1 3 5 7 9 10 1 3 5 7 9 10

NITROGEN -PHOSPHnRUS-CALCIUM MAGNESIUM -POTASSIUM -

0.056 J? • 0.064 0.015 0. 084

0.045 J?.Q13_ 0. 072 0.018 0.052

0.051 0.014

0. 044 0.021

0.056 0.014

0. 080 0.014


0. 076 0.018 0.052

0. 081 0.020 0. 060

0.063 0.012 0.054

0. 052 0. 010 0.062

CALCIUM MAGNESIUM -POTASSIUM -

0.218 _0._083_ 0.308" 0.017 0. 092

0.136 0.060 0.344 0.013 0.087

0. 144 0. 054_

" 0.243 0.014 0.115

0.147 0.046_

"0.226 0.014 0. 085

0.229 0.307 0. 06b 3.0 79 0.261 " 0~. 2 73" 0.019 0.028 0. 143 0.181

NITROGEN

BRANCH BRANCH >1

0. 114

1-1/4

0.156

(TOP I

0.477

-TWIGS- -FOL I AGE * (MID)

0. 390

(LOWI

0. 3 72

(TOP)

0. 683

(MIDI

0. 866

(LOWI

0.819

CONES

0.000

? r ~ I _ I G J . i F O l - _ *_E - -"(TOPI (MIDI" ( LOWI

0.62S O.^&^y PHOSPHORUS-CALCIUM MAGNESIUM -POTASSIUM -

0. 025 0. 162 0.013 0. 069

0.029 0.184 0.023 0.091

0.067 0.191 0.042 0.156

0. 059 0. 156 0. 038

0.061 0.163 0.043

0. 112 0. 394 0. 077

0. 121 0.335 0. 074

0. 114 0.335 0. 067

0.176 0.192 0.281 0. 332 0.311

0.000 0.37 0.000 0.293 0.000 0 0. 000

440 0.249

055

• ••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPECIF I CAT I ON******** * * * * * * * * * * * * * * * • » * * * # * * « • * * • « * * • • * • • * • • • • * • * * ••••••^•••* »•••"•"* •»"*"•"

-W00D**SECT10N- -BARK**SECTION-1£2 3£4 5(16

NITROGEN 40.90 26.50 17.44

7£8

8.90 PHOSPHORUS-CALCIUM MAGNESIUM -

14.60 54.87 12.97

7.53 40.66

9. 30

6.37 28.63

6. 96

3.12 12.82

2. 85

1.26 0.33 1.43 0.28

10

0.29 0.05 0.19 0.04

TOTAL

95.29 32. 00

138.61 32.89


1E2

15.01 68. 69

3.13

3 £4

37^31 26.42_ 1077 3 55.47 2.56

5 £6

15. 18 5.22

24.50 1.47

7£8

J2 .06 "3.59 15.64

1.07

1. 2 6 "0'. 36 1,44 0. 11

10 TOTAL

0. 37 92. 60 O.'l'O 35700" 0.33 166. 07 0.03 8.33

POTASSIUM 55.25 28.77 20.60 10.18 1.23 0.22 116.25 POTASSIUM 18. 79 19.00 10.40 7770 0"77~B~ 0771 56.49

BRANCH BRANCH •- -TWIGS->1 1-1/4 (TOPI (MIDI (LOWI

NITROGEN - 32. 99 8.41 15. 70 6. 97 7. 18 PHOSPHORUS- 7. 25 1.56 2.19 1. 06 1. 19 CALCIUM 46. 83 9.91 6.27 2. 80 3. 15 MAGNESIUM - 5. 13 1. 26 1. 37 0. 69 0. 82 POTASSIUM - 19.99 4.39" "5. 13 3. 16 "3. 70"

• *. TOTAL (TOP)

— F O L I A G E — - ; (MIDI" "(LOW)

CONES

29.85 21.10 20.53 29.39

TOTAL

71. 02

TWIGS+.FOLIAGE--"(TOPI" (M10 I (LOW I TOTAL"

0.00 36.80 27. 50 36. 57 100.88

TREE TOTAL"

330. 17 4.43

12.22 2.88

11.98"

3.45 12. 19 2.38

"8.67

2.87 7. 94 1.74

"7.86"

4.10 12.04 2.42

11.18

10.42 32.16

6. 54 '27.71"

0. 00 0. 00 0.00

"0. 00"

5.64 18.46 3. 75

13.80 "

3.92 10.74 2.43

11.01

5.29 147T5" 15.18 44.38 3.24 9.42

14.88 " 39.69

90.65 405.85

57.05 237. 32

TREE NO. 80

SPECIES NO. 1

LENGTH ( M) 31.2

DBH CM)

0. 564

CROWN DIA(M) 6.2

CROWN BUTT L.tM) L.tMl 25.8 5.5

1 3 5 7 9 10 1 3 5 7 9 1 0

NITROGEN - 0. 05 3 0.036 0.052 0.049 0.064 0.071 NITROGEN - 0.199 0.158 0. 168 0. 186 0. 264 0.319 PHOSPHORUS- 0. 020 0. 006 0.010 0. 023 0.0 24 0. 013 PHOSPHORUS- 0.073 0.055 0.047 0.051 0. 058 0.072 CAICIU" 0.056 0.058 0.053 0. 078 0.065 0. 043 CALCIUM 0.3 71 0.208 0. 147 0.235 "" 0; 155 0.185 MAGNESIUM - 0. 01 7 0. 015 0.014 0.024 0.019 0. 014 MAGNESIUM - 0.016 0.020 0. 017 0.021 0. 023 0.034 POTASSIUM - 0.091 0.051 0.052 0. 069 0.093 0. 063 POTASSIUM - 0.072 0.115 0.093 0.104 0. 129 0. 187

BRANCH BRANCH —TW1GS- * -FOL IAGE- * CONES * TWIG S+FOLI AGE * >1 1-1/4 (TOPI ( MID) ( LOW) ' (TOP ) ( M ID) " (LOW) (TOP ) (MID) 1 LOWI

NITROGEN - 0. 135 0. 162 0.471 0.353 0.386 0. 902 0. 854 0. 845 0. 000 T_xijS6 0.6 04 O.bJJf PHOSPHORUS- 0. 033 0. 037 0. 351 0. 061 0.066 0. 121 0.133 0.132 0.000 0.475--. --0T456 CALCIUM 0. 146 0.172 0.204 0. 194 0.183 0. 222 0. 208 0.265 0.000 0.213 0.224 MAGNESIUM - 3. 022 0. 027 0.040 0. 042 0.048 0.060 0. 057 0.072 0.000 0.05JJ---"0. 050^ -»J3_t060

POTASSIUM - 0. 37 3 0.083 0.164 0.139 0. 173 0. 307 0.357 0.278 0.000 j8<?35 0.243 o7z>6.

•••PERCENT NUTRIENT CONTENT BY SAMPLE SPECIF! -WOOD^SECT ION- « BAP.K**SECTION-

•••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPECIFICATION**************************************************************************

-WOOD**SECTION- -BARK**SECTI ON-T O

0.38 ""07"0"9~ 0. 22 0.04

TOTAL

50.59 ~T5784~

63. 64 5.31 2 8. 04

1C2 3C4

NITROGEN - 21.0313.62

5C6

9.05

7t8

4.65 0.68

10 TOTAL

0.15 49.22

1C2 3S4

NITROGEN - 21.11 12.45

5C6


6.39 2.51 2.90 27.06 17. 15 11.66 7.64 4.51 3.44

7C8

8.30 1.92 0.25 0.03 14.00 PHOSPHORUS- 7.57 3.92 2.03 2.0l~ 5.87 0.68 0.09 62.50 CALCIUM - 34. 19 13.52 7.37 7.21 1.78 0.20 0.03 17.60 MAGNESIUM - 2.16 1.43 0.78 0.80

1.06 "0723" 0. 62 0.09

POTASSIUM 31.45 15.97 10.88 6.67 0.98 0.13 66.09 POTASSIUM - 11.03 7.93 4.05 4.2 9 0. 51 0. 22

BRANCH BRANCH •- -TWIGS- -FOLIAGE- CONES >1 1-1/4 (TOP) (MID I (LOW) TOTAL (TOP I (MID)

NITROGEN - 14.47 0. 79 7.72 8.91 3. 90 20. 53 15.51 30. 23 PHOSPHORUS- 3. 51 0. 18 0.84 1.53 0.67 3.04 2.08 4.71 CALCIUM 15.62 0. 34 3. 35 4. 90 1. 85 10.10 3.83 7.37 MAGNESIUM - 2.37 0. 13 0.66 1.07 0. 49 2.22 1.02 2. 01 POTASSIUM - 7.47 0. 43 2.68 3.50 1. 75 7.93 5.29" 12.62

"(LOW) TOTAL TWIGS•FOLI AGE

(TOP) (MID) (LOW)

14.78 60.53 0.00 23.23 39.14 18.69 1 96.13 9. 09 0. 00

15.83 0.00 4.29 0.00

"22777 7j;-ffO"

2.91 7.18 1.69

"7.97

~6"7Z4" T T W 12.27 6.48 3.08 1.75

16. 12 " "6761

"T27TT" 25.93 6. 51 30.70"

45.67 168. 53 31.92

T32. 73'

2.31 4.64 1.26

"578 7""

TREE NO. 105

SPECIES NO. 1

LENGTH <M> 31.9

OBH (H ) 0.5 36

CROWN DIA(MI 4.7

CROWN BUTT L.(H) L.(MI 23.5 8.4

•••PERCENT NUTRIENT CONTENT *

I 3__

BY SAMPLE SPECIFI CAT I ON** ********* ****••» *»»**»«**«****«*•* *»****•»* «*###**** — WOOD**SECTION -* * 6ARK**SECT ION *

5 1 2 10_ 1 3 5 7 g 10 NITROGEN -PHOSPHORUS-

0.030 _0i.031 CALCIUM

MAGNESIUM -POTASSIUM -

0.052 0.015

0.052 0.057 0.015 0.015 0.065 0.054

0.055 0.02 0

0. 061 0.030

0.062 0.024

0. 075 0.019

0. 062 0.014 0.056

0. 074 0. 023 0.071


0.208 0. 083 0.057

0. 019 0.073

0. 04 8 0. 015 0. 064


0.280 0.01 8 0. 083

0.174 0. 072 0.275 0.018 0.090

0. 161 0. 071 0. 184 0.017 0. 095

0.179 0. 066 " 0". 2 36" 0.021 0.098

0.222 0.074 0.152 0 0.025 0.034 0. 128 0.169

0.275 0.081 1 59"


BRANCH BRANCH 1-1/4 >1

0. 127 0. 03 3 0. 177 0.02 2

POTASSIUM - 0.075

0. 194 0.043 0. 161

0.092 "

(TOP)

0.462

-TWIGS-(MID)

0.483

( LOW)

0. 396 0.063 0. 173 0.038 0.146

0.079 0. 187 0. 043

0.069 0.163 0.053

(TOP)

0. 102 0. 322 0. 068

-FOLIAGE-(MIDI

0.870 0.845

0. 148 0.176 0.273 0. 278

(LOW I

0.854

CONES _* TW_LGS*FOl I*.G_E

(TOP) (MID)"" " (LOW)

• ••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPECIFICATION**"*************************"***»• ******"****-*TS***^»T(.^^t?»^«^S1«T«W*^*

-WOOD** SECT I ON * 162 3E4 5 66 7C8 10 TOTAL

-BARK**SECTION-1£2 3C4

NITROGEN - _29.71 16.27 11.08 6.38 PHOSPHORUS-CALCIUM MAGNESIUM -

1 0.49 24.65 6. 86

1.34 5.39 18.09 4.41

4. 84 12.96 3. 55

0.42 65.20 NITROGEN 2.83 6.81 2.20

0.52 1.23 0.41

0.11 0.26 0.08

24.17 63. 99 17. 52

POTASSIUM- 26.72 16.71 12.17 7.54 1.59 . 0.35 .65. 07


21.42 _6.05 6.8 5 21.96 1 .69

8.68 31. 09 2.02

5C6

9.70 "3.91 12.01 1.08

768

5_.3 7_ "1.86 5.20 0.62

POTASSIUM

9 10

1.27 0.27 "0.42 0.08" 0.86 0.16 0,14 0.03

TOTAL

54. 08 '21."8 0" 71.28 5. 59

9.68 8.82 5.51 3.03 0.73 0. 17

_BRANCH_BRANCH_*:

" " 1 - 1 / 4 >1 r r — r — T W I G S ^ * *—-----FOLIAGE-— * CONES * TWIGS*FOLIAGE * (TOP) (MID) (LOW) TOTAL (TOP)"" (MIDI (LOW I TOTAL (TOPI (MID) " (LOW) "TQTAL"

NITROGEN 10. 72 7. 36 6. 43 FHOSPHORUS- 2. 83 1.64 0. 95 CALCIUM - 14.92 6. 12 2.40 MAGNESIUM - 1. .33 1.11 0.52 POTASS IUM - 6.33 3.49 2.03

6.09 .00

4.79 17.30 15.14 21.72 25.55 62.40 1 . 2.36 0.54 1.87

0. 83 2. 04 0.64 2. 13

2.73 6.79 1.70 6.03

1.78 5. 60 1.19 4.76

1.92 21.57 27. 81 30. 34 2. 95 8.64 1.81 7.14 "

3.30 11.61 2.29 7.69"

79. 71 8.03

25. 85 5.30 19.59

0.35 0.05 0.10 0.64

2.73 8.01 1.71 6.79

27.94

TREE TOTAL"

218.99 3.95 10.99 2.35 9. 01

4. 13 10.81 13. 65 32.65 2.93 6.99 9.62 25.62

61 . 6l 189.01 33. 13

"129.09

TREE NO. 110

SPECIES NO. 2

LENGTH (HI

41.2

DBH ( M)

0. 904

CROWN DIA(M) 8.4

CROWN BUTT L.(M) L.(M) 28.4 12.8

1 3 5 • 7 9 10 1 3 5 7 9 10 — i

NITROGEN - 0, 045 0. 045 0.036 0. 046 0.048 0. 059 NITROGEN - 0.212 0. 178 0. 177 0.175 0. 235 0. 251 £

PHOSPHORUS- 0. 00. 0. 004 0. 004 0.007 0.006 0. 009 PHOSPHORUS- 0.051 0. 042 0.035 0.045 0. 052 0. 046 1

CALCIUM 0. 054 0. 058 0.05 5 0.070 .0.06 5 0. 062 CALCIUM 0.298 0. 487 0. 507 ' 0.433 " 0. 426 "" " "0. 566 t'

MAGNESIUM - 0. 010 0. Oi l 0.009 0.014 0.013 0. 012 MAGNESIUM - 0. 022 0. 029 0. 030 0. 033 0. 041 0. 042 POTASSIUM - 0. 060 0. 053 0.051 0. 057 0.068 0. 068 POTASSIUM - 0.108 0. 116 0. 130 0.1 21 0. 143 0. 1 26

BRANCH BRANCH -TWIGS — « *- -FOLIAGE- _CQNES__ • ---TWIGS+FOLIAGE *

i j i

>1 1-1/4 (TOPI (MID) ( LOW) (TOP) (MID) (LOW 1 (TOP) (MID) (LOW) •

NITROGEN - 0. 157 0. 231 0.000 o.odo 0.000 0. 000 0. 000 0.000 1.041 0. 715 0. 814 0.794 | PHOSPHORUS- 0. 020 0. 036 0. 000 0. 000 0.000 0. 000 0. 000 0.000 0.226 0. 094 0. 104 0.110 't CALCIUM 0. 440 0. 331 0.000 0.000 0.000 0. 000 0. 000 0.000 0. 049 0. 345 0.410 0.457 MAGNESIUM - 0. 031 0. 02 7 0.000 0.000 0.000 0. 000 0.000 0.000 0. 070 0. 045 0. 053 0.067 I

I POTASSIUM - 0. 10J 0. 137 0.000 0.000 0.000 0. 000 0. 000 0.000 0.694 0. 369 0.311 0.372

• ••PERCENT NUTRIENT CONTENT BY SAMPLE SPEC IF ICAT I ON****************************************************************************** * -WOOD**SECTION- -BARK**SECTION-

•••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPECIFICATION*******************************************************

•wnOD*«SECTION- -BARK**SECTION-

U2 364 5£6 7C8 10 TOTAL 1£2 3£4 5C6

NITROGEN 55.00 37.37 21.69 13.78 3.83 1.52 133. 18 NITROGEN 30. 42 41.77 29.80 PHOSPHORUS-CALCIUM MAGNESIUM -

4.54 3.65 67.82 52. 22 13.10 9.63

3. 35 33.36 6.26

1.92 0.45 19.77 5.14 4.00 1.06

0.24 13.84 1.61 179.92 0.32 34.40


7.23 9.00 61.24 116.92 4.01 6.94

6.73 79.43 5.28

7£8

18.49 4. 38.71 3.30

9 10

5. 19 3.42_ 1". 14 "0.62 9,41 7.69 0.90 0.57

TOTAL

129.10 " 29.08 313.41 21 .00

POTASSIUM 63.21 48.03 23.70 18.35 5.43 1.76 170.49 POTASSIUM - 17.53 28.94 21.22 11.92 3. 16 1.71

BRANCH 8RANCH *- -TWIGS- -FOLI AGE- CONES *• >1 1-1/4 (TOPI (MID) (LOWI TOTAL (TOP I (MIDI "(LOW) TOTAL

JWIGS+FOLIAGE— (TOP I (MID)(LOW) TOTAL'

84.50

TREE "TOTAL"

NITROGEN - 52.66 16.32 0.00 0.00 0. 00 0.00 0. 00 0.00 0.00 0.00 1.25 54.59 63.66 31.90 150.15 482.66 0.00 0.2 7 0.00 0.06 0.00 0.08 "0.00 0.83"

4.41 T577T" 18.37 76.82 2.70 10.25 14.97 67.45

7TT2T" 741. 04 77.91

"3 66."53"


6. 77 147. 45 10. 26

2. 53 2 3. 38 1.91

POTASSIUM - 33. 57 9. 70

0. 00 0. 00 0. 00 0.00"

0.00 0.00 0.00 0.00 0. 00 0. 00 "0.00 0.00"

0.00 0.00 0.00 0.00 q.oo o.oo 0.00 0.00

0.00 0.00 0.00 0.00 0. 00 0.00 0.00 0.00

7.21 26.35 3.42

"28;i7"'

a. io 32.10 4.13 24. 31

W.*E9 1E*ST . O r » . . : » 0 ' l . M * V 00*0 Ei. * t T S0*£ 9»7*S ZZ*£ 8S*E _50* I 85*1 St'0 _ 6 i M 69**>Z 6T*^ 90*1 96 *T 8 f l 00*0 80*E ZA*0 L^'l 06*6 11 ° I "\l *6"" 6<7 '0"~" 6 Z *0"' £9 °0 15*021 TZ'ST. 8 i c E 03*9 £9**/ 00 *0 l^'OT LZ'Z 09** <?**£ 6£**> T V T 6t "Z 61*1 »<9 *S 59*5Z _.8'S 8£*T 86*Z OS'T 00*0 »*>"7 £6*0 S£*Z 9TT, £*>*! S*/*0 £9 '0 SE*0 99'0

*S '£ - KPISSVlOd £ T T. - '(.IHSaNOVh OZMl - kfllDlWD e i ' I -SOMOHdSOHd

18*911 *>8*3E 98*6 *0Z'AT 8A*II 00*0 9!'0£ £6*9 8S*EI 99*6

j w i o i _ i v i c i 3381 *

89*8 £6 °Z Z9*E £ T Z 8/ *E SE "9 N3'JfJblIN

I MOT )....•( 01 W)...(d01). jy.l01_._.(MDl)_...(OIW)_ (dOl) 1V101_ (MOD ( 01W ) J dOl) *t I 1-1 --39VI103+S9IM1 * S3N03 * 39VI103 * * S9 I Ml - — . ; _ » HONVfcB '"HONVoB

9*r'<7 U*S - wnissviod TO t-8*0 1Z*T 6 T *S £8*6 68 * ET 56*iT - Hfl l SSVlOd

86*6

i6*0 V9 *5T

_09*Z 9T*£I

- rtfllSSNOVW wnmvD

-SflTOHdSOHd - N390H1IN

80*91 60*0S

9E*££

60*0 61*0 60*0 8Z*0

8E*0 £1*1 8E*0

56*1 8i*S SS'T

i9*E T*7*TT LX'Z

LZ'<7 61 *£T 8Z*Z

ZA'S 1V8T. £?**

•;6*0 86*£ 65 *9 6«7*8 6 C E I

- Wfll S=M)VK

-snautidSOHd - N390»1IN

1?101 01 83A 93 5 *3£ Z3I "1V101 01 83A 93 5 <;3£ -NOU03S*#X'dV8- -N0li33S*»Q0OM-

Z31

_jt*aA»*JL»»-»»JL«JJL»«^ d I 03d5 3TdHV5 A8 SHVMOV330 NI S1NDCHV lN3 IK inN»»»

1 •")

o

OATO SAO'C A90*0 - wnissviod

65 I *0

- HOI S3N'9Vh

-SnsOHdSOHd - N320biIN

(MOT ) t a i w ) IdOl) -39»nO3+S0IMl » S3N03

(MOD IOIW) (dOl) (MOD (01W) ( d C U ) Wl-T T< -39VI103- —S9IM1- HDNV H3NS'o8

901*0 ZZO'O 91 £ *0_ SSO'C <7AZ*0

160*0 610*0

_80_£_!0_ ^6 *b 6ZZ*0

080*0 <7l0*0 08T *0_ 5 £0*0 £8T*0

580*0 M0*0 T i l *0 AEO *0 8AI *0

EAO'O

ZIZ'O 8E0' 9AI'

901*0 910*0 6ZZ*0 WO *0 A£Z*0

- wnissviod - wniS3N9VW

wnmvo -SfldOHdSOHd - N390dilN

AIZ*0 zzo *0 BW *0 £Z0*0 ZAO *0

*<A0*0 ZZ0°O 590*0 ZZO'O */S0*0

190*0 TEO *0 ?60 *0 OZO °0 *50 *0

Z90*0 EZO*0 VAO *0 UC'O Z<70*0

8A0 *0 OZC'O 850*0 Z7.6~*0 £<70*0

£60 *0 I zo • c AAO *C ZZO *0 *>50*0

- wnissviod - KflIS3N9VH

-sn-aoHdSOHd - N390M1IN

01 6 A 5 £ 1 01 6 Z : I £ I „ NOI103S**iiaV8 * » N01i03S**OOOM o

31dWVS AB 1N31N0D !N'3HiinN lN33«3d*»»

S**/ 9*0Z i ' V lt<?'0 T S Z T 611 (W)*1 (H)*T (W)VIO (H> (H) *0N *0N 1108 NMOBD NMOdO H8J H19N31 S3 ID3dS 33U1

o V

r

TREE NO. 159

SPECIES NO. 2

LENGTH ( Ml

29.9

D8H ( M ) 0. 485

CROWN DIA(M) 8.1

CROWN BUTT L.IMI L.(M) 18.1 11.8

• ••PERCENT NUTRIENT CONTENT BY SAMPLE SPEC IFICATION****************•*•*•••***••*»**•****•«•**•*••»•**»••••••••*«**•*••••»»»*****•* • WOOD**SECT ION * * BARK**SECTI ON *

1 3 5 7 9 10 1 3 5 7 9 10

NITROGEN -JIHOSPHORUS; CALCIUM MAGNESIUM -POTASSIUM -

0. 048 jj.00_6_ 0.056 0. C03 0.075

0.047 _0.CJ7_ 6.064 0.011 0.054

0.053 . JJ.006 0.06 7 0.014 0.031

0.042 _0_.„005 6. 067 0. 012 0.076

0.050 _0.009 0.051' 0.010 0.061

0. 068 J ) . 010_ 0. 061 " 0.011 0. 07 0


0.2 54 0.037


0.631 0.018 0.112

0.203 _C_PJ33 0.596" 0.028 0.100

0.232 0. 048_ 0.632" 0.038 0. 127

0.254 0.044 6.540 0. 046 0.132

0.369 0.440 0. 056 0.066 0.795" 0.616" 0, 051 0.053 0.199 0.263

BRANCH BRANCH i - i / 4 >1

_ • - - - —

' (TOP) -JWIGS-( MID)

-----*_ "(LOW) (TOP I

-FOLIAGE-\ (MIDI "" (LOW)

CONES _* TWIGS* FOL I AGE * ""(TOP) """ (MID) "" (LOW)

NITROGEN - 0. 177 0.249 0.000 0.000 0.000 0. 000 0. 000 0.000 1.211 0.543 0. 881 0. 835 PHOSPHORUS-CALCIUM MAGNESIUM -POTASSIUM

0. 024 0. 282 0.024

0. 035 0.274 0. 030

0.099 0.123

0. 000 0.000 0.00 0

0. 000 0.000 0. 000

0.000 0.000 0.000

0. 000 0. 000 0.000

0.000 0. 000 0. 000

0.000 0. 000 0.000

0.216 0.030 0.075

0.000 0.000 0.000 0.000 0.000 0.000 0.468

0.076 0.509 0.063 0.442

0.117 0.123 0.536 0.534 0. 064 0.063

0.397 0.383

• ••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPECIFICAT] »•••*•«••••••

-W000+* SECTION— -BARK**SECTION-TOTAL 162 364 566 768 10 TOTAL 162 364 566 768 T O -

NITROGEN - 15.97 11.32 7.26 3.8 8 0.99 0. 3e 40. 30 NITROGEN -PHOSPHORUS-CALCIUM MAGNESIUM -

2.15 20.15 3,24

1.50 15.47 2. 89

0.85 10.23 1.96

0.59 4.97 0.91

0.17 1.00 0. 19

0.05 0.34 0.06

5. 32 52. 15 9.25


17.42 13.78 13.18 2. 68

46.77 1. 78

2.55 38.88 2.11

2.48 31.76 2.30

8.48 1. 36

18.14 1.32

POTASSIUM - 21.56 15.91 11.96 5. 75 1. 0.39 56.76 POTASSIUM - 8.06 7.18 7.03 4.50

2.11 "0."3'2 4. 53 0. 29 1. 13

1.14 56.11 **07i7 '9757"" 1.59 141.67 0.14 7.93 0. 68 28. 59

BRANCH BRANCH •- -TWIGS * * >1

NITROGEN - 9.46 10.44 0. 00 0.00 0. 00

FOLI AGE -1-1/4 (TOP) (MID) (LOW) TOTAL (TOP) (MID) (LOW) TOTAL

0.00 0.00 0.00 0. 00

CONES • TW I GS • FOL I AGE — (TOP I (MID) (LOWI

0.00 0.00 15.87 18.33 7.68

TOTAL"

41.88

TREE "TOTAL

158.19 PHOSPHORUS- 1. 30 1.47 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0. 00 2.23 2. 43 *"", 1 3" 5*779 2"2.44 CALCIUM - 15.05 11.46 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 14.84 11.15 4.91 30.91 251.24 MAGNESIUM - 1.29 O._00'_ 0.00_ 0.00 0.00 0.00 0.00 0.00 0.00 0. 00 _ 1.85 1. 33 0.58 3. 76 23. 51 POTASSIUM - 5. 26 5. 36 0.00* 0.00 0. OO" "6.66 0.66" 0.00"* 0.00 0.60 "070"0" "12.90" 7.97 3.65 " 24. 53 " 1 20.5 0

TREE N O . 1 7 5

S P E C I E S N O .

2

L E N G T H (Ml

3 8 . 4

OBH (Ml

0 . 7 1 1

CR OWN 01 A I M )

8 . 5


2 8 . 9 9 . 6

• • • P E R C E N T N U T R I E N T CONTENT BY SAMPLE S P E C I F I CAT I 0 N » * * * * * * * * * * * * * * * * * * # * * * * * * * * » * * * * * « » « ^ ^ * WOOD*^SECT ION * * B A R K * * S E C T ION »

1 2 5 7 9 10 1 3 5 7 9 1 0

N I T R O G E N -P H O S P H O R U S -

0 . 0 4 4 _0_. 0 0 3

C A L C I U M MAGNESIUM POTASS IUM

0 . 0 6 2 0 . 0 1 2 0 . 0 5 3

0 . 0 4 5 <L.°OJ_ 0 . 112 0 . 0 2 5 0 . 0 8 1

0 . 0 5 3 0 . 0 0 9 0 . 0 6 7 0 . 0 1 3 0 . 0 5 1

0 . 0 3 8 _0_.J3 03_

0 . 0 7 6 0 . 0 1 3 0 . 0 5 4

0 . 0 6 1 0 . 0 1 0 0 . 0 4 9 0 . 0 1 1 0 . 0 6 7

0 . 066 0 . 0 1 1 0 . 061 0 . 015 0 . 0 6 4

N ITROGEN -PHOSPHORUS-

0 . 2 3 8 0 . 0 3 8

0 . 2 3 4 0 . 0 5 0

0 . 2 7 2 0 . 0 5 7

C A L C I U M MAGNESIUM -P O T A S S I U M -

0 . 4 6 5 0 . 0 1 9 0 . 1 3 4

0 . 4 3 1 0 . 0 3 3 0 . 1 4 3

0 . 4 1 1 0 . 0 4 3 0 . 126

0 . 2 6 4 0 . 0 5 9 0 . 4 1 8 0 . 0 4 4 0 . 165

0 . 4 0 0 0 . 075_

' 0 . 4 1 5 0 . 0 4 5 0 . 199

0 . 4 1 5 _ 0 . 0 83_

6 . 321 0 . 0 4 5 0 . 2 5 2

BRANCH BRANCH >1 1 - 1 / 4 ( T O P )

- T W I G S -( MID) (LOW)

- F O L I A G E -IT OP) (MID) I LOW)

CONES * TWIGS *FOLI AGE (TOP I "(MID)' ( LOW)

N I T R O G E N - 0 . 1 6 0 0 . 2 4 6 P H O S P H O R U S -C A L C I U M MAGNESIUM -P O T A S S I U M

0. 02 5 0 . 252

_ 3 . 3 2 7_ 0 . 101

0 . 0 3 6 0 . 2 3 9 0 . 0 2 9 0 . 1 3 7

0 . 0 0 0 0 . 0 0 0 0 . 0 0 0 0 . 0 0 0 0 . 0 0 0

o. oop_ o. odo o. obo 0 . 0 0 0

0 . 0 0 0 0. 0 0 0 0 . 0 0 0 0 . 0 0 0 1 . 5 6 1 0 . 7 5 9 , 0 . 501 0 . 0 0 0 0 . 0 0 0 0 . 0 0 0

0 . 0 0 0 o. ooo 0. 0 0 0

0 . 8 3 3 0 . 0 0 0 0 . 0 0 0 0 . 0 0 0

0 . 0 0 0 0 . 0 0 0 0 . 0 0 0 0 . 0 0 0

0 . 0 0 0 0 . 0 0 0 0 . 0 0 0 0 . 0 0 0

0 . 2 7 7 0 . 0 3 0 0 . 088 0 . 734

0 . 1 0 6 0 . 4 4 3 0 . 0 7 6 0 . 3 2 8

0 . 0 9 2 0 . 1 11 0 . 4 6 2 0 . 4 9 4 0 . 0 6 0 0 . 0 5 8 0 . 2 76 0 . 3 0 6

• ••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE S P E C IF ICAT I 0N + * * * * * * » •••********•***•*>*• * * » * * » * * * * » * « * " " ^ ^

- W P 0 0 ^ * S E C T I 0 N -

_NI T R 3 G E N P H O S P H O R U S -C A L C I U M MAGNESIUM -

1£2

2 . 2 6 6 4 . 3 1 1 4 . 0 3

3 £4

3 . 1 0 4 7 . 3 6 1 0 . 2 7

5 £6

3 3 . 0 3 2 6 . 0 0 1 5 . 6 6

7E8

9 . 0 8 1 . 9 7

2 4 . 3 1 4 . 5 0

1 . 1 9 1 1 . 4 4

2 . 2 3

2 . 3 1 0 . 3 9 1 . 8 7 0 . 4 3

1 0

J?i6JL 0 . 1 1 0 . 6 2 0 . 1 5

TOTAL

8 6 . 7 6 9 . 02

1 5 0 . 4 1 3 1 . 6 0

P O T A S S I U M - 5 0 . 1 0 3 5 . 0 5 1 7 . 9 2 1 1 . 0 4 2 . 5 3 0 . 6 5 1 1 7 . 30

- B A R K * » S E C T I O N -1£2

P H O S P H O R U S -C A L C I U M MAGNESIUM -

3 £4

N ITROGEN - 3 7 . 25 2 2 . 8 2 6 . 9 8 4 . 8 3

7 0 . 80 3 8 . 0 0 4 . 0 8 3 . 4 4

5 £6

J 4 . 7 0_ " 3 . 1 8

2 2 . 7 5 2 . 3 9

7 £ 8

_ 8 ' 0 3 _ " " 1 . 6 2 1 0 . 0 7

1 . 0 7

9 1 0

2 . 5 6 1 . 0 0 0 . 4 8 0 . 2 0 2 . 6 5 0 . 7 7 0 . 2 9 0 . 1 1

TOTAL

8 6 . 3 5 1 7 . 2 8 "

1 4 5 . 0 5 1 1 . 3 8

P O T A S S I U M 2 1 . 9 0 1 2 . 1 3 7 . 9 8 4 . 4 0 1 7 2 7 0 . 6 0 4 8 . 2 9

BRANCH J 3 R A N C H _ ">1 1 - 1 / 4 ( T O P )

- - T W I G S • (MID) "(LOW)

* *. TOTAL ( TOP")

- F O L I A G E - - - - -(MID) " (LOWI «

" T O T A L "

_CONES_ • - TWIGS* FOL I AGE — " ( T O P ) " (MID) " (LOW) T O T A L "

T R E E TOTAL"

N I T R O G E N - 2 7 . 76 1 8 . 11 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 5 1 . 5 9 3 9 . 32 3 7 . 0 8 1 2 7 . 9 8 P H O S P H O R U S - 4 . 37 2 . 6 6 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 7 . T 8 7 , 22 C A L C I U M - 4 3 . 32 1 7 . 6 1 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 3 0 . 1 2 3 6 . 2 8

_M.AGNESXU_M_J: 4._7.1_ 2 . 1 4 0 . 0 0 _ 0 . 0 0 _ 0 . 0 0 _ 0 . 0 0 ^ 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 5 . 1 4 4 . 74 P O T A S S I U M - 1 7 . 5 7 1 0 . 1 1 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 ' " " " 6 . 0 0 ' 0 . 0 0 " " 6 . 0 0 " 0 . 0 0 "0". 0 0 " ' 2 2 . 3 3 ' " 21". 64 '

4 . 9 3 1 9 . 3 3 2 1 . 9 9 8 8 . 3 9

2 . 5 9 1 2 . 4 8 1 3 . 6 1 5 7 . 5 8

3 4 6 . 9 6 5 2 . 6 7

4 4 5 . 2 8 6 2 . 3 1

"2 5 0 . 8 5

TREE SPECIES LENGTH DBH CROWN CROWN BUTT NO. NO. (M) (M) DIA(M) L.(M) L.(M) 176 2 26. 3 0. 361 0.0 19.6 6.6

*** PER CENT NUTR I ENT CONTENT BY SAMPLE SPECIF I CANTON ********************************** ***'"* * WOOD** SECT I ON * * BARK** SECT I ON *

1 3 5 7 9 10 1 3 5 7 9 10

NITROGEN - 0. 063 0.041 0.045 0.050 0. 059 0. 063 NITROGEN - 0.242 0.226 0. 246 0. 265 0. 359 0.409 PHOSPHORUS- 0. Oil 0. 004 0. 007 0.008 0. 009 0. 012 PHOSPHORUS- 0.041 0.044 0.051 0. 053 0. 074 0.095 CALCIUM 0. 043 0.061 0.060 0. 059 0. 060 0. 051 CALCIUM 0.675 0.480 0.482 "0. "591" b. 413 0.550 MAGNESIUM - 0. 007 0. OIJ 0.007 0. 009 0. 009 0. 010 MAGNESIUM - 0.010 0.022 0.022 0. 023 0. 02 5 0.021 POTASSIUM - 0. 054 0.054 0. 059 0. 065 0. 073 0. 073 POTASSIUM - 0.097 0.158 0.134 0. 146 0. 143 0.237

BRANCH BRANCH -TWIGS-- --* -FOLIAGE- CONES * TWIGS+ FOLIAGE---* >1 1-1/4 (TOP) (MID) ( LOW) (TOP) (M ID) (LOW) (TOP) (MID) ( LOW)

NITROGEN - 0. 00) 0.000 0. 000 0.000 0. 000 0. 000 0.000 0.000 0. 000 0.000 0. 000 0. 000 PHOSOHORUS- 0. 000 0. 000 0. 00 0 0. 000 0. 000 0. 000 0.000 0.000 0.000 O.COO 0.000 0. 000 CALCI'JM 0. 000 ' 0.000 0.000 0. 000 0. 000 0. 000 0. 000 0. 000 0. 000 0.000 0.000 0. 000 MAGNF SIUM - 0. 000 0.000 0. 000 0.000 0. 000 0. 000 0. 000 0.000 0.000 0.000 0.000 0. 000 POTASSIUM - 0. 000 0.000 0. 000 0. 000 0. 000 0. 000 0. 000 0.000 0.000 0.000 0.000 0. 000


a WCOD**SECT ION * * BARK**SECTION * 162 3£4 5C6 768 9 10 TOTAL 112 3C4 5C6 7C8 9 10 TOTAL

NITROGEN - 9.25 4. 64 3.19 1.95 0.41 0.10 18. 55 NITROGEN - 6. 58 3.76 2.74 1.50 0.43 0.16 15. 17 PHOSPHORUS-CALCIUM "AGNESIUM -

1.17 9.25 1.27

0. 56 6. 55 0.90

0.49 4.01 0.55

0.2 9 2.14 0.3 3

0.06 0.42 0.07

0.02 0.08 0.02

2.60 21.46 3. 13


1.20 16.23 0. 44

0.76 7.65 0.35

0.55 5.74 0.24

0.30 2.41 0.11

"OTO'9" 0. 5 0 0.03

0.0"4 0.22 0.01

2.94 32.74 1.19

POTASSIUM - 3. 53 6.09 4.19 2.49 0.51 0. 12 21.97 POTASSIUM . 58 2.32 1.49 0. 69 0. 17 0.09

BRANCH BRANCH *- -TWIGS-

NITROGEN

>1

0.00

1-1/4 (TOP) (MID) (LOW) TOTAL - - _- FOL I AG E "(TOP) (MID) (LOW)

_«. CONES * TWI GS«-FOL I AGE * TOTAL ' "(TOP)'"""( MID)' (LOW) TOTAL

0.00 0.00 0. 00 0. 00 0.00 0.00 0. 00 0.00 0.00 0.00 0.00 0.00 0.00 0.00

e.35

TREE "TOTAL""

33. 72 PHOSPHORUS- 0. 00 0. 00 0. 00 0.00 0. 00 0. 00 0. 00 0.00 0.00 0.00 0. 00 0.00 0.00 0. 00 0.00 5.53 CALCIUM 0. 00 0. 00 0.00 0. 00 0. 00 0. 00 0. 00 0. 00 0.00 0.00 0.00 0.00 0.00 0. 00 0.00 54.20 MAGNESIUM - 0. 00 0. 00 0.00 0.00 0. 00 0. 00 0. 00 0. 00 0.00 0.00 0. 00 0.00 0.00 0. 00 0.00 4.32 POTASSIUM - 0. 00 0. 00 0.00 0.00 0. bo" 0. 00 b. bo " 0.00 0.00 0. 00 ' 0. 00 " "0.00 " "0. 00 0. 00" " " 0.00 30.32"

TREE NO. 178

SPECIES NO. 2

LENGTH (Ml

33.9

DBH (Ml

0.884

CROWN DIA(Ml 0.0

CROWN L.IMI 28.7

BUTT L. (M) 10.2

•••PERCENT NUTRIENT CONTENT BY SAMPLE SPEC IF I CAT 10N»**«»*»«»»»»» ••*»*»•*••»«••••»»•••••••••»»»••••••»»»>

• -NITROGEN -J?HO_S_PHOR_US:L_ CALCIUM MAGNESIUM -POTASSIUM -

0. 067 _9i00_9_ 6. 04d O.OOd 0. 043

0.037 _9 .Q_05_ 6.076 0.013 0.048

-WOO0**SECTI0N * _> 7 9 10

0.036 0.005

0. 052 0. 006

0.095 0.013

0. 071 0.01 5 0.05 6

0. 058 0. 013 0. 093

0.080 0.025 0.1 39

0. 064 _0. 041_ 0."654 0. 012 0.064

• — —


0.303 0. 045 0.496 0.023 0. 130

• • * • « • « • * * • • • • • » « • • • • • • A A A A

8ARK^*SECTI0N * 5 7 9 10

0.172 0.035 673 53 ' 0.030 0.109

0.168 0. 038_ "67313 0.025 0. 100

0.180 0.035_ ' 0.314 0.031 0.115

0.214 0.376 _0. 034_ _;0.O83_ 0.319 " 0.427 0.045 0.040 0. 172 0.267

BRANCH BRANCH >1

NITROGEN - 0.000 PHOSPHORUS-CALCIUM .MAGNESIUM -POTASSIUM -

0. 00 J

0.000 0.000

1-1/4

0.000 0.000 0.000

• TWIGS * (TOP ) (MID)

FOL I AGE • (LOW) (TOP )

0.000 0. 000 0.000

0. 000 0. 000 0. 000

0.000 0.000 0.000

o.ooo ' o.ooo o.ooo 6.o66~ o.ooo

0. 000 0. 000 0. 000

(MID)

0.000 0.000 0.000 0.000 0. 000 0. 000 0.000 0. 000 0. 000

( LOW)

0.000 0.000 0.000 0.000

0.000 0.000 0.000

CONES

0.000 0.000 0.000

_* TWJ.GS+FOLIAGE--(TOP I (MID)

0.000 0.000 0.000

o.ooo o.ooo"

I LOW)

0.000 0.000 0.000 0.000 o.ooo o.ooo o.ooo o.ooo 0.000 0.000 0. 000 "o.ooo"

•••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPECIFICATION*****""""""""***

• WOOO**SECTI ON • • 162 3 64

• 5 66 768 10 TOTAL 162

BARK**SEC TION 3 64 566 768


_6_*b_18 30.09 25.22 2 0.57 2.27 3.13 75.36 12.33

4.10 61.11 11.31

3.35 36.69 8.05

2.68 19.19 5.27

0.31 1.92 0.60

_0.26_141.60_ 0.T7" ' 1 8. 74 0,22 194.99 0.05 38.59

NITROGEN - 57.64 31.28 34.87 19.98 2.05 PHOSPHORUS- 9.71" 6".76" '7.36 3751 0."33" CALCIUM - 102. 88 61.28 62.91 32.13 3. 06 MAGNESIUM - 6.40 5.16 5.69 3.86 0. 43

POTASSIUM - 53.32 43.30 42.70 32.31 3.31 . 0.26 180.20 POTASSIUM - 29.00 19.22 21.57 14. 59 JTT!

10

0.68 0.15 0. 77 0.07

TOTAL

146.49 27. 32""

263.03 21.62

J ^ A N 1 " ! 3RANCH_*R-:---------TWIGS • • >1 *"' 1-1/4 (TOP) (MID) (LOW) TOTAL (TOP)

^ F O L I A G E - — - * CONES * T Wl GS* FOL I AGE--(MID) (LOW) TOTAL "'" ( TOP) (MID) (LOWI

NITROGEN - 0.00 PHOSPHORUS-CALCIUM _MAGNES IUM ^ POTASSTUM -

0.00 0.00 0. 00 0. 00 0. 00 0. 00 0. 00 "o'. 00*

0. 00 0.00 0. 00 0. 00*

0. 00 0.00 0. 00 0. 00 6. oo •

0. 00 0.00 0.00 0.00 0. 00 0. 00 _ 0. 00 6. o o 6 . oo

0.00 0.00 0.00 0.00 0.00 0.00 0.00 0. 00 6.00* " 0.00

0.00 0.00 0.00 0.00 0.00 0.00 "0.00*"" 0.00"

0.00 0.00 0.00 0.00 0. 00 0.00 0.00 "0. 00

0. 00 0. 00 0.00 "0. 00"

0. 00

0. 48

TOTAL

0.00 0.00 0.00 0.00 0.00

86. 52

TF EE TOTAL"

288. 09 0, 00 0.00 0.00 0.00 0.00 0.00 0. 00 * 0. 00

0.00 0.00 0.00 0.00

4 6.55 458.02 60.21

266. 7?

T R E E

N O .

2 1 3

S P E C I E S

NO. 2

L E N G T H

(Ml 30.0

0 3 H

( M )

0 . 5 6 4

C R O W N

01 A( Ml 5 . 6

C R O W N B U T T

L . ( M ) L . I M I

2 5 . 1 4 . 9

«««p£RCgNT NUTRIENT CONTENT BY SAMPLE SPECIFICATION*"****************************^ • WOOD**SECT ION * * BARK**SECT ION *

1 - 3 5 7 ? 10 1 3 5 7 9 10

0.046 0.052 0.063 0. 076 NITROGEN - 0.259 0.230 0.230 0.306 0.365 0.419 _0_.006 Oj.008 0..014 0_.01_2 PHOSPHORUS- 0.036 0.040 _ 0. 041 0.052 _ 0.062 0.072 0. 072 6.057" 0.057 6.054 CALCIUM - 6.666 0.621" "0.515 0.690 0.579 0.463 0.011 0.009 0.009 0.009 MAGNESIUM - 0.016 0.026 0.031 0.032 0.034 0.031 0.063 0.048 0.061 0.065 POTASSIUM- 0. 132 0.117 0. 148 0.131 0.172 0.226

NITROGEN -_P_HQ_$ PJ±ORUi^_ CALCIUM MAGNESIUM -POTASSIUM -

0.054 0.0 07_

051 0 0. 011 0. 061

0.047 0.006_ 6.05_4 0.010 0.055

BR ANCH BRANCH

NITROGEN

> 1

0. 172 PHOSPHORUS-CALCIUM MAGNESIUM -

0. 022 0. 330 0.020

1 - 1 / 4

0.209 0.031 0. 313 0.022

( T O P )

0.000

-TWIGS-( MID) ( LOW)

0.000 0.000

(TOP I FOL I A G E -

(M I D I

0. 000 0. 000

(LOW)

0.000

CONES * TWIGS + FOL I AGE • "" '"""(TOP) "• (MID) (LOW)

1.493 0.795 0.829 0.829 0.000 0. 00 0 0 . 6b o

0. 000 0.000 0. 000

0.000 0.000 0.000

0.000 0. 000 0. 000

0. 000 0. 000 0. 000

0.000 0.000 0.000

0. 261 0.025 0. 076

0.106 0.398 0.062

0. 099 0.099 0.510 0.510 0.058 0.058

P O T A S S I U M - 0.090 0.103 0.000 0. 000 0.000 0.000 0. 000 0.000 0.564 0.384 0. 341 0.341

~**VNUTRIENT A M O U N T S I N O E C A G R A M S B Y S A M P L E S P E C I F I C A T I O N * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * R * * * * * * * * * * * * * ^ * * * *

- W O O O * * S E C T I O N - B A R K » * S E C T I O N -

TC5 T O T A L 1 C 2 3 C 4 5 £ 6 768

N I T R O G E N 19.31 12. 05 7.70 4.43 0.80

10

0 . 1 9

TOTAL

4 4 . 9 9 NITROGEN -

1£2


2.53 1.65 1.09 20.35 16.39 10.20 3.95 2.66 1.56

0.33 0.18 4.36 0.72 0.6 7 0.11

0.03 0.14 0.02

6.30 52.66 8.97


3.51 59.22 1.92

3 £4

11.16 — n s T " 27.57 1.38

5 £6

10.41

T O "

TT&TT 23.33 1.22

5^22 1. 42 0.8 9 "0.24 9.84 2.26 0.52 0. 13 2.35 0.67

0.59 51.32 "OTTO -"e."5i"" 0.65 122.86 0.04 5.22

POTASSIUM - 22.74 15.39 8.35 4.22 0.78 0.16 52. 14 POTASSIUM - 11.46 6.42 5.39 0.32

BRANCH BRANCH *- -TW1GS->1 1 - 1 / 4 (TOP) (MIDI (LOW) TOTAL ( T O P )

^FOLIAGE--— (MID) (LOW)

- * _ C O N E S _ _

" T O T A L

TWIGS+FOL1AGE--"(TOPT"""(MID)' ' (LOW)" TOTAL"

NITROGEN 15.81 10.74 0, 00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 2 5.92

12.98 2.03

12.51

43. 17 — 57

26. 18

26.58 16.1 2 3.00 1.82

"17.75 -"10.77

9 5 . 2 7

T T T 7 T "

26.60

TREE "TOTAL"

218.13 PHOSPHORUS-CALCIUM _MAGNE SIUM -POTASSIUM -

2. 05 1. 57 0. 00 35.04 16.04 0.00 1.35 1. 1 4 0.00 8.25 5.55 0.00"

• 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 _o1oo 0 . qo_

"0.06 o.'oo"

0 . 0 0 0 . 00 0 . 0 0 0 . 0 0 o.oq 0 . 0 0 "oio'o 6 . 0 0 '

0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 __o_oo_ 0 . 0 0 0 . 0 0

0.66 0.06 0 . 0 0 0 . 0 0 0 . 0 0 0 . 00

"0V00 o.oo

55.68 282.29 6. 65 24. 03

"41.04" 133. 58"

TREE SPECIES LENGTH OBH CROWN CROWN BUTT NO. NO. (Ml (Ml DIAIMI L.(M) L.(M) 215 2 38.2 0. 805 0.0 25. 7 12.5

•••PERCENT NUTRIENT CONTENT BY SAMPLE SP EC IF I CAT I ON*** ******************* ****^^ * -WOOO**SECTION * * -8AP.K**SECTI0N *

1 2 : § 7 9 10 1 3 5 7 9 1 0

NITROGEN - 0. 061 0.044 0.046 0.045 0.086 0. 085 NITROGEN - 0.267 0.194

J

0. 208

1

0.216

1

0. 307

I u

0.391 PHOSPHORUS- 0.003 0. 006 0. 009 0.009 0.015 0. 021 PHOSPHORUS- 0.039 0.038 0.037 0.043 0. 041 0. 075 CALCIUM 0.055 0.055 0.055 0.071 0.090 0. 060 CALCIUM 0.265 0.350 0.296 0.311 0. 743 0.356 MAGNESIUM - 0.C09 0.003 0. 01 0 0. 012 0.017 0. Oi l MAGNESIUM - 0. 016 0.020 0. 022 0.031 0. 030 0. 029 POTASSIUM - 0. 051 0.042 0.042 0.054 0.060 0. 058 POTASSIUM - 0.121 0.099 0.083 0.090 0.093 0.196

BRA MCH BRANCH —TWIGS- -FOLIAGE- CONES * Tul r.^+Fni T AGP * >1 1-1/4 ( TOP 1 ( MIDI ( LOW) (TOP) (MIDI (LOW 1 ( TOP 1 (MIDI ( LOW)

NITROGEN - 0. 000 0. 000 0.000 0. 000 0.000 0. 000 0.000 0.000 0.000 0.000 0. 000 0. 000 PHOSPHORUS- 0. 00) 0. 000 0. 00 0 0. 000 0.000 0. 000 0.000 0.000 0.000 0.000 0.000 0.000 CALCIUM 0. 000 0. 000 0.000 0. 000 0.000 0. 000 0. 000 0. 000 0. 000 0.000 0.000 0.000 MAGNESIUM - 0. 000 0. 000 0.00 0 0. 000 0.000 0.000 0. 000 0.000 0. 000 0. 000 0. 000 0. 000 POTASSIUM - 0. 000 0. 000 0.000 0.000 0.000 0. 000 0. 000 0.000 0. 000 0.000 0.000 0.000

• ••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPEC IF ICAT'l0N*"*>*************** *******"*T*~*T^

-WOOO»«SECTION-1£2 3£4 5(16 7£8 10 TOTAL

-BARK**SECT ION-1£2 3C4 5 C 6


41.12 25.73 17.46 13.07 2.93 5.43

42.61 6.85

4. 20 31.42 5.39

0.44 100.75 3.53

24.48 4.38

2.45 16.12 2.96

0.51 3.06 0.59

POTASSIUM - 36.20 24.01 18.58 11.36 2.04

0.11 0.31 0.06

NITROGEN - 47.51 31.73 34.34 16.28

118.00 20.23


7.93 5.97 63.40 50.98 3.69 3.28

0.30 92.49

6.52 49.12 4.30

SRANCH_ BRANCH J*j; " > 1 1 - 1 / 4

~z—r—TWIGS — * * (TOPI" TMlb") (LOW I " TOTAL ' (TOP)

POTASSIUM -

--FOLIAGE—— (MID)" (LOW I

7£8 9

_20.63 3.99 3.32 0.53"

41.59 9.66 2.43 0.40

10

0.47 0.0'9 0.43 0.03

22.68 14.36 14.01 7.26

TOTAL

1 38. 66 ~ 24."37 215.17 14.12

1.21

* _CONES T O T A L * "

• TW I GS*FOL I AGE — "(TOP) (MID) " (LOW)

NITROGEN - 0.00 0.00 PHOSPHORUS-CALCIUM _M_AGMES IUM_ _ POTASS I UM -

0. 00 0.00 0.00 0. 00 0. 30 0. 00 o". 66

o. oo o. oo o. oo o. oo"

0.00 0. 00 0. 00 0.00 6. oo"

0.00 0.00 0,00 0.00 0. 00 _ 0. 00 b.oo". "6".o6r

0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 "" 0.00

0.00 0.00 0.00 0.00 0.00 0.00 0.00 "0.00 "0.00

0.00 0.00

0.24

•

TOTAL '

0.00 0.00 0. 00 0.00 0.00 0.00 0.00 "0.00 "' "0.00"

0.00 0.00 0.00 0.00 *

0.00 OT075 0.00 0.00 0.00 0.00 0.00 0.00

59.76

TREE TOTAL"

239.42 0.00 0.00 0.00 0.00

40.65 333. 18 34.35

"152.25

TREE NO. 235

SPEC!ES NO. 2

LENGTH < M )

15.1

09H (M)

0.3*3

CROWN DIA(Ml 0.0

CROWN L.(M)

8.4

BUTT L. (M)

6. 8

• ••PERCENT NUTRIENT CONTENT BY SAMPLE SPEC I F I CAT I ON********************************************** • WOOD** SEC T I ON * * B ARK**SECT I ON *

1 3 5 7 9 10 1 3 5 7 9 10


0.053 JJ.Ol 0_ 0. 052 0.009 0. 053

0.051 _0.0 07_

0. 06 0 0.011 0. 050

0.049. _0_.0i)6_ 0. 06 7 0.012 0.054

0. 061 J).008_ 0. 062 0. 013 0. 055

0.054 _0_.009 0.054* 0.010 0.062

0. 066 0. 012_

"0.~"053 0. O i l 0.069


0.229 0. 037_ 0.627 0. 02 0 0. 122

0.224 _0_039_ 0.583 " 0.029 0.112

0. 251 0. 046

' 0.492" 0.042 0. 157

0.254 0.043 "O". 431 0.037 0. 182

0.313 0. 056 0.418"' 0.037 0. 180

0.377 0.058

" 0. 6 34" 0.038 0.217

_BR_ANCH_ >1

_3RANCH_ 1-1/4 (TOP)

-J WIGS-(MID)

* (LOW) (TOP)

-FOLIAGE-(MID) (LOW I

CONES * TWIGS* FOLI AGE * "(TOP)" (MIDI " (LOW)

NITROGEN - 0. OOP 0.000 0.00 0 0. 000 0.000 0. 000 0. 000 0.000 0.000 0.000 0.000 0.000 PHOSPHORUS-CALCIUM MAGNESIUM -POTASSIUM

P. 000 0. 000 0.000

0.000 0.000 0.000

0.000 0.000

0.000 0. OJO 0.000

0. 000 0. 000 0. 000

0.000 0.000 0.000

0. 000 0. 000 0. 000

0. 000 0.000 0. 000

0.000 0.000 0. 000

0. 000 0. 000 0. 000

0.000 0.000 0.000

0.000 0.000 0.000 0.000 0. 000 0. 000 0. 000 0.000 0.000

0.000 0.000 0.000 0.000 0.000 o_oo_o_

0.000

• ••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPECIFICAT ION*********************************************************************»**»*

-WOOD««SECTIQN- -BARK**SECTION-162 364 566 768 9 10 TOTAL 162 364 5 66 768 9 10 TOTAL

NITROGEN - 4.65 3.33 2.68 1.57 0.44 0.10 12.77 NITROGEN - 3.67 3.61 2.63 2.18 0. 44 0. 19 12.71 PHOSPHORUS-CALCIUM MAGNESIUM -

0.77 5.02 0. 94

0.43 4.22 0.79

0.34 3.14 0. 62

0.24 1.58 0.32

0.08 0.44 0.08

0.02 0.08 0.02

1.87 14.48 2.76


0.61 9. 80 0.40

0.65 8.16 0.54

0.47 4.80 0.41

0.38 3.27 0.29

0.08 0, 59 0. 05

0".03 0. 32 0. 02

" 2 . 21" 26.93

1. 72 POTASSIUM - 4.36 3.48 2.66 1.59 0.50 0.10 13.19 POTASSIUM - 1.90 2.04 1.76 1.39 0. 25 0.11 7.46

BRANCH BRANCH * TWIGS->1 1-1/4 (TOP) (MID) (LOW) TOTAL

FO LIAGE — (TOP) (MIO) ' (LOW)

TOTAL CONES *- TWIGS* FOL1 AGE —

(TOP )" (MID) (LOW) TOTAL"

TREE TOTAL"

NITROGEN - 0. 00 0. 00 0. 00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0, 00 0. 00 0.00 25.48 PHOSPHORUS-CALCIUM MAGNESIUM -POTASSIUM -

0. 00 0. 00 0. 00 0.00

0. 00 0. 00 3. 00 0. 00

O.OP 0.03

_0. 00 o. do"

0.00 0.00 0.00 0.00

0. 00 0. 00 0.00 0.00

0.00 0.00 q.oo_ "b.dd

0. 00 0. 00

_0.00_ 0. 00

0. 00 0. 00 0.00

0.00 0.00 0.00

0.00 0.00

0.00 0.00 0.00

"0.00"

0.00 0. 00

o.'oo

0.00 0.00 0.00 0.00

0.00 0.00 0.00 0.00 0, 00 0. 00

"0.00 0.00"

"7j~70"0" 0.00 0.00 0.00

4.08 41.41

4. i.8 "20.65"

f ' r

T R E E

N O .

2*3

S P E C I E S

N O .

2

L E N G T H

( M »

17. a

O B H

( M )

0.310

CROWN 01 A(H ) 5.5

C R O W N B U T T

L . ( M ) L . I M I

11.5 6.3

•••PERCENT NUTRIENT CONTENT 8Y SAMPLE SPECIFICATION******************»»*****»************^ -WOOD** SECT I ON * * BARK** SEC TI ON *

_5 7 9 10 1 3 5 7 9 1 0 * V


0. 0*8 0.006


0. 060 0. 010 0.0*6

0.0*1 _0_._0_0*_ 6.05 7 0.012 0.061

0. 0*2 0. 00*

0. 0*9 0. 010

0.0*7 0.006

0.053 0. 013 0.055

0. 052 0. O i l 0. 062

0.050 0.012 0.055

0. 067 ___P_1_2_ 0. 0** 0. 01* 0. 067

NITROGEN -PHOSPHORUS-CALCIUM MAGNESIUM POTASSIUM

0.301 0.0*3 0. 5*5 0.026 0.085

0.236 0.036 0.629 0.029 0.101

0. 261 0.0*2_ "0.72 5 0.0*0 0. 131

0. 2 88 0.0** 0.5 99 0.038 0.151

0. *01 0.075 6. "76 2" 0.0*5 0.263

0. 522 0.091^ "6". 56"! 0. 0*7 0.271

BRANCH BR AN C H * T WIGS * *j- FOLIAGE » CONES * — - TW_I GS+ F OLIAGE — - * >1 1-1/* (TOP) (MID) (LOW) (TOP) (MID) (LOW) (TOP") " (MID) (LOW)

NITROGEN - 0. 127 0.213 0.000 0. (JOO 0.000 0.000 0.000 0.000 0.000 0.852 0.816 0.750 PHOSPHORUS- 0.021 0.032 0.000 0.600 0.000 0. 000 0.000 0.000 0.000 0.110 0.121 0.086 CALCIUM - 0.228 0.25* 0.000 O.dOO 0.000 .0. 000 0. 000 0.000 0.000 0.370 0.339 0.269 MAGNESIUM - 0.02* 0. 03* 0.000 0.600 0.000 0. OOP 0.000 0. 000 O.J)00 0.1 00 0. 087 0_. 067 POTASSIUM - 6.071 0.115 0.000 0.000 0.000 0. 000 0.000 0.000 0.000* 0.3*6" 07311 0.271

• ••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPECIFICATION************************************************"**"************'************

-WO00**SECTION- BARK**SECTION-162 36* 566 768 9 10 TOTAL 162 36* 566 768 9 10 TOTAL

NITROGEN - 3.*0 2. 10 1.3* 0.72 0.16 0.07 7. 80 NITROGEN - 3.36 3.13 2.17 1.21 0.28 0. 16 10.31 DHOSPHORUS-CALCIUM MAGNESIUM -

0.37 *.*6 o.a*

0.21 2.76 0.62

0.21 1.53 0.35

0.12 0. 76 0.17

0.02 0.17 0.0*

0.01 0.05 0.02

0.9* 9. 7* 2. 0*

PHOSPHORUS-CALCIUM MAGNESI UM -

0.50 7.3* 0.35

0.*9 8.53 0.**

0. 3* 5.23 0.31

0. 21 2.38 0.15

" ""6." 05 0. 53 0.03

0.03 0. 17 0.01

1.61 2*. 18 1.28

POTASSIUM - 4. J9 2.92 1.71 0. B7 0.19 0.07 9.85 POTASSIUM - 1. 17 l.*7 1.12 0. 72 0. 18 0.03

_8R_ANCH_BRANCH ~>V 1-1/'* (TOP)

- - T W I G S

(MID) " (LOW)" TOtAL (TUP)" • - F O L I A G E ; - ^ - - ;

' " ( "MIDT~'"""L0W» CONES *-

NITROGEN *. 29 3. 30 0.00 0.00 0. 00 0.00 0.00 0.00 0.00

TOTAL

0.00

-TWIGS + FOLIAGE— (TOP I (MID) " (LOW)

0.00 5.80 1*.53 *.05

TOTAL"

2*. 38

*. 7*

Tf.EE "TOTAL"

50.57 PHOSPHORUS-CALCIUM _MAGNES IUM_-POTASSTUM -

0. 73 0. 53 7.71 4.54 0.81__ 0.61 2. 41 2. 06

0.00 0. 00 0. 00 6. 00

0.00 0.00 0.00 0.00 0.00 0. 00 6.00 o.oo

0.00 0.00 0.00 0.00 0.00 0.00 o.ooP.OP

0.00 P. 00 0.00 0.00 0. 00 0.00 o.oo " o.oo"

0.00 0.00 0.00 0.00 0.00^ 0.00 "0.00 ** "o.oo"

0. 75" 2.51 0.68 2.36

2. 16 6.03 1. 56 5.5*"

"07*"7~ l.*5 0.36 "1. *6

T7TT 9.99 2. 60 9. 3 6

7.23 56. 16 7.3*

28.41"

http://Tf.EE

TREE NO. 2 5 6

S P E C I E S NO. 2

LENGTH ( M )

2 T . 9

DBH (M)

0 . 4 7 5

CROWN O I A I M ) 0.0

CROWN BUTT L . ( M ) L . ( M ) 2 0 . 3 7.6

•••PERCENT NUTRIENT CONTENT BY SAMPLE S P E C I F I C A T I O N * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * MOOD** SECT ION * * BARK **S ECT I ON *

1 3 5 7 9 10 1 3 5 7 9 10


0 . 0 5 5 0. 004

0 . 0 4 0 0 . 0 0 3

0 . 0 4 6 0 . 0 0 5

0. 044 0 . 0 0 8

C A L C I U M MAGNESIUM P O T A S S I U M

0 . 0 5 7 0 . 0 0 9 0. 049

0 . 0 7 0 0 . 0 1 0 0 . 0 4 9

0. 061 0 . 0 1 0 0. 0 5 4

0. 065 0. 0 1 0 0. 0 6 4

0. 0 5 1 _0 . 0 1 3 _ 0."0 57 0 . 0 0 9 0 . 0 6 0

0. 067 0.015


0.259 0.03 7

0.217 0.038

0. 048 0. O i l 0. 0 73

C A L C I U M MAGNESIUM -POTASSIUM -

0. 4 6 0 0.015 0. 0 9 9

0.348 0.025 0.086

0.226 0. 0 4 1 _

" 0 . 3 4 1 0.029 0.104

0 . 2 8 4 0.04 7 6. 524* 0.035 0. 143

0. 3 1 5 0 . 0 6 2 _ ""0.390 0.030 0. 1 5 7

0.454 _0.0 98 0 . 4 1 6 " 0.031 0.239

NITROGEN -

BRANCH BRANCH >1

0 . 0 0 0

1-1/4

0 . 0 0 0

( T O P )

0 . 0 0 0

-TWIGS-( M l b )

* 7 L b w ) "

o.ooo o.ooo

_* --j-FOL I AGE-(TOP) "(MID)' "

0. 0 0 0 0.000

(LOW I

0 . 0 0 0

CONES

0.000

* TWIGS+FOL I AGE * ""( TOP) " ' (MID) (LOW )

0 . 0 0 0 0.000 0 . 0 0 0

PHOSPHORUS-C A L C I U M MAGNESIUM -

0. 0 0 3 0 . 0 0 3 0 . 0 0 0

0 . 0 0 3 0 . 0 0 3 0 . 0 0 0

0 . 0 0 0 0 . 0 0 0 0 . 0 0 0

0. 0 0 0 0 . 0 0 0 0. 0 0 0

0 . 0 0 0 0 . 0 0 0 0 . 0 0 0

0.000 0. 0 0 0 0. 0 0 0

0. 0 0 0 0 . 0 0 0 0. 0 0 0

0 . 0 0 0 0 . 0 0 0 0. 0 0 0

0. 0 0 0 0. 0 0 0 0 . 0 0 0

0 . 0 0 0 0.000 0 . 0 0 0

P O T A S S I U M - 0 . 0 0 0 0 . 0 0 3 0 . 0 0 0 0 . 0 0 0 0 . 0 0 0 0.000 0 . 0 0 0 0 . 0 0 0 0. 0 0 0 0 . 0 0 0 0.000

0.000 0.000 0.000 0 . 0 0 0 0.000 ?5 0 0 0 _

0 . 0 0 0

• • • N U T R I E N T AMOUNTS IN DECAGRAMS BY SAMPLE S P E C I F I C A T I O N * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * *

* WOOD**SECTIGN * * B A R K * * S E C T I O N * 162 3 64

NITROGEN - 1 3 . 9 5 8. 51

5 66

5.92

768

3.42 0.55

1 0

0.18

TOTAL

3 2 . 5 4

162 3 £4

NITROGEN - 1 3 . 5 2 1 1 . 0 2

566

7.09

7£8

5.55

10 TOTAL

0. 5 0


1.31 1 3 . 7 1 2.79

3. 71 1 2 . 9 2 1 .95

0.83 8.25 1.28

0.7 5 4.35 0.7 0

0. 14 0.61 0. 10

0.04 0.13 0.03

3.49 4 4 . 9 8 6.85


2.14 2 2 . 94 1.13

"~T79'7" 1 7 . 12 1.35

~ 1T24"-1 2 . 0 3 0.90

"TToi o r n r 8.45 0.62 0.60 0.05

0.32 3 8 . 0 0 ""OTO 7 6 7 5 3 0.29 6 1 . 4 6 0.02 4. 05 0.17 1 6 . 6 1 P O T A S S I U M - 1 4 . 3 0 1 3 . 1 2 7. 73 4.44 0.65 0.20 3 7 . 4 4 P O T A S S I U M 5.24 4 . 7 3 3.45 2.7"8 0.2 5

BRANCH BRANCH * TWIGS- - F O L I AGE- CONES *•

NITROGEN

>1

0. 00

1-1/4 ( T O P ) ( M I D ) (LOW) TOTAL (TOP) (MID) " (LOW) TOTAL

0.00 0.00 0.00 0. 00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00

" 0 7 0 0 0700""

T W I G S * F O L I A G E ~ TP. EE (TOP) ( M I D ) (LOW) TOTAL "TOTAL

0.00 0.00 0. 0 0 0.00 7 0 . 5 4

o o o b

o o o o

o o o o

0 .00 0.00 0.00

"'"0. 00

0.00 0.00 0. 0 0 "0.00

0.00 0.00 0. 0 0 0.00

10 . 0 2 1 0 6 . 4 3 1 0 . 9 0 " 54". 05


0.03 0. 03 0. 00

0. 0 0 0. 0 0 0. 0 0

0.00 0.03 0. 0 0

PO T A S S I U M - 0 . 0 3 0.00 0.00 0.00

0.00 0.00 0.00 0.00 q. qq 0.00

o. bo"

0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 o.oo o.oo o. oo_ __o_ q_ 0.00 0 7 0 0 0.00 0.00

A P P E N D I X 7

N U T R I E N T CONCENTRATION AND N U T R I E N T D I S T R I B U T I O N

W I T H I N 1 3 SAMPLED T R E E S ON THE HANEY SAMPLE PLOT

S P E C I E S 1 - WESTERN HEMLOCK

S P E C I E S 2 - WESTERN RED CEDAR

S P E C I E S 3 - YELLOW CEDAR

TREE NO. 8

SPECIES NO. 3

LENGTH ( Ml

19.0

OBH IMI

0.231

CROWN 01 AIM) 5.8

CROWN L. (M) 13.5

BUTT L. I Ml 5.5

•••PERCENT NUTRIENT CONTENT BY SAMPLE SPECIFICATION******************************************************8

* WOOO**SECTION * * BARK**S ECT I ON * 1 3_ 5 7 __9 1_0 1 3 5 7 9 10

NITROGEN -_PHO_SPifOgJAS-„ CALCIUM MAGNESIUM -POTASSIUM -"

0.050 _0, 005_ 6. 107 0.003 0. 052

0.062 _0.006 0.0 94 0.009 0.050

0.05 3 _o.oi?_ 0.091 0. 009 0.057

0. 063 J>_. 014 6. 089 0. 009 0. 069

0.085 0. 076 _0.012 0.01 0_ 0.107 0.095 0.007 0. 007 0.058 0.068

NITROGEN -_£.H° SPHORUS-CALCIUM MAGNESIUM -POTASSIUM -

0.323 _P_._0_43_ 2. 570 0.048 0.280

0.270 0. 045_ 2.270" 0.050 0.266

0. 323 _0. 048 2.265 * 0.052 0. 375

0.4 01 0.055 1.705" 0.050 0. 325

0.433 _0o06 8_ 1.471 0.049 0. 350

0.533 0.061 T.2 56" o.04e 0.291

BRANCH BRANCH >1 1-1/4

-TWIGS- -FOL IAGE-(TOP ) (MID) ( LOW) (TOP) (MID) (LOW)

CONES _* TWIGS + FOL I AGE-(TOP) (MIO) ( LOW)

NITRQG;N !46 0. 243 0. 00 0 0. 000 0.000 0. OOP 0. 000 0.000 0.000 0.727 0. 752 0.728 PHOSPHORUS-CALCIUM _MAGN£S_IUM POTASSIUM

0.023 0.029 0. 747 0. 751 0. 023 0.028

0.000 0.00 0 0.00 0

0. 000 O.OOP 0. 000

0.000 0.000 0.000

0. 000 0. 000 0. 000

0. 000 0.000 0. 000

0.136 0.135 0.000 0.000 0.000 0.000 0.000

0. 000 0.000 0.000 0.000

0.000 0.000 0.000 0.000

0.079 1.142 _0.061 0.314"

0. 072 0.072 1. 196 1.407 0.054 0.066_ 0.237 0.2 81

***NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPECIFICATION**************************************************************************

* WO0D**SECTI0N * * BARK** SECT I ON * 162 364 5 66 768 10 TOTAL 162 364 566 768 10 TOTAL

NITROGEN - 3.00 PHOSPHORUS-CALCIUM MAGNESIUM -

0.31 5.37 0.43

2. 38 0.37 3.30 0.33

6. 39 2.66 0.25 "

0.19 1.39 0.12

0.19 0.03 0.23 0.02

0.03 8.37 NITROGEN 0.00 0.04 0.00

1.29 13.48 1.15


1.45 0.21 11. 86 0.24

_1_36_ 0.2"i"

10.43 0.23

1.05_ "0.15 5.76 0.15

0. 92_ "P."l4 3.49 0.11

POTASSIUM 2.71 2.19 1.36 0.91 0. 13 0.03 7.82 POTASSIUM 1. 34 1.47 1.02 0.74

0. 26_ "o; o"4 0= 88 0.03 0. 21

0.16 5.20 0"."0"2"" "0.7 7" 0.38 32.60 0.01 0.77 0.09 4.87

_B_ANCH BRANCH « TWIGS- _ * *_->'i 1-1/4 "(TOP) ("MID") ("LOW) TOTAL

--"-FOL I AGF * JCONES * TWI GS* FOL I AGE--: * (TOP)' (MIDI (LOW) "TOTAL" (TOP) (MIDI " (LOW) TOTAL"

TREE "TOTAL"

NITROGEN - 1. 03 3. 39 0.00 0.00 0,00 0.00 0.00 0.00 0. 00 0. 00 0.00 2.55 5. 87 2. 98 11 . 40 29.44 PHOSPHORUS- 0. 12 0. 39 0.00 0.00 0.00 0.00 0. 00 0. 00 0. 00 0. 00 0.00 0.28 0. 56 0. 30 i . 14 3. (i CALCIUM 3. 29 10. 28 0.00 0.00 0.00 0.00 0. 00 0. 00 0. 00 0. 00 0. 00 4.00 9. 33 5. 77 19. 10 78.95 MAGNESIUM - 0. 12 0. 38 0. 00 0. 00 0.00 0.00 0.00 0.00 0. 00 0. 00 0. 00 0.21 0.42 0. 27 0. 90 2.34 POTASSIUM - b." 60 1. 87 0.00 "o.oo 0. 00 0. 00 * 0.00 " " 0.00" "0. 00"" "" 0. oo "o.oo "1.10 1.85 " " 1 . 15 " 4. 10 "'" 19.26

f cv

r

TREE SPECIES LENGTH D8H CROWN CROWN BUTT NO. NO. (Ml (HI DIA(M) L.(M) L.(M) 50 1 17.1 0.160 4.9 14.3 2.9

• ••PERCENT NUTRI ENT CONTENT BY SAMPLE SPECIF i CATION ************* ****^ • WOOD**SECTION * * B ARK**SECT I ON *

1 3 5 7 9 10 1 3_ 5 7 ? lp_ NITR0G5N - 0. 043 0. 039 0.041 0. 049 0.055 0. 056 NITROGEN - 0. 166 0.210 0.233 0.2 82 0. 331 0. 352 PHOSPHORUS- 0. 003 0. 003 0. 01 0 0. 009 0.012 0. 013 PHOSPHORUS- 0.024 0.039 0.041 0. 054 Oc 063 0. 068 CALCIUM 0. 045 0. 043 0.042 0.046 0.048 0. 045 CALCIUM 0.412 0.502 0.295 0.314 0. 349 0. 3 37 MAGNESIUM - 3. 00 7 0. 003 0.008 0. 009 0.009 0. 009 MAGNESIUM - 0.016 0.023 0.025 0. 034 0. 045 0. 047 POTASSIUM - 0, 053 0. 055 0. 061 0. 064 0.0 54 0. 068 POTASSIUM - 0. 086 0.118 0. 184 0.195 0. 223 0. 2 63

BRA NCH BRANCH • —TWIGS-- • *- -FOLIAGE- • CONES * TWIGS*FOLIAGE — * >1 1- 1/4 ( TOP I (MIDI (LOW) (TOP) (MID) (Low) (TOP) (MID) ( LOW)

NITROGEN - 0. 117 0. 123 0. 589 0.437 0.515 0. 759 0.709 0.735 0.000 •6-W174 0. 573 0. bj&-PHOSPHORUS- 0. 019 0. 024 0. 080 0. 082 0.071 0. 128 0. 138 0.161 0.000 0.4 fT-*^ s0.396 > . -3C403 CALCIUM 0. 17 3 0. 179 0.305 0. 313 0.275 0. 432 0.479 0. 542 0. 0 00 0.368 0.409 MAGNESIUM - 0. 015 0. 021 0.05 5 0. 050 0.050 0. 105 0. 113 0. 121 0. 000 •m«085 POTASSIUM - 0. 072 0. 083 0.195 0. 168 0. 185 0. 334 0. 289 0.303 0.000 fl<?64 0.229 0?2>4»

•••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPECIF]

-W0OD*«SECTI0N- -BARK**SECTION-162 364 5 66 768 10 TOTAL 162 364 566 768 10 TOTAL

NITROGEN - l.OJ 0.72 PHOSPHORUS-CALCIUM MAGNESIUM -

0.21 0.16 1.24 0.81 0.19 0.14

0.60 0.13 0.53 0.11

0.34 0.08 0.07 0.02 0.31 0.07 0.06 0.01

0.03 2.85 NITROGEN 0.01 0.02 0.00

0. 59 3.04 Oo 51


0. 83 0. 14 2.01 0. 09

0.86 0.16 1.55 0.09

0.64 0.12 0.76 0.07

0.40 0.08 0.43 0.05

0.10 0.11 "070_ """D'70"2~ 0„10 0.10 OoOl 0.01

2.94 ""0753" 4. 96 0.33

POTASSIUM 1.45 1.04 0. 33 0.3 3 0.08 0.03 3.82 POTASSIUM - 0.45 0.59 0.47 0.27 Oo 07 0.08 1.93

BRANCH BRANCH *- -TWIGS->1 1-1/4 (TOP) (MID) (LOW I TOTAL

-FOLIAGE-(TOP) (MID) (LOW)

CONES TOTA'L

THIGS*FOLI AGE—

(TOP) """ (MID) u o w r TOTAL"

TR EE TOTAL"

NITROGEN - 0. 16 2.12 0. 88 1.62 0. 93 3.43 0.61 1.49 0. 74 2.83 0.00 1.49 3.11 1.66 "07*59 0.2 9 2.16 1.04 0.42 0.21 1.23 0.64

6.26 1.10 4. 00 0.80 2. 43

14.34 PHOSPHORUS-CALCIUM _MAGNES I_UM„; POTASS"!UM -

0.03 0.42 0. 24 3. 08 0.02 0. 36 6. 10 1. 38

0. 12 C. 46 0.J38 0.29"

0.30 0. 13 1.16 0.49 0. IB 0.09 "0.62 "0.33"

0.55 0.10 2.11 0.35 0.36 0. 08 1.25 0.2 7"

0.29 0.16 1.01 0.54 0.24 0^12 0.61 0.30

0. 55 0. 00 1.89 0.00 0.44 0.00

T . 18" "0700"

0.22 0.80 0.17 '0.56*

2.66 15.32 2.03

"9.6'5

1 3 5 7 9 10 1 3 5 7 9 10

NITROGEN -PHOSPHURUS-

0.040 0. 015

0.045 0.007

0.035 0.006

0.057 0.020

0.072 0.018

0. 066 0.014


0. 171 0. 027

0.130 0.023

0.151 0. 025

0.206 0.027

0.288 0. 048

0. 357 0. 062

CALCIU* MAGNESIUM -POTASSIUM -

0.041 0. 010 0. 08 i

0. 053 0.013 0.066

0. 076 0.019 0.069

0. 064 0. 016 0. 074

0.050 0.012 0.081

0. 040 0. O i l 0. 086


0.603 0. 019 0, 096

0.624 0.020 0.124

0.482 0.019 0.085

07496 0.019 0.109

0.270 0.037 0. 201

07242" 0.045 0.22e

BRANCH 3RANCH -TWIGS-- * -FOLIAGE- CONES * T WI GS.FOL I AGE * >1 1-1/4 (TOP) (MIO) ( LOW) (TOP) (MID) (LOW) ( TOP) (MID) ( LOW)

NITROGEN - 0. 114 0. 153 0.484 0. 363 0.323 0. 776 0. 766 0.759 0.000 •6^30 0. 564 o.>w PHOSPHIIRUS-CALCIUM MAGNESIUM -

0. 01 7 0. 20 3 0.019

0.024 0. 135 0.022

0.067 0.265 0. 066

0. 052 0. 259 0. 053

0.055 0.227 0.044

0. 100 0. 486 0. 105

0. 114 0. 538 0. 117

0. 155 0.478 0. 133

0. 000 0.000 0. 000

0. 4 2>«^ 0. 376 0.08>-'

vO.409,

•"0.085

^-Tj.407 0.352

^0^088 POTASSIUM - 0. 034 0.093 0.246 0. 162 0.149 0.372 0. 309 0. 302 0.000 0. 236 07?>6»

TREE NO. 54

SPECIES NO. 1

LENGTH (M)

22.4

DBH (M )

0.394

CROWN 01 A( M) 4.7

CROWN L.IMI 14. B

BUTT L. (M) 7.6

• ••PERCENT NUTRIENT CONTENT BY SAMPLE SPECIF I CAT I ON ****************************************************************•**»»» -WOOD**SECTION * -BARK*•SECT ION-

4

• ••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPECIFICATI ON **************************** ********»*~**»~*~*~*^

-WOOD**SECTION * 162 3 64 5 66 768 9 10 TOTAL 162 364 566 768 9 10 TOTAL

NITROGEN - 6.34 5.13 4. 31 2.93 0.22 0.06 19.48 NITROGEN - 4.71 3.67 4.11 2.27 0. 72 0. 14 15. 61 PHOSPHORUS-CALCIUM MAGNESIUM -

1.77 7 .23 1.75

0.84 8.44 2.02

1. 33 7.29 1. 80

0.86 2.59 0.64

0.05 0.15 0. 04

0.01 0.04 0.01

4. 86 25. 79 6.26


0.77 19. 20 0.60

" 0763" 14.43 0.50

0.60 0.34 11.25 3.53 0.43 0.26

0 .T2 0 ,67 0, 09

"0 .02 0. 10 0. 02

2. 49 49. 18 1 .91

POTASSIUM - 11.47 8.53 7.46 3.52 0.24 0.08 31.29 POTASSIUM - 3.44 2.73 2.24 1.43 0. 50 0 .09 10.43

BRANCH BRANCH *• •—TWIGS- -FOLIAGE * CONES *- TWIGS+FOLIAGE— TREE

NITROGEN

>1

4. 05

1-1/4 (TOPI (MID) (LOW) TOTAL (TUP) (MID)(LOW I " TOTAL" "(TOP) (MID) "" (LOW) "TOTAL"

8.24 1. 02 2.03 1.13 4.18 PHOSPHORUS-CALCIUM _MAGNES IUM _-POT ASSIUM -

4.81 11.64 5. 39 21. 83 0. 62 1.23 7. 35 9.66 0_. 66 1. 14 1. 99 5.14

0. 14 0. 56 0. 14 0. 52

0.00 5.83 13.67 6. 52 0.29 0. 19 1.45 0.79 0.30 0.15 0.91 0. 52

0.62 0.62 2.80 3. 01 0.59 0.65 1.95 " 2.30

1.73 8. 17 1.7 8

' 4.69'

1.10 3. 39 0.94 2.15

3.45 14.58 3.37 "9.14

0.00 0. 00 0.00 0.00

0.76 3.57 0.79 2.82

2. 03 9.62 2.07 5.60

T 7 2 9 " 4.1 9

26. 01 ros""

TOTAL"

73.41 4.

17.38 1.10 3.96 2.67 11.09

13.2T" 109.36

13.93 60.93

TREE SPECIES LENGTH OBH CROWN CROWN BUTT NO. NO. (Ml CM) DIA(M) L. (M) L.(M) 60 2 20.4 0.5 33 0.0. 14.3 6.1

• ••PERCENT NUTRIENT CONTENT *

: 1 3

BY SAMPLE SPECIFICATION******************************************************************************* — WOOD**SECT ION * * BARK**SECTI ON *

5 7 9 1 0 1 3 5 7 9 1 0


0. 057 _0_._003_ 0.094 0. 005 0.056

0.081 0. 009 0.095 0.005 0.050

0.048 0.004 0. 129 0.007 0.050

0.072 _C_006_ 6. 1 24 0.003 0. 047

0.051 0.005 " 0. 119 0.009 0. 044

0 . 053 g. oo 5_

"o. n~8 0. 009 0. 04 4


0.253 0.028_ "~0.'546 0. 018 0.142

0.229 _0.022_ 0.592 ' 0.017 0.088

0. 203 0.019_ 0,590 0. 018 0.07e

0.063 0.004 "0 .11 3 0.007 0.043

0.063 0. 081 0. 004 0.005 0.116 6.099" 0.009 0.007 0.059 0.040

BRANCH BRANCH >1 1 - 1 / 4 (TOPI

rJwiGS-rr *

CMIDT (LC'KI

^FOL IAGE-(M ID I (LOW I

CONES * TWIGS+FOLIAGE * "(TOP) ""'" (MID) (LUW)

NITROGEN 0. OOP . 0. OOP 0.000 o.oqo o.ooo o.ooo 0.000 0.000 0. 000 0.000 0. 000 0.000 PHOSPHORUS-CALCIUM MAGNESIUM -POTASSIUM -

0. 003 0. 003 0. 033 0. 003

0.000 0.003 0.003 0.000

0. 000 O . O P O 0.000 6.000 0. 003

0.000 0.000 0.000

0. 00 0 0. 000 0. 00 0

0. 000 0. 000 0. 000

0.000 0.000 0.000

0.000 0. 000 0.000 0.000

0.000 0. 000 0.000 0.000

0.000 0.000 0.000 0.000 0.000

0.000 0.000 0.000 0.000 0. 000 0. 000^

0.000"

• ••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPEC I F I CAT ION**************************** ************"************"**************** ******

« . . . WOOD**SECT ION * * BARK**SECTI ON * 162 364 566 768 9 10 TOTAL 162 364 566 768 9 10 TOTAL

NITROGEN - 14.51 11.41 8.67 3.81 0.70 0.25 39. 36 NITROGEN - 6.61 3.91 1.26 0.00 0.00 0.00 11.78 PHOSPHORUS- 1.65 1.04 0.73 0.3 7 0.08 0.03 3.89 PHOSPHURUS- 0. 68 0.37 0. 11 0.00 oroo- o."co- T.16 CALCIUM 19.93 19. 80 18.23 7. 50 1.63 0.57 67. 63 CALCIUM 15.59 10.70 3.34 0.00 0. 00 coo 29.63 MAGNESIUM - 1.13 1 .09 1.10 0.52 0.12 0.04 3. 99 MAGNESIUM - 0.48 0.32 0.12 0.00 0,00 0.00 0.92 POTASSIUM - 11.21 8.37 7.03 2.84 0.61 0.21 30.77 POTASSIUM - 3. 16 1.51 0. 58 0.00 0. 00 0. 00 5.25

BRANCH BRANCH *- TWIGS * * — FOLIAGE * CONES * TWIGS + FOL IAGE — TF EE >1 1-1/4 {TOPI ( MIDI (LOW I TOTAL (TOP I (MIDI ( LOW) TOTAL (TOP I" (MIO) "(LOW) TOTAL "TOTAL "

NITROGEN - 0.00 0.00 0.00 0.00 0. 00 0.00 0. 00 0. 00 0. 00 0. 00 0.00 0.00 0.00 0.00 0.00 51. 14 PHOSPHORUS- 0. 00 0. 00 O.PO 0.00 0. 00 . 0.00 0.00 0.00 0.00 0. 00 0. 00 0.00 0.00 6.00 0.00 5.03 CALCIUM 0 . 03 0. 00 0. 00 0. 00 0. 00 ' 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0 . 00 97.25 MAGNESIUM - 0. 00 0. 00 0.00 0.00 0. 00 0.00 0.00 0. 00 0.00 0. 00 0.00 0.00 0.00 0.00 0.03 . 4.91 POTASSIUM - 0. 30 0 . 00 o. 66" 0.00" 0. 00 " " 0.00 6.00" 6.66 0. 00 "6. 00 " "o; oo" 0.00 0. 00 0.00 0. 00 3 6:. 02

TREE NO. 6)

SPECIES NO. 3

LENGTH ( M )

27.2

DBH < M I

0.401

CROWN OIA(M) 8.3

CROWN BUTT L.(H) L.(H)

15.4 11.8

•••PERCENT NUTRIENT CONTENT BY SAMPLE SPECIFICATION************************************************ • WOOD**SECT ION * * BARK**S ECT I ON *

1 3 5 7 9 10 1 3 5 7 9 10


0.053 0. 009

0.049 0.005

0. 126 0.123 0.013 0.012 0.04 9 0.046

0.06 0 0.005

0. 069 0. 009

0.073 0.007

0. 072 0. 012

0. 119 0.014 0. 046

0. 053 0.006 0. 027

0.111 0.135 0.017 0.014 0.068 0.056


0.255 0. 045

0.250 0.032

3.509 2.182 0.041 0.039 0.168 0.113

0.268 0, 043 2. 326 0.050 0.212

0.339 0. 064_ T788 2" 0.055 0.241

0. 365 0^070 1.477 0.069 0.220

0.429 _(K0 31_ 1.490 0.080 0.279

BRANCH BRANCH -TWIGS- -FOLIAGE->1 1-1/4 (TOP ) ( MIU) (LOW) (TOP) (MIO) (LOW)

CONES J * r — T J i ! GS+FOL L A G i - : - * (TOP] ~ (MTOT ( LOW)"

_NJTRGGEN ^ _ PHOSPHORUS-CALCIUM MAGNES IUM_Z_ POTASSIUM" -

0. 107 0. 013 0. 523 0. 029 0.062

0.250 0.000 0.000 0.000 0. 000 0. 000 0.000 0.000 0.846 0.703 0.841 0.031 0. 305 _9_'_°43_ 0. 154

0.000 0. 00 0 0.000

0.000 0.000 0. 000

0.000 0.000

0.000 0.000 _q._o_oo_ "o.ooo

0.000 0. 000 0. 000 0.000

0. 000 0. 000 0. 000

0.000 0.000 0.000

0.000 0.000

0. 000 0.000 0.000 0.000"

0. 094 1.222 0.092

0. 091 1. 313 0.091

0.476 0. 354

0.091 1.447 0.094 0.476

• .•NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPECIFICATION*****************************************"

-WPOU** SECTION- -BARK**SECTION-162 364 566 768 9 10 TOTAL 162 364 566 768 9 10 TOTAL

NITROGEN - 10.63 7.95 7.45 3.98 0.68 0.19 30.84 NITROGEN - 2.45 3.32 2.83 2.96 0. 51 0. 34 12.40 PHOSPHORUS-CALCIUM MAGNESIUM -

1.33 26.01 2.64

0.67 17.66 1.94

0. 31 9.94 1.16

0.45 4.59 0.64

0.06 1.03 0. 16

0.03 0.35 0.04

3.41 59. 53 6.57


0.37 2 7. 60 0.39

0.48 28.85 0.57.

0.49 19.57 0.49

0.56 14.11 0.52

""O.T'O"" 2.07 0. 10

'• 0.06 ' 1.19 0.06

'2 .07 93.39 2.13

POTASSIUM - 9.91 6.73 4.25 2.67 0.63 0.15 24. 33 POTASSIUM - 1.36 2.08 2.11 1.94 0.31 0.22 8.01

BRANCH BRANCH * - TW I GS- •* - F O L I AGE CONES *- TWIGS*:FOLIAGE « TREE

NITROGEN

>1

5. 21

1-1/4 (TOP) (MIO) (LOW) TOTAL

5. 93 PHOSPHORUS- 0.87 CALCIUM - 25.31 MAGNESIUM -__ 1.40

' POTASS IUM - ' "3.05'

0. 73 19. 09 1. 02

"3.65

0. 00 0. 00 0.00 0. 00 0. 00 "

0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0. 00 "" 0. 00

( TOP) (MID) (LOW) " TOTAL "(TOP)' (MID.) '" (LOW)

0.00 0.00 0.00 0.00 0. 00 0.00 4.57 12. 70 0.00 0. 00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0. 00 0.00 0.00

0.00 0.00 0.00 0.00 0.00 0.00 0.00 '" 0.00

0.51 6.60 0. 50 2.57

15.67 2.04 nre~

29.28 21.85 2.03 1.42 7.90 7.19

TOTAL

32.94 s~7T2"

57.73 3.95

17.66

TOTAL

87.33 11.00"

255.59 15.OS 56.70

9 L

TREE NO.

65

SPECIES NO. 1

LENGTH ( M)

23. 5

DBH ( M)

0. 343

CROWN DIA(M) 6.1

CROWN BUTT L.IM) L.(M) 18.5 4.9

•••PERCENT NUTRIENT CONTENT BY SAMPLE SPECIFICATION**************************************************** • WOOD**SECT ION * * BAPK**SECTI ON *

1 3 5 7 9 10 1 3 5 7 9 10


• . 045 J2- 01 2_ 0.051 0.00? 0. 06 0

0.042 JJ.-P14_ 0.055 0. Oil 0.035

0.045 _0. 011_ 0.04 7 0.010 0.054

0.047 0.008 0.041 0. 008 0. 061

0.052 0.014 0.045 0.009 0. 069

0. 067 _0. 01__ 0. 043 0. 009 0. 079

NITROGEN -_P>HOSPHORUS _ C AlC IUM '"-MAGNESIUM -POTASSIUM -

0.151 0.158 0.036 0.020 0.3 f6" 0.423"" 0.013 0.011 0.062 0.069

0. 167 0.027_ 0. 464 0.015 0. 108

0.201 0.032 0.284 0.017 0.137

0. 275 0.311 0. 041 0. 044_ 0.383 "0.3 63 0.024 0.026 0.154 0.180

_BRA'CH BRANCH_ 1-1/4" >1 (TOP)

-TWIGSr ( MID) ( LOW)

* — --FOL IAGE-(TOPf (MID) (LOW )

CONES * TWIGS+FOL I AGE * "(TOP) (MID) " (LOW)"

NITROGEN - 0. 091 0. 161 0.479 0.423 0.423 0. 856 0.814 0.814 0.000 PHOSPHnRUS-CALCIUM MAGNES IUM -POTASSIUM -

0. 014 0. 173' 0. 013 0. 059

0.025 0.249 0. 020 0.094

0. 06 5 0.42 3 0.04 6

0. 054 0.419 0.042

0.054 0.419 0.042

0. 106 0. 472 0. 085

0. 124 0. 510 0. 107

0.220 0.219 0.219 0. 467 0.43 2

0.124 0. 510 0. 107 0.432

0.000 0.46 0.000 0.447 _0.000 0.06 0.000"

***NUTPIE~NT AMOUNTS IN DECAGRAMS. BY SAMPLE SPECIFICAT I ON****************************A*********************************************

* WOOD**SECTION * * BARK**S EC Tl ON —

.NITROGEN PHOSPHORUS-CALCIUM MAGNESIUM -

1C2

5.99 1.77 7.3) 1.4J

POTASSIUM - 10.01

3C4

4.52 1.32 5. 35 1.08 7.24

5 66

2.72

7t8

1.30 0.28

10

0.12

TOTAL

14. 93 NITROGEN

1C2

4.94

3E4

3.64

5£6

2.30 0.58 2.63 0.52 3.43

0.2 9 1.12 0.22

0.07 0.24 0.05

0.03 0.08 0.02

4.07 16.73 3.28


0. 89 11.80 0.38

0.53 9.94 0.28

0.3 7 4.67 0.20

7£8 9

1.55 0^33_ 0.24 0. 05 2.17 0.46 0.14 0.03

10

0. 12 "'0.0'2 0. 15 0.01

"TOTAL

12. 89

ro"9~ 29. 18 1.04

1.72 0.37 0.14 22.91 POTASSIUM - 2.09 1.98 1. 53 0.94 0. 13 0.07 6.60

BRANCH BRANCH *->1 1-1/4 (TOP)

NITROGEN - 7.32 11.85 2.06

•— TW IGS

"(MT6) (LOwf

.* * FOLIAGE-TOTAL (TOP) (MID) (LOW) TOTAL

3. 09 2.41

CONES * TW I GS+ FOL I AGE— ""( TOP)"" ( MID) "" ( LOW)

7.56 6.33 10. 34 8.79 25.46 0.00 PHOSPHURUS- 1.10 CALCIUM - 14.34 MAGNESIUM- 1.09 POTASSIUM - 4.77

1. 84 18. 2 5 1. 43 6. 3 9

0. 23 1. 82 0. 20 "0. 95 "

0.39 0.31 3.06 2.39 0.31 0.24_ I'.bO 1.25

0.98 0.79 7.26 3.49 0.74 0.63 3.79 3.46

1.58 6.48 1^36 5.49

1. 34 5.51

JL. 15 4.67

3. 70 15.48 3.14

T3." 6 r

0. 00 0.00 0.00 "0. 00

8.39 13.43 1.06 5.31 0.83 "4". 40"

11.20 T797 1.65 9.54 7.90 1.66 1.39 "7.09" 5*. 91"'

"TOTAL"

33. 02 4.68 22.74 3.88

17.40"

TREE TOTAL"

80.02 13.79

101.25 10.76

•"58.78 "

TREE NO. 69

SPECIES NO. 1

LENGTH t M )

18.9

OBH (Ml

0.221

CROHN OIA(M) 3.5

CROWN BUTT L.(MI L.(M) 14.0 4.9

• ••PERCENT NUTRIENT CONTENT BY SAMPLE SPECIF I CAT ION*********************************************************************»***»*•»** -WOOD**SECTION- --BARK**SECTI ON-

1 3 5 7 9 10

NITROGEN - 0. 042 J . 040 0.047 0. 045 0. 060 0. 06 9 PHOSPHORyJ- 0. 01J 0. 013 0.012 0. 013 0. 014 0. 014 CALCIUM 0. 03 9 0. 049 6. 042 0. 042 0. 038 0. "038 MAGNESIUM - 0. 00 3 0. 013 0.012 0. 012 0. 010 0. O i l POTASSIUM - 0. 06 0 0. 054 0. 055 0. 070 0. 074 0. 073

10

NITROGEN -J>_HOSPHORUS-CALCIUM MAGNESIUM -POTASSIUM -

0.150 0. 0 2 4_ 0.2 33 " 0. 019 0. 100

0.185 _0._028_ " 0.222 0.028 0.158

0.201 0. 0 3 3_ "6". 2 28 0.028 0. 179

0.245 0.042_ 0.191' 0.037 0.201

0.288 0.322 0.048_ _0.053 "0.204 "" 0.217 0.038 0.040 0.217 0.212

BRANCH >1

_3R_ANCH_ 1-1/4

-TWIGS- -FOL I AGE-( TOP ) ( M I D I I LOW) (TOP ) (MID) (LOW)

CONES •r-rTWIGS*FO_L I AGE — (TO PI (MIDI ( LOW)

NITROGEN 0. 105 0. 159 0.472 0. 564 0.345 0. 793 0. 809 0.790 0.000 PHOSPHORUS-CALCIUM _MAGNES IUM_r POTASSIUM -

0.02 0. 0. 109 _0.022_

,074

0. 024 0. 143 _°«027_ 0.100

0.074 0.199 _0. 0_58_ 0.183

0. 072 0. 158 0. 047

0.046 0. 346 0.035

0.169 0.169

0. 122 0. 331 0. 106

0. 134 0.412 0. 128

0.336 0.315

0. 162 0.509 0.108 0.408

•••NUTRIENT AMOUNTS IN OECAGRAMS BY SAMPLE SPEC IF ICAT I ON**************************** **«*'*** + *****"*T'*l>*"********«îrV*>*"i^*

; « WOO0**SECTI0N * * BAR K**SECT ION * 1C2 3C4 . 5&6 7C8 9 10 TOTAL 1&2 3£4 5C6 7E8 9 10 TOTAL

_NITROGEN _ - 1.72 1_.38 1.08 0.72 0.16 0.06 5.11 NITROGEN- 1.91 1.49 1.27 0.60 0. 20 0.10 5.77 PHOSPHORUS- 0.47 0.38 O.T§ 0.18 07o4 0.01 1.37 PHOSPHORUS- 0.30 0.24 "0".2'1" 0.13" 0.03 """0.02 0.93 CALCIUM - 1.86 1.44 0.99 0.55 0.10 0.03 4.97 CALCIUM - 2. 59 1.73 1.20 0.59 0.14 0.07 6.32 MAGNESIUM - 0.45 0. 39 0.27 0.15 0.02 0.01 1.29 MAGNESIUM - 0.27 0.21 0.18 0.11 0.03 0.01 0. 81 POTASSIUM - 2.33 1.72 1.47 0.99 0.19 0.06 6. 81 POTASSIUM - 1747 1730 1708 b"7"""3~ 0TT5 0"70"6 4"77b"

BRANCH BRANCH * TWIGS

>1 1-1/4 (TOO) (MIDI

* * ( LOW I TOTAL ( TOP)

— FOLIAGE--—-(MIO)"" fLOW) TOTAL

JZONES *- TWI GS+ FOL I AGE — (TOP I (MIDI (LOW I' TOTAL"

TREE TOTAL"


0. 13 3. 93 0.99 1 .35 0. 24 2.59 4.92 4.69 0.95 10.56 0. 00 5.91 6. 04 0. 02 0.60 0. 13 3.59 0. 03 0. 66 '"o".""09"" 2.50'

0. 16 0,42 0, 12 0. 38

0.17 0.03 0.38 0. 24 0.11 0.02 0.41 " C. 12

0.36 0.75 1.04 2. 05 0.26 0.66 0.91" " 2. 08

0.78 0.19 2. 39 0.61 0.74 0.13

T.82" 0.49"

1.73 5.05 1.53 4.40"

0.00 0.00 0. 00 "0.00

0.91 2.47 0.78 2.47

0.95 2.77 0.85 2.23

1.19 0723"" 0. 85 0. 15 0.61

13.14

6.09 1. 79 5. 31

5.01 21.13 4. 58

19. 40

TREE NO. 72

SPECIES NO. 1

LEN GTH ( Ml

24.2

D8H (HI

0. 396

CROWN 01A(Ml 6.6

CROWN BUTT L.(M) L.(H) 17.3 6.9

•••PERCENT NUTRIENT CONTENT BY SAMPLE SPECIFICATION******************************************************************************* * W0O0**SECT ION * * BARK**SECTI ON * _1 _ J 5 7 9 10 1 3 5 _7 9 10

NITROGEN -PHOSPHCRUS-

0. 046 0.._030

0.043 . 0. 0 2_2_


0.053 0. 014 0. 104

0.057 0. 014 0.153

0. 042 _0.-pl2 0. 042 0.010 0. 03 4

0.042 _0. 006_ 0.048 0. 009 0. 064

0.044 o- o n_ 0.047 0.009 0.073

0. 060 j0.j011_ 0. 038 0. 009 0. 05 5


0. 159 0.026_ "67332" 0. 015 0. 062

0.118 _0._0_19_ 0.327 ' 0.013 0.083

0. 141 0.023 0.301" 0. 014 0. 109

0. 160 0.031 "6.2 21" 0.015 0.114

0.211 0.302 0. 040 0.045 "0.256"" " 0.255' 0. 019 0. 029 0.165 0.221

BRA NCH BRAN C H_ 1-1/4

NITROGEN

>1

0. C77 0. 132

_*r-_- ~~I WIGS- -(TO PI (MI Of" ( LOW I

0.718 0.484 0.444

(TOP)

0. 821

-FOLIAGE-" "(MID)

0. 824

(LOW)

0.820

CONES

0. 000

* TWIGS*FOL I AGE * "(TOP)" (MID) " (LOW)

PHOSPHORUS-CALCIUM MAGNESIUM

0. 012 0. 134 0. 014

0.025 0.173 0. 020

0. 075 0.242 0.04 5

0. 068 0.25i 0. 049

0.073 0.246 0.053

0. 112 0. 317 0.072

0.121 0.438 0. 037

0.144 0.430 0.082

0.000 0. 000 0. 000

POTASSIUM - 0.030 0.033 0.131 0.168 0.220 0.431 0.297 0.311 0.000

•••NUTRIENT AMOUNTS IN DECAGRAMS 3Y SAMPLE SPECIFICATION**************************************************************************

* WC;OD**SECTION * * BARK*»SECT I ON

NITROGEN -

1C2

7.70

3£4

4.94

5C6

3.35

7£8

1.52 0.28

10

0.08

TOTAL

17. 86 NITROGEN

U 2

7.23

3C4

4.22

5£6


4.45 9.45 2.41

1.96 5.76 1.42

0.71 3.65 0.79

0.30 1.68 0.32

0.07 0.2 9 0.06

0.01 0.05 0.01

7.50 20. 88 5. 01


1 . 1 6

17. 18 0.73

0.69 10.21 0.43

0.5 8 5.60 0.31

7£8 9

2.08 0.38_ 0.40 0. 07 2.67 0.46 0.19 0.03

T P " "TOTAT

POTASSIUM - 22.04 13. 73 5,96 2.42 0.46 0.07 44.67 POTASSIUM 3. 80 3.12 2.39 1.56 0, 30

BRANCH BRANCH *- -TWIGS-

NITROGEN -

>1

4. 31

1-1/4 (TOP) (MIDI (LOWI TOTAL ( TOP) ^FOLIAGE---— ( MIO')" "I LOW I

9.81 4. 09 5.38 3. 19 12.66 7.23 17. 55 11. 73

TOTAL

36. 51

CCNES *- TWIGS•FOLI AGE--(TOP)"~"(MID) (LOW)

0.00 11.32 1.42 4.17 0.89 "4.82"

14.92 2.56

22.93 ~T73"4" 12.11 7.92 2.39 1.56

" "8. 19 6.03

' TOTAL"

49. 17

TREE "TOTAL"

98.45 —irvr 24. 20 4. 84

"19.04 "

20.30 101.62 13.65

~ 8 5.41"

PHOSPHORUS-CALCIUM _MAGNES IUM__-POTASSTUM -

0. 69 7. 49 0.81 4. 49

1.35 12. 81 _1._43_

5. 93

0, 43 1. 38 0. 26 1. 03"

0.75 2. 78 0.54 1 .86

0. 52 1. 77 0. 38 1. 58"

1.71 5.93 1.18 4.4 8

0.99 2.79 0. 64-_ '3.79 "

2.58 9.33 1. 85_ "6.3 3

2.06 6.14 1.18 4.44

"T76"r 18.27 3.66

TV. 5 6"

"1570 0 0 . 0 0 0 . 0 0 0700

TREE SPECIES LENGTH DBH CROWN NO. NO. ( M) (HI DIA(M) 76 3 14.8 0.180 5.1

CROWN BUTT L.(M) L.IM)

9.1 5.7

•••PERCENT NUTRIENT CONTENT BY SAMPLE SPECIFICATION*************************************************** • WOOD** SECT I ON * * BARK**SECT ION *

1 3 5 7 9 10 1 3 5 7 9 10

NITROGEN -__PH OS PHORUS-CALCIUM "AGNESIUM -POTASSIUM -

0.043 _0.0.09_ 0. 153 0.011 0. 045

0.053 J>_.JH>9_ 0. 149 0.008 0.050

0.056 0.036 0. 121 0.009 0.044

0. 057 0.006

0.071 0.009

0. 064 0. 011


0.261 0. 043

0. 121 0.008 0. 053

0.138 0.008 0.054

0. 140 0. 007 0.054


2.934 0. 026 0. 162

0.289 0. 048 2.995 0.041 0.173

0.226 0. 047' 2.839 0.032 0. 159

0.276 0.075 2.A43 0.036 0.223

0.348 0.071

0.313 _0.079 2/537 1/9 59 0.047 0.044 0. 329 0.306

BRANCH BRANCH -TWIGS-- * FOLIAGE- CONES >1 1-1/4 (TOP) (MID) ( LOW) (TOP) (MID) (LOW)

* TW!GS*FOLIAG_E---*_ (LOW)" ( TOP) (MID)

NITROGEN 0.113 0.190 0.000 0.000 0.000 0. 000 0.000 0.000 0.000 0.766 0.675 0.766 PHOSPHORUS-CALCIUM VAGNEJJUM^-" POTASSIUM -

0. 316 0. 573 _0.013_ 0. 363

0.025 3. 882 0.023_ 0. 14 7

0.030 0. 30 0 0.000 0.003

0. 000 0.000 0. 000

0.000 0.000 0.000

0.003 0.000

0. 000 0. 000 0. 000 0.000 0.000

0.000 0.000 0.000 0.000 o oip o o. o oq_

o.'boo

0.000 0.091 0.082 0.088 0.000 1.180 1. 1 18 1.346 _0.000 0.059 0.05 8 0.060_ 0.000 "b"/"296" 0/310 0. 338

• ••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPECIF 1

-WOOD*«SECTION- -BARK**SECTION-


1 £2 3£4 5 £6 7£8 9 1 0 TOTAL 1£2 3£4 5 £6 7£8 9 10 TOTAL

1.53 1. 04 0. 74 0.39 0.06 0.03 3.79 NITROGEN - 0.47 0.33 0.30 0.3 5 0, 09 0.00 1. 55 0.25 4.42 0.26

0.14 2.44 0.15

0.03 1.57 0.11

0.05 0.79 0.05

0.01 0.12 0.01

0.01 0.07 0.00

0. 55 9.42 0.59


0.08 5. 04 0.06

0.06 3.79 0.05

0.07 3.41 0.04

"0/09 3.2 3 0.05

0. 02 0. 76 0. 01

o.'o'o 0.00 0. 00

0. 33 16.23 0.21

1.34 0.85 0.63 0.32 0.05 0.03 3.25 POTASSIUM - 0.29 0.22 0.23 0.33 0.10 0.00 1.16 . POTASSIUM

8RANCH BRANCH *- -TWIGS- -FOL I AGE- CONE S *- -TWIGS* FOLI AGE- TREE >1 1-1/4 (TOP ) "(MID) ( LOW) TOTAL (TOP) (MIDI (LOW) TOTAL (TOP) (MID) "(LOW) TOTAL TOTAL

NITROGEN - 0. 55 2. 11 0. 00 0.00 0.00 0.00 0.00 0.00 0.00 0. 00 0. 00 1.84 3. 51 7. 58 12.93 20.93 PHOSPHORUS- 0. 03 0.29 0. 00 0. 00 0. 00 0.00 0. 00 0. 00 0.00 0.00 0.00 0.22 0.43 0. 87 1.52 2.76 CALCIUM 2.63 9. 79 0.03 0. 00 0. 00 0.00 0. 00 0. 00 0.00 0.00 0.00 2.83 5.81 13.32 21.97 60.09 MAGNESIUM - 0. 06 0. 2 5 0. 00 0.00 0. 00 0.00 0.00 0.00 0.00 0.00 0.00 0.14 0. 30 0. 60 1.04 2. 15 POTASSIUM - 0.32 1.63 0. 00 0.00 " "b.'oo 0.00' "' o. bo " "0. 00 0.00" "0.00" "0.00 0.71 1.61 3. 34 5.67 12.03

TREE NO. 102

SPECIES NO. 3

LENGTH ( Ml

25. 5

DBH (Ml

0.602

CROWN OIAI Ml 9.1

CROWN L.IMI 15.0

BUTT L.IMI 10.5

• ••PERCENT NUTRIENT CONTENT BY SAMPLE SPECIF ICATI0N***************^***^*******^**+****^***^***********+*************^***»»****«**» • WOOD*«SECT ION * * BARK»*SECTI ON *

1 _ 2 5 7 9 10 1 3 5 7 9 L9

NITROGEN -JLHILSPjHOJi!JJi£_ CALCIUM MAGNESIUM -POTASSIUM -

0. 052 J)._C0 5_ 0. 105 0.011 0. 0*4

0.054 _0.op3_ 0.095 0.009 0.046

0.051 0.005 0.090 0.01 0 0.04 7

0.067 _0.0 ? a

o.io'o" 0.010 0. 049

0.064 0.006 0.100 0.011 0.053

0. 085 0. 009 o".T6i 0. 01 0 0. 056

NITROGEN -_PJjOS_PHORUS- _ CALCIUM "'-MAGNESIUM -POTASSIUM -

0. 300 0.05_5_ "2.951 0. 039 0.173

0.298 JL'J?73

2.160" 0.045 0. 166

0. 322 0. 086 "2.543 * 0. 050 0.255

0. 365 0.094 2.015" 0. 044 0.300

0. 417 0. 110 1.989"' 0, 064 0.296

0.516 0. 1 04 1.772 0. 062 0. 326

BRANCH BRANCH >1 1-1/4 (TOPI

-TWIGS-( MIDI ( LOWI

*-- — r f OL I AGE-IT OP) (MIDI (LOW I

CONES * TWI GS + FOL I AGE * (TOP) ' (MID) " (LL'W I

NITROGEN - 0. 155 0. 259 0.000 0.000 0.000 0.000 0.000 0. OOP 0. OOP 0. 8P7 0.776 P. 665 PHOSPHORUS-CALCIUM MAGNESIUM -

0.023 0.037 0.719 0.735 0. 024 p. 032

0. 000 O.POO 0.000

0. 000 0. 000 0.000

O.OPP 0.000 O.OOP

0. 000 0. 000 0.000

O.PPO 0.000 0. 000 0.000 O.OOP 0.000

O.OOP 0.083 0.082 0.091 0.000 1.432 1.697 1.905 0. 000 0.084 0.088 0.114

POTASSIUM - 0. 036 0.138 0.000 0.000 0.000 0. 000 0.000 0.000 0.000 0.394 0.385 0.330

• ••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE S PEC IF ICAT I ON*********************************************"* ***»**«***•«*«*« »*»r***4*"*~

* WOOD * SECT ION- * * 8ARK««SECTI0N * 1£2 3(14 5 £6 7E8 10 TOTAL 1£2 3(14 5£6 7E8 10 TOTAL

NITROGEN - 21.33 13.22 10.34 5.49 0.73, 0.23 51. 84 NITROGEN 7.77 PHOSPHORUS-CALCIUM MAGNESIUM -

2. 32 41.21 4.21

1.65 23. 33 2.45

1.12 16.70 1.80

0.6 0 8.39 0.88

0.07 1.15 0.12

0.02 0.27 0.03

6.28 91.06 9. 50

PHOS'PHORUS* "" CALCIUM MAGNESIUM -

1. 66 66.44 1. 10

7.96 """2.04 60.44 1.22

5.94 ~"1756" 39.43 0.81

2.66 P.'6"9

12.61 0.37

1.04 "0727" 4. 97 0.16

0.36 25.73 "0707 673*0" 1.24 186.14 0.04 3.70

17.60 POTASSIUM 13.63 11.74 8.39 4.2 3 0.62

BRANCH BRANCH •- -TWI GS->1

0.15 43.81

1-1/4 (TOP) (MID) (LOW) TOTAL

POTASSIUM

-FOLIAGE-

4.40

TTUP) (MID) (LOW) TOTAL

5.40 4.80 2.03 0. 74

CONES *- TWIGS* FOLI AGE— (TOP)~ (MIDI (LOW)

0.23

TOTAL" TP EE

"TOTAL'

NITROGEN 19.61 13.65 0. 00 0.00 0. 00 0.00 0.00 0.00 0. 00 0. 00 0.00 13.48 38.19 37.88 89.55 200.39 PHOSPHORUS-CALCIUM MAGNESIUM -POTASSIUM

2. 95 90. 73 3.00

10. 34

1.95 41. 29 1.71 7. 25

0.00 COO 0.00 0.00"

0.00 0. 00 0.00 O.OP

0. PO P. 00 0. 00 0.00

0.00 0.00 jO.OO^ 0.00

0.00 0.00 0. 00 O.JO

0. 00 0. OP P.PO P.PO 0.00 0.00

" 0 7 0 0 O 7 0 ' 0 ~

0. 00 0. 00 0.00. 0.00 0.00 0.00

" " 0 7 0 0 0"70'0"

1.39 23.91 1.41

""6.57"

4.01 83.51 4.31

"18". 96"

57015 97"41T 83.45 190. 86 4.99 10.70

"14.46 40.00

26.83 600.C9 28. 61

"115.50 "

TREE NO. 103

SPECIES NO. 1

LENGTH (Ml

15.5

DBH (M)

0.183

CROWN OIA(M) 2.9

CROWN BUTT L.IM) L.(M) 10.3 5.3

•••PERCENT NUTRIENT CONTENT BY SAMPLE SPECIFICATION****************************************************** • WOOD**SECTION * * BARK**S ECT ION *

1 3 5 7 9 10 1 3 5 7 9 10

NITROGEN -PHOSPHORUS-. CALCIUM MAGNESIUM -POTASSIUM -

0.03 7 _0_.-P_39_ 0. 047 0.011 0. 056

0.040 _0_._012_ 0. J43 0.010 0.051

0.036 0.008

0.046 0.015

0.056 0.016

0. 063 0. 014


0.200 0. 037

0.217 0.045

0. 047 0.010 0.051

0. 048 0.010 0. 055

0.039 0.011 0.062

0. 04 5 0. 010 0. 062


0. 574 0.029 0. 120

0.552 0.041 0. 192

0.236 _0.06 0_ 6. 500 0. 059 0. 257

0.268 _0.06 3 0. 399 0.059 0.213

0.342 3.075

0.329

o.jno 0. 41 7 674 00" 0.067 0.060 0. 237 0.232

BRANCH BRANCH * TWIGS- -FOL IAGE-

NITROGEN -

>1

0. 113

1-1/4

0. 161

(TOP)

0.454

(MID)

0. 514

( LOW)

0.413

(TOP)

0. 788

(MID)

0. 792

(LOW)

0.779

CONES

0.000

* TWIGS + F0LI AGE-- • (LOW) ( TOP) (MID)


0.013 0.115 0. 025

3.024 0.223 0.031

0.036 0.340 0. 08 7

0.078 0. 350 0. 089

0.065 0.299 0.075

POTASSIUM - 0.079 0.093 0.208 0.221 0.179

0. 124 0. 363 0. 126 0. 341

0.119 0. 49 7 0. 130

0.122 0.537 0.137

0.292 0.246

0.000 0 0.000 0.352 _0.000 0.10 6. oob


•W000**SECT10N- -BARK**SECTION-

JNJJROGEN ^_ FHOSPHORUS-CALCIUM MAGNESIUM -

162

6.27

3E4 5 C 6

0.76 0.53

7C8

0.41 0.09

10

0.03

TOTAL

2.80 NITROGEN

1C2

0. 90

1.16 0.27

0.20 0.90 0.20

0.15 0.52 0. 13

0.13 0.3 5

0.09

0.03 0.07 0.02

0.01 0.02 0.01

0. 79 3.11 0.71


0. 18 2. 42 0.15

3G4

0.61_ 0.14 1.42 0.13

5 £6

0.58 "0.14" 1.03 0.14

7£8 9

0.33 0. 13 0.07 0.03 0.45 ' 0.17 0.07 0.03

10

0.14 "O'VO'S" 0. 16 0. 02

TOTAL

2. 68 0.59"" 5.65 0. 54

POTASSIUM 1.36 1 .01 0.63 0. 4 7 0.11 0.03 3 . 66 POTASSIUM - 0. 67 0.61 0.54 0.25 0. 09 0.09

JSRANCHJ3RANCH * I- 1/4 >1

NITROGEN 0. 92 2. 65

(TOP )

0.68

•--TWIGS --(MID) "(LOW)

1 .08 1. 16

TOTAL

2.92

( TOP)

0.55

•-FOL I AGE "(MID) (LOW)

0.79 1.01

* "TOTAL"

2.36

CONES *- TWIGS+FOLIAGE— (TOP) (MID) (LOW)

0. 00 1.23 1. 87 2.17

TOTAL

5.27 "0.94 3.53 0. 92 2.13

2.25

TREE "TOTAL"

14.33 2T7T" 17.62 2.90

' 10.24

PHOSPHORUS-CALCIUM ^MAGNESIUM _-_ POT AS S1UM -

3.15 0.39 1.64 3.68 3.21 0.51 0.67 " 1.53

0. 13 0. 51 0. 13 0. 31

0.16 0.16 0.73 0. 84 0.19 _ 0.21 0.46 6. 50"

0.47 0.09 2.08 0.25 0.53 0.09 1.28 " 0.24"

0.12 0.16 0.50 0.70 0.13 0.18 0. 29 0. 32

0.37 0.00 1.45 0.00 0.40_ 0.00_ "0.65 " "0.00

0.22 0.77 0.22 0.55

0.26 0.34 1.23 1.54 0. 32 0.39 0.76 " 0.82

TREE NO. 10*

SPECIES NO. 1

LENGTH ( .Ml

17.2

OBH (Ml

0.325

CROWN 01 A(Ml 4.0

CROWN BUTT L.(M) L.(M) 10.4 6.9

•••PERCENT NUTRIENT CONTENT BY SAMPLE SPECIFICATION*************************************** • WOQO**SECT ION * * BARK**SECT 1 ON *

1 3 5 7 9 10 1 3 5 7 9 10

NITROGEN -J_HI3SPM9_RUS:L_

C A L C I U M -MAGNESIUM -P O T A S S I U M -

0. 0*3 J). 03 3_ 0.0 5 7 0. 016 0.03 3

0.039 0.014. 0.076 0.015 0.058

0.037 _0.012 0.028 0.00 6 0. 0*9

0.0*5 _Q.011_ 0.03B 0. 009 0. 055

0.053 .0.011 6.0 38 0.009 0.056

0. 07* .0. 01 6_ 6." 0*3 0.011 0. 073


0.153 0.029 "~0. 5Of" 0.020 0. 081

0.161 _0._032 6.63 9 0.023 0.098

0. 197 0.033 "0.5 25 0. 025 0.101

0.267 0. 050 6.27* 0.036 0.190

0.290 0.313 0.051 0.057 0.260 ' 0.266" 0.035 0.0*2 0.178 0.199

J3RANCH BRANCH 1-1/*

NITROGEN -

>1

0. 117 0.167

I TOPI

0.633

-JWJGS-( MI Dl

0.536

( LOW)

0.533

(TOPI

0. 6*3

-FOLIAGE-" (MID)

0. 6*9

CONES (LOW )

0.632

* TWIGS* FOL I AGE * "(TOP) " (MIO) (LOW)

0. 000 0. 593 PHOSPHORUS-CALCIUM MAGNESIUM -

0. 018 0.22* 0. 021

0.032 0.2 0* 0. 02*

0. 09* 0.239 0. 055

0. 073 0.273 0. 052

0.066 0.251 0.050

0. 125 0. *13 0. 131

0. 130 0. 505 0. 1*1

0. 127 0. 5*3 0. 139

POTASSIUM - 0. 090 0.100 0.221 0.192 0. 158 0. *15 0.371 0.368 0.000

• ••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPEC IF ICAT I ON**************************************************************************

«- WOOD** SECTION * * 8ARK**SECT I ON * 1C2 3t4 5&6 7(18 9 10 TOTAL 1(12 3d* 5&6 768 9 10 TOTAL

NITROGEN - 4.09 3. *9 0.7* 0.*5 0.07 0.0* 8. 87 NITROGEN - 2.16 2.52 1.** 0.53 0.12 0. 13 6.89 PHOSPHORUS-CALCIUM MAGNESIUM -

2.35 6.66 1.55

1.19 *. 76 0.9*

0.21 0.60 0.13

0.10 0.3 5 0.08

0.01 0.05 0.01

0.01 0.02 0.01

3.87 12. *3 2. 72


0.*2 7.87 0.30

0.*5 8.20 0.33

0.26 2.*8 0.19

0. 10' 0.51 0.07

0. 02 0, 10 Oo 01

0.02 0. 11 0.02

1.27 19.27 0.92'

POTASSIUM - 7.35 *. 90 0.9* 0,51 0.07 0.0* 13.81 POTASSIUM - 1. 23 l. * l 0.90 0.35 0. 07 0. 08 * .o*

BRANCH BRANCH *-

NITROGEN

>1

2.07

1-1/* (TOP) --TWIGS (MIDI (LOW)"'

*.99 0.51 0. 97 1 . 4 9

TOTAL

2.96

( TOP I

1. 09

•j^F OL I AGE— (MIDI (LOW I

CONES *-

3. 63 4. 11

TOTAL

8. 83

TW I GS* FOL I AGE * (TOP) (MID) (LOW) TOTAL"'

0.00 1.60 4.60 5.60 11.80

TP EE "TOTAL"

3 4 . 6 2 PHOSPHORUS-CALCIUM MAGNESIUM -POTASSIUM -

0. 32 0.95 3. 96 6. 07 0.37 0.73 1 , 60

0.03 0. 23 0.0*

2. 99 0. 13

0.13 0.19 0. *9 0.70 .0. 09 0. 1* "0.3 5 0.**"

0.39 0.21 1.42 0.70

JO. 28 0.22 0.97 0.71"

0.73 0.83 2.83 3.53 0.7 9 0.J91_ 2.08" "2.39

7.06 1. 92 5.17

0. 00 0.00 0.00 "0.0*0"

0.29 0.93 0.27 ""0.88"'

0. 86 3. 32 0. 89 "2.42

TTor" 4.23 1.05 "2". 83

""2776""" 8.49 2.20 "6. 14"

8.57 50.22 6.93

28.58'

TREE NO. 108

SPECIES NO. 1

LENGTH (Ml

27.7

OBH (M )

0.490

CF OWN DIAIM) 7.3

CROWN L.(M) 23.2

BUTT L.IM) 4.6

• ••PERCENT NUTRIENT CONTENT BY SAMPLE SPEC IF I CAT IQN***********»******************************************************************* « WOOO** SECT ION * * BARK **S ECT ION *

1 3 5 7 9 10 1 3 5 7 9 10

NITROGEN PHOSPHORUS-

0.049 0.012

0.064 0.02 9

0. 050 0.011

0.065 0.016

0. 063 0. 016


0. 146 0. 021

0.155 0.023

CALCIUM MAGNESIUM -POTASS IUM -

,03 3 0.059 0.009 0.012 0.059 0.067

0. C46 0.011 0.092

0. 047 0. 010 0. 069

0.038 0. 037 0.009 0.010 0.071 0.076


0.235 0.255 0.016 0.015 0.071 0.097

0. 164 _0.J>26_ 0. 23 9 0.012 0. 095

0.191 _0_. 02 8 0.2 30" 0.012 0.098

0.255 _0.04 0_ Oc 176 0.016 0. 145

0.348 _0.054_ 0". 216 " 0.019 0.1 94

BRANCH BRANCH -TWI-GS- -FOLI AGE- CONES * TWIGS+FOLI AGE-

NITROGEN -

>1

0. 115

1-1/4

0. 146

(TOP)

0.609

(MID)

0.447

(LOW I

0.363

(TOP)

0. 905

(MIO)

0. 834

(LOW)

0.813

( TOP) (MIO) ( LOW)

0.000 PHOSPHORUS-CALCIUM _MAGNES IUM_-POTASSIUM -

0.02 3 0.155 _0.015_ 0. 071

0.023 0. 149 _0. 01? 0.098

0.035 0.292 _0.J>4 7_ 0. 218

0. 087 0. 3'i3 0. 043

0.055 0.194 0. 036

0.124 0.282 0. 092

0. 127 0.462 0. 101

0. 182 0.402 0. 123

0.201 0.121 0.234 0.343 0. 334

0.000 0.4 0.000 0.287 _0.000 0.06,

61 odd


-WO0D**SECTICN- -BARK**SECTION-1C2 3C4 5(16 7 £8 9 10 TOTAL 1£2 3 £4 5£6 7£8 9 10 TOTAL

NITROGEN - 14.61 9. 55 5. 96 2.27 0.38 0.08 32.85 NITROGEN - 8.92 8.27 5.00 2.67 0.41 0. 14 25.41 PHOSPHORUS- 3.72 3.53 2.12 0.54 0. 10 0.02 10. 03 PHOSPHORUS- 1 . 30 1.26 0.7 5 O.'l ~ "0.06" 6 ;o'2 3.8i CALCIUM 13.66 8.87 4.33 1.69 0.22 0.04 29.37 CALCIUM 14. 54 12.79 6.61 2.43 0.28 0.O9 36.74 MAGNESIUM - 2.91 1.87 1.09 0.3 8 0.05 0.01 6.31 MAGNESIUM - 0.92 0.68 0.33 0.17 0. 03 0. 01 2.13 POTASSIUM - 17.53 13.39 8.43 2.77 0.41 0.09 42.63 POTASSIUM - 4.99 4.97 2.72 1.46 0.23 0.08 14.45

BRANCH BRANCH * - TWIGS * * — F O L I A G E * CONES_ * TWI GS* FOL I A G E - - TP EE >1 1-1/4 (f OPT (MID) (LOW) TOTAL (TOP) (MID) (LOW) "TOTAL ~ (TOP)" (MIO) "' (LOW) TOTAL " "TOTAL

NITROGEN - 14.05 10.48 2. 74 4.47 3.49 10.70 9.96 17.34 13.08 40. 38 0.00 12.70 21. 81 16.57 51.08 133.87 PHOSPHORUS-CALCIUM MAGNESIUM-"POTASSIUM -

2.77 1.68 18.99 10.67 1 . 9 3 1 . 3 6 3. 69 6. 33

0. 38 1.32 0. 21 "C.98'

0.87 0.53 3.13 1.86 0.43 0. 34 2. 0 1 1 . 16

1.78 1.37 2.63 2.93 6.31 3. 10 9.62 6.47 0.99 1.01 2.09 1.98 "4.15 " 2.57 7. 13 " 5.38

6.93 0.00 19.19 0.00 5.07 0.00

15.08" 0.00

1.75 4.42 1.22 3.55

3.50 3.46 12.74 8.34 2. 52 2. 32 9.14 6.55

8.71 25.50 6. 06

19.24

T 7 7 W 121.28 17.74 91. 34

A P P E N D I X 8

PROGRAM FOR C A L C U L A T I N G MOISTURE CONTENT OF

SAMPLES FOR C H E M I C A L A N A L Y S E S

P r o g r a m p r e p a r e d b y M r . K e n T w a i t e s

.COMPILE C PROGRAM FOR CALCULATING MOISTURE CONTENT AND STANDARD DEVIATION OF UP TO C **120*« UNORYfD TREE SAMPLES CLASSED IN UP TO **9*« SPECIFICATIONS AMD C ORYE D AT TWO TEMPERATURES<TI,T2) C PROVIDE DATA CARCS AS FOLLOWS C 1. 4 INTEGERS ON 1ST CARD- NO. OF SAMPLES(M), NO. OF SPEC I F IC AT IONS ( N ) , C 1ST DRYING TEMP.(Til, 2ND DRYING TEMP.(T2) C 2. N CARDS WITH CCMENTS UP TO 12 CHARACTERS FOR EACH SPEC. - DATA IN QUOTES C_ 3. M DATA CARDS FORMAT A3 ,11 , A2 , Al ,4F7.3 , 13 (TREE NO., SPECIFICATION,

"C DISC NO., LOCATICN'WITHIN'CROWN, EM PT Y ROT TLE WE I GHT(A) , BOTTLE•UNDRYE6 C .SAM PL5(BI, ROTTLE+S AMPLE DRYEO AT 1ST TEMP.(C), BOTTLE+SAMPLE DRYED AT C 2ND TEMo. (Ol, BOTTLE NO.

•INTEGER M, N,Tl,T?,SPEC(120l,ROTTLE(120),COUNT!9)/9*0/ RE AL* 9 A(l20),B(120),C(120 I,D(120) ,WB(120 I,WC(1201,WD(1201, 1MC T 1 ( 9 I /9* C . 000 /,MC T2 ( 9) /9*C. CDC/, MC 1U 20 )_, MC2 U 20 I ,

(9J79*6.obd7",bsoRf 1 SUMT1 (9) /'-(*0.0()0/, SUMT CHARACTER CHAR*12(9 I,TREE*3(120 I,DISC*2(120I ,CR0WN*1(120 I

INITIAL READ IN CF ALL DATA

i i

I

!

30

REA0,M,N,T1,T2 DO 30 1 = 1,N RE AO, C H AR ( I) RE All 5 ,1 .END = 50 I ( TRE E ( I I , SPEC ( I 1 .DISC (I) .CROWN (I ), AI l l ,

IB( I),(. ( I),0( I ), BOTTLE ( I ) , I=1,M) FORMAT! A3, 11, A2 ,41 ,4F7.3, 131

10_ "i l

_L*L 13

GO Tl 40 C CONTROL TO H£RE IF END OF DATA ENCOUNTERED DURING READ IN BEFORE M CARDS READ 50 WRITE (6,21 M _ ^ ___

"2" FOR MA T ( • 1' , • E "f6 OF DATA" ENCOUNTERED BEFOR E '7 13, ' CARDS READ"1! 1'. ..RECOUNT CATA CARDS') GO TO 51

14 "1 5 16 1 7

40 . . CLNTI MUE C CALCULATE WEIGHTS

DO 100 1=1 ,M LESS BOTTLE WEIGHT AND MC.IN PER CENT FOR EACH SAMPLE

W3(11= B(11-A( 11 WC. ( I I = C ( I ) - A ( I ) WD ( I I = D ( I ) - A ( I )

18 19 20

MCI(I I= (W S(I)-WC(I)l/WBII 1*100.DO MC2( I l = (Wtl ( I l-WCI I I l/WBI I 1*100.00 CONTINUE _

C PR I NT OUT OF "GRIG INAL DATA. WEIGHTS MINUS BOTTLE WEIGHT,-~070 MC FOR EAtH SAPPT" C AT TEM^ 1 ANO 2 C **HE AOI NG ORINTOUT'

100

21 WRITE!6,10 I 22 10 FORMAT!'1' ,T2,' INITIAL PRINTOUT OF DATA READ IN', T69,

^ I >**«• ,T75, 'CALCULATIONS-WEIGHTS, 0/0 MOISTURE CONTENT' I 2 3 ' " 'WRITE (5, 11 I " " " ' ' 24 WRITEI6.12I T1,T2,T1,T2 25 11 FORMAT) 'U' ,T2, ' TREE ', T8,' SPEC ,T14,' PI SC , T20 , ' CP.CWN' ,T29, '*A*',

lT3a,'*3*',T47,•*C*',T56,<*D*',T63,'BOT',T69,'***',T75, l'WTS. MINUS EMPTY WT . • , T98 ,' MC (0/0 ) •, T 10 7 , • MC (0/0 ) • )

26 12 FORMAT!' • ,T3, 'NO. ' ,T15, 'NO. ' ,T21, 'LOC. • ,T28, • EMPTY' ,T36_, "1 'UNJRYET ,'T45, I 3'i ' ~ORY' ,^54,13", • DR Y • , T 63",'' NO. ' ,"T69, •***r7T7"57 1' *H* *C* *D*',T98,13,• DRY' , T107, 13,' DRY')

C **P«tNTOi.lT OF ORIGINAL DATA AND CALCULATIONS 27 WRITE (6,13 I 28 13 FORMAT ( • 0' I 29 wR ITE (6,14 ) ( TRE E( I ) ,SPEC( I I ,DI SCI 1 ) , CROWN (I ) ,A( I I ,R( 11 ,CI I I ,D( I ), """ : " 1 S 0 T T L E ! I I, W*< T I ,WC( 11 , WDI I ), MC 1 ( I) , MC2 ( I ) , 1=1, Ml""

30 14 FORMAT!' ' , T 3 , A 3 , T 10, 11 , T15 , A 2 , T22 , Al , T27 ,F 7 . 3 , T36 , F7 . 3 , T45 , 1F7.3,T54,F7.3,T63, I3,T69,'***',T76,F5.3,2X,F5.3,2X,F5. 3.T9 8,

© r 1F7.3,T107,F7.3)

C CALCULATING AVERAGE MOISTURE CONTENT o 31 00 99 9 1=1 , M SI

32 COUNT! SPEC ( I ) l=C0UNT( SPEC I 1)1 + 1 33 MC T i l SPEC! I 1 I = MCT1 (SPEC! I ) l+MCK I)

o 34 MC T>( SPEC( I ) 1 =MCT? ( SPEC! I) )*MC2 ( I 1 > © 7 35 999 CONTINUE

36 00 10 0 0 1= 1, N o 37 IF ( CJ 'JNTI I 1 . LE. Ot GO TO 1 0 0 0 o

3 3 MC T I ( I ) = MC Tl ! I 1 /COUNT I I ) 3 9 MO. T2( I )=MC T2 ( I 1/COUNT < I 1

o 40 1P00 CONTINUE o. C CALCULATING STANDARD OEVI AT ICN FROM FORMULA... C STO OEV = SORTISUM FROM 1 = 1 TO M OF ( X < I t-A VG( X ) )** 2/( M-11 )

*s 41 DO 1 ) 0 2 1=1, M C1

4 2 S'JMTl ( S P E C I I ) I = SUM T 1 ( SPEC ( I I 1•(MC1( I 1-MCTK SPEC! I ) 1 )**2 43 SUMT2(SPEC I I I t=SUMT2(SPEC( I1 1 + IMC2!I)-MCT2(SPEC( I)I 1**2

-> 4 4 1002 Cf N T I N U t f ' 45 DO 100 3 1=1, N 46 IF(COUNT!Il.LE.1) GO TO 1 0 0 4

o ' 47 SUM Tl ( I l=DSORT( SUM T1 ( I ) /(COUNT! 11-11) <~. o ' 48 SUMT2 ( I l=OSQRT( SUMT2! I ) / (CCUNTI 11-1)1 4 9 GO TO 1 0 0 3

Q 50 1 0 0 4 s i r m ( i )=e .000 O 51 SU MT2 ( I ) =0 .0 DO 52 1003 CONTINUE

n C FINAL °R I NT OUT c n 53 WRITEI6.2rOITl.T2 54 200 FORMAT!'-' ,T4,'SPFCIFICAT ICN' ,T23,'NUMBER',T33,,**,,13,' DEG. C ',

O 1'DRYING**'.T56, •**•', 13. • DEG. C DRYING**') c O 55 WR I TE ( 6 » 20 11 56 201 FORMAT!' ',T22,'IN GROUP')

O 57 WRITE I 6, 20 2) 53 202 .FORMAT!' ',T35,' AVG MC S TO DEV ' »T58, • AVG MC STD DEV •) 59 WRITE (6, 20 21 )

o 60 2021 FORMAT! ' 0 ' 1 61 00 100 1 1 = 1, N 62 IF ( C O U N T ! Il.LE.0» GO TO 800

D 63 WR I TE ( 6 ,2C 3 1 I ,CHAR ( I) ,COU^( I 1 , M , MCT 1 ( I 1, SUMT1 ( I 1.MCT2I I 1, c 1SUMT2III

64 203 FORMIC ',T2,I3,' - ' , A12 , T23, 13, • / •, I 3, T35, F 7. 3 ,T44, F 7. 3 , T58 , J> 1 F 7 . 3 . T 5 7 . F 7 . 3 )

65 GO TO 1 0 0 1 6 6 800 WR I TE ( 6 , 2 0 4 ) I ,CHAR ( I I .COUNT! I ) , M , I 67 204 FO RM\T(1 ',T2,I3,' - ' ,A12,T23,13,*/',I3.T75,

l>***|f| OAT A GIVEN FOR GROUP ',11,'***') 68 1001 CONTINUE

C CALCULATING TOTAL FOR ALL GRCUPS 69 MC T l ! 1 1 = 0. 000 70 MCT?! 1 1=0.000 71 DO 2 0 0 0 1=1, M 72 MCTK I ) = MCT1 ( 1 1 + MC 1(1) 73 MC T2( 1 ) = M 0 T 2 ( 1 1 *MC2 (I 1

v j : 74 2 0 0 0 CONTINUE' 75 MCTK 1 ) = MCT1 ( 11 /M

5 76 MCT2I 1 1 =MC T2( 1 ) /M 77 DO 2 0 0 1 1= 1,M \^

73 SUMTK 11= 0 . 0 0 0 . 79 SUMT2 ( 1 ) = 0 . 0 0 0 80 SUMTK 1 >=SUMT1( 1H-IMCK I )-MCTl(l)l**2 >

http://WRITEI6.2rOITl.T2

81 SUM T2 ( 1 ) = S UMT2(11•< MC2(I I-MCT2I 1) )**2 82 2001 CONTINUE 83 SUMTK 1 )=OSQRT( SUMTH l l / I M - l l ) 84 SU MT2 ( 1 I =0 SORT ISUMT2(1 I/(M-lI)

C PRINTOUT OF TOTALS 8 5 WR irE16.?06l",M, MCTK 11, SUMT1 ( 1 I ,MCT2( 1), SUMT2 I 1) 86 206 FORMATCO' , T 2 , ' * * * * A L L GROUPS****•,T23,13,•/•.13,T35,F7. 3,

1T4*,F7..3,T53 ,F7.3,T67 ,F7.3) 87 51 WRITE ('6,20 51 •_ 8 3 205 FORMAT P I ' I 8 1 RE TURN go__ EN 0 .

tDATA

157

A P P E N D I X 9

S P E C I F I C G R A V I T Y PROGRAM


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A P P E N D I X 10

PROGRAM FOR C A L C U L A T I N G BIOMASS D I S T R I B U T I O N

FOR SAMPLED T R E E S

SCOMPHE C - M O P E L OP BIOMASS DISTRIBUTION FOR 20 TREES FROM HIGH ELEVATION SAMPLE C - P L O T N E A R S Q U A M I S H .

I N T E G E R T R E E , S E C ( 1 0 ) , 8 P , A G E , B U T L 1 , R U T L 2 , C R O P t , C R O D 2

R E A L V O L W ( 1 0 ) , V O L B ( 1 0 ) , S P G W ( 1 0 ) , S P G B ( 1 0 ) , L , l . T 0 6 , L T 0 1 , D B H , B U T L , I C R O D . F K B I , F w T N S ( 3 ) , F W S B l , F W S B U , F W S T N ( 3 ) , F W B N H ( 3 ) , F W H 4 S ( 3 ) , F W C O ,

2 F W S C 0 . P W B 1 , B w S B 4 , D W S T ( 3 ) , P W S N ( 3 ) #DWSCO,DWWSC 1 0 ) , D W B S U O ) ,[)WW,DWB, 3 M C 8 l , M C R « , D W B U ( 3 ) . M C T N ( 3 ) , D W T N ( 3 ) , D W T ( 3 ) , D W N ( 3 ) , D W B « T , D w T N T , D W T T , 4P W N T,0W C0,D W H1,M C C0

I T K E E=0

C - C A L C U L A T I O N FOR A T R E E B E G I N C - R E A D I N T R E E P A R A M E T E R S A N D C R O W N C O M P O N E N T S F R E S H W E I G H T S

DO 1 0 0 1 = 1 , 2 0

H E A D ( 1 , 1 0 1 ) T R E E . S P ; A G E , L 1 , L 2 > L T 0 6 t , L T 0 6 2 , L T 0 1 1 , L T 0 1 2 , R B H , B U T L 1 »

"" ' i J U U T L 2 , C R 0 n i , C R 0 D 2 , F W H 1 , C F W T N S ( M ) , M s 1 , 3 ) , F W S B l , F W S B 4 , ( F W S T N ( K j

2 K = 1 , 3 ) , ( F W B H R ( J ) , J s l , 3 ) , < F W R 4 S ( N ) , N = 1 , 3 ) , F W C O , F W S C O

1 0 1 F Q R M A T { 1 3 L LL I 3_X j j . 3 , 3 ( I J , Hl,±; i , 1 , 4 ( 1 2 ) , 4 X , F 6 . 1 , 3 X , i ( F S . l ) , 5 ( F 3 . 1 ) /

1 3 X , i ( F 5 .1 ) , 3 ( F 4 , 1 ) , F 4 , 1 , F 1 , 1 )

C - R E A D I N D R Y W E I G H T S O F C R O W N C O M P O N E N T S

7 R E A D ( 5 , 1 0 2 ) D W S B 1 , D W S H « , C D W S T ( K ) , K s l , 3 ) , ( D W S N ( J ) , J = 1 , 3 ) , D W S C O

F b R M A T ( « X , B ( M , 3 ) , F 3 , 3 ) " ' • " " '

D W « s D W B = D W R U T = P W T N T i D W T T = O W N T = D W C O a D W B l s O , 0

102

5 , 0 0 0

6 , 0 0 0

7 , 0 0 0

8 , 0 0 0

_ ' ' , 0 0 0

1 0 , 0 0 0

1 1 , 0 0 0

1 2 , 0 0 0

1 3 , 0 0 0

1 1 , 0 0 0

1 5 . 0 0 0 _

i 6", 600 1 7 , 0 0 0

1 8 . 0 0 0

1 9 , 0 0 0 "

2 0 , 0 0 0

J ' • ooo 2 2 , 0 0 0

2 3 , 0 0 0

2 U , 0 0 0

" 2 5 , 0 0 0 "

2 6 , 0 0 0 _ * E J t T E N 3 I 0 N * O T H E R C O M P I L E R S MAY NOT A L L O W M U L T I P L E A S S I G N M E N T S T A T E M E N T S

• E X T E N S I O N *





' E X T E N S I O N *

1 0

1 1

' 2 — 1 3

O T H E R C O M P I L E R S M AY NOT A L L O W M U L T I P L E A S S I G N M E N T S T A T E M E N T S


O T H E R C O M P I L F R 3 M AY NOT A L L O W M U L T I P L E A S S I G N M E N T S T A T F M E N T S

O T H E R ' C O M P I L E R S ' M A Y * NOT A L L O W M U L T I P L E A S S I G N M E N T S T A T E M E N T S "



DO 2 Ma i ,3 O W T ( M ) = 0 , 0

D W N ( M ) = 0 , 0

1« 1 5

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1 8

1 9

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2 4

2 5

2 6

2 "* 0 W T N ( M ) 3 O , 0 '

C-DATA T R A N S F O R M A T I O N TO M E T R I C S Y S T E M

L = 0 , 3 0 « f l * F L O A T a i ) + 0 , 0 Z 5 « * F L O A T ( L 2 )

L 1 6 6 6 , 3 6 « * * r-L o A t " L"T 0 6 i ) • b , o "2 S a * F L b A T (L T o 6 2 y ~

L T O 1 * 0 , 3 0 4 8 * F L O A T ( L T 0 1 1 ) • 0 , 0 2 5 4 * F L O A T ( L T 0 1 2 )

_ _ _ B U T L = 0 , 3 0 i 4 8 * F L O A T ( B U T L l ) + 0 , 0 2 5 « * F L O A T ( B U T L 2 )

'" D B « i O , 0 2 5 < 4 * O b H : '[

C R n D s ( 0 . 3 0 i 4 B * F L O A T ( C R O D l ) + 0 , 3 0 a 8 * F L O A T ( C R 0 0 2 ) ) / 2 , 0

; F W H l s 0 , « 5 i 5 9 2 * F W H l ' •

" F W S B l = 0 , 4 5 3 5 9 2 " i F " w S B " i ~ ~

F » S B 4 s 0 . 4 5 3 5 9 2 « F ^ S B 4

C R O L = L - H U T L

00 1 0 3 M = 1 , 3 " •" ""' " ' • ' - '

F W T N S ( M ) s 0 , « 5 3 5 9 2 * F W T N S ( M )

F W S T N ( H ) s 0 , 4 5 3 5 9 2 * F W 3 T N ( M ) • ' \- ' -.27 F - H N R ( M ) s O , a S J 5 9 2 * > W H N R ' M ) .. ,}

28 103 F w B 4 S ( M ) s 0 , 4 5 3 5 9 . 2 * F W . R 4 S ( M )

29 F W C P = 0 , u 5 3 5 9 2 * F W C O ' _ -••'

3 0 '" " • F W S C O s 0 . 4 S 3 5 9 2 « F * S C O ""'

C - B I O M A S S F O R T R F E T R U N K (WOOD A N D BARK) C - R E A D I N S P E C I F I C G R A V I T I E S O F S E C T I O N S

3 1 R E A D ' ( 3 , t 0 6 ) ( S P G W ( J ) , S P G B ( J ) , J « 1 , 1 0 )

- 3 2 1 0 6 F 0 R M A T ( 2 I X , F 5 , 3 , l i X , F 5 , 3 )

3 3 0 0 1 0 4 j = !, 1 0 . . . '. ?.'

C - R E A D I N V O L U M E S OF S E C T I O N S • ~r;-; ; - ' ~; lU R E A P ( 2 , 1 0 5 ) V O L W ( J ) , V P L B ( J ) -'-

3 5 1 0 5 F O R M A T ( 1 5 X , F 5 , 3 , 2 X , F 5 , 3 ) .

36" " " " V O L W ( J ) s i 0 0 0 * v b L W ( J ) .

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3 6 , 0 0 0

3 7 , 0 0 0

3 8 . 0 0 0

3 9 , 0 0 0

4 0 , 0 0 0

4 1 , 0 0 0

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4 4 , 0 0 0

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A P P E N D I X 11

PROGRAM FOR C A L C U L A T I N G N U T R I E N T D I S T R I B U T I O N


iCOMPILE C **NUTR1ENT DISTRIBUTION PROGRAM** C FUR UP TO 99 TREES

INTEGER TREE,SPECIE.NUMBER,NTREES REAL L.DBH.CROD.CROL,BUTTL,DWB1,DW84T,DWCO,DWTT,DWNT,DWTNT, 1DWTI3) ,DWN(3),DWTN(3),TRET,DWWSI 10),OWW,DWBS110).DWB,MCDATA( 8)

1. 000 2.000 3. 000 4. 000 5.000 6.000

REAL NPCR!21),PPER(2i),CAPER!211.MGPEP (21),KPER(21 I,WT(6),NUT(6) R = AL NW0HDC7,51,NBARK(7,5 I ,NB1(5),NB4T(5 I,NT(4,5),NN(4,5),NC0(5>,

J.NTN! 4,5), PE-RCNT.. TOTAL (5.) , JNN ( 3 (., TN P ( 3 I ,TNCA( 3 )., TNMG ( 3 ), TNK (.3 ) , 1ZER0I 3 l/3»0. 0/ COMMTN N!JM3;R/AREA2/MC0ATA COMMON /AREA! / N°ER . P PEP., CAPE R, MGP ER . K P ER

7. 000 8.000 9 . 000 10.000 11.000 1 2. 000

99

CHARACTERS 1 CHAR(5) / 'NITROGEN 1'MAGNESIUM -',«POTASSIUM -'/

_REAO \ \ OF. NUMBER OF TREES REAM 5 , 99 INTREE S FCRMATI 12 I

R£AO IN OF .MOISTURE CONTENT DATA

-• , • PHOSPHORUS- ', 'CALC IUM 13,000 14.000 15. 000 16, 0;)0 17, 000 18. 000

10 REVO! 5 ,100) (MCDATA! I) ,1 = 1 , 11 100 FCRMATI3IF5.3.1 X) )

_XZ 0) 1 1 =1 j.N.TREE S

8)

READ IN OF BIOMASS DATA FOR ONE TREE L3 READ!5,1011 TRE E,SPEC IE,L,DBH,CROD,CROL,BUTTL,DWB1,DWB4T, DWCO,

1DWTT, OWNT,DWTNT,(OWT! J) , DWNI J ) ,DWTN(J) , J = l ,3), TRE T, DWWS ,DWW, DWBS ,

19. 000 20. 000 21,000 '"22.000" 23.000 24. 000

inwB ' 14 101 FORMAT! I 3,1 I ,F4.1 ,8X, F5. 3 , 3F4.1 ,2F6. 1, F4. 1.3F6. 1/7X.9F6.1, F9. 1/

17X, 10F6. 1 ,F9. 1 /7X.1QF6.1.F9.1) ;

15 NUM3ER=21 ' 16 IF! S°ECIE.E 0.2. OR.SPECIE. EQ. 3 I NUMB ER =1 8

C CALL SJ-iPRUGRAM TO REAP IN NUTRIENTS N AND P FOR ONE TREE AND RETURN

25. 000 26. 000 27.000 "28. 000" 29. 000 30.000

PERCENTAGES FOR EACH SPECIFICATION ( EG. WOOD , BAR K. . . ) 17 CALL SNP(NPER.PPER)

C CALL SJBPRUGRAM TO READ IN NUTRIENTS CA ,MG,K FOR ONE TREE AND RETURN C • PERCENTAGES FOR EACH SPECIFICATION

18 CALL SCAMGK (C AP ER ,MGP ER, KPER ) 19 1 F( SPECIE.EQ. 2. OR.SPECIE.EQ.3 ICQ TO 1004

31. 000 32.000 _3_3_. 000_ 34. 000 35. 000 36. 000

?0

C CALCULATING PERCENT NUTRIENTS FOR T WIGS CFOLI AGE FOR SPECIES 1 BY AVERAGING 37.000 C THE PERCENTS FOR TWIGS AND PERCENTS FOR FOLIAGE 38.000

N1 = 14;N2 = 17 • • 39.000 22 23 24

00 29 K=l,3 TNNIK ) = ( NPER1K + N1 ) «-NP ER ( K+N2 ) )/2 T.NP I <) = ( PPER (K>N1 l+NP£R(K»N2) 1/2

40.000 41.000 42. 000

25 26 27

TNCAIK) = (C*PER(K + NlH-CAPER(K + N2))/2 TNMG! K 1= ( MGPERI K+Nl ) +MGPER (K+N2) ) / 2 TNK!<)=(K0ER!K+N11.KPERIK+N2)1/2

28 29 CONTINUE 29 1004 CCNTINUE

C *««CALCULAT1NG NUTRIENT CONTENT BY SAMPLE SPECIFICATION*** C SSPEC - WOOD C CALCULATES NUTRIENT CONTENT FOR WOOD FOR THE 5 NUTRIENTS AND STORES IN ARRAY C NWOOO!7,51.COMBINES BIOMASS WOOD DATA FOR SECTIONS 1C2.3C4,5tb,7C8,9,10 .

30 31 32

DO 2 J = 1,4 WT! )l =DWWS(2*J-1)*DWWS(2*J) CONTINUE ' . . ":

43.000 44.000 4 5. 000_ "4 67 000 47. 000 48. OOP 49.000 50. 000 51. 000 52.000" 53. 000 54.000

33 WT!5)=DWWSI91 34 1T( 6) = DWWS( 10)

C GETS NUTRIENT PERCENTS FOR APPROPRIATE SEC TlONS,AVER AGES,PUTS IN ARRAY NUT(6) 3 5 "00~i J=l,5 36 NWOOO! 7, J I =0.0 37 N = l - -

"55. 000 56. 000 57.000 5 8.000" 59.000 60. 000

f 38 GO TOt 10,20,30,40,501 , J 61.000 39 10 CALL CALC(N,NPER,NUT,£60) 62.000 40 20 CALL CALC(N,PPER,NUT, £60) 63.000 41 30 CALL CALCIN,CAPER,NUT,660) 64.000 42 40 CALL CALCIN,MGPER,NUT,660) 65.000

y- 43 50 CALL CALCIN,KPER,NUT,660) 66. 000 44 60 00 4 K=l,6 67. 000 < 45 NWOOO IK,J )=WT IK ) *NUT 1 K ) 68. 000 46 NWOOO( 7, J)=NW00D(7, J) «-NWOOO(K,J) 69.000 47 4 CCNTINUE 70.000 48 3 CONTINUE 71. 000

© C SSPEC - BARK 72.000 C CALCULATES NUTRIENT CONTENT FOR BARK FOR THE 5 NUTRIENTS AND STORES IN ARRAY 73.000

© c NBARKI7,5),COMBINES 310MASS BARK DATA FOR SECTIONS 1£2,3£4,5£6,768,9,10. 74. 0 00

© 49 DO 5 J = l ,4 75. 000 50 WTI Jl = DW3S~< 2*J-1) *DWBS(2*J ) 76.000 51 5 CCNTINUE 77. 000 52 •Ul 5) = D*3S( 9) 78. 000 53 WTISI=DWBS(10) 79.000

C GETS NUTRIENT PERCENTS FOR APPROPRIATE SECTIONS,AVERAGES,PUTS IN ARRAY NUT I 6) 80. 000 0 54 no ••> .1 = 1,5 81.000

55 N6AUI7, J) = 0.O 82. 000 56 N = 7 e3.000 57 GO TlI 11,21,31,41,51),J 84.000 59 11 CALL CALCIN,NPER,NUT,£61> 85. 000

© 59 21 CALL CALCIN,PPER,NUT,£61) 86. 000 © 60 31 CALL CALCIN,CAPER,NUT,£61) .87.000 61 41 CALL C ALC I N, MGPER ,.NUT , £61 ) 88. 000 62 51 CALL CALC(N,KPER,NUT,£61) 89. 000 63 61 DO 1 K=l,6 90.000 64 NBARKIK,J) =WT I K)*NUTIKI . 91. 000 65 NiOAR< I 7, J )=NBARK( 7, J) tNBARKIK.J) 92.000 66 7 •CONTINUE 93. 000 67 6 : CCNTINUE 94. 000

C SSPEC - DWB1 95.000 68 K=0 - 96.000 69 N = 13 97. 000 70 DO 3 J = 1, 5 98.000 71 CALL c IXI N, J,K, PERCNT ) 99. 000 72 NdllJ)=0W91*PERCNT 1 00. 000 73 8 CONTINUE 101.000

C SSPEC-JW34T 102.000 74 K = 0 103.000

U 75 N = 14 1 04.000 U 76 • 0 9 J = l ,5 105. 000 77 CALL =IXIN.J.K,PERCNT) 106.000 ' • ~ - • - - —

to 73 N34TI J ) = DW34T*PERCNT 107.000 to 79 9 CCNTINUE 1 C8. 000

C SSPEC-D^T 109.000 80 DO 12 K=l,4 110.000

«» 81 DO 10 J=l ,5 111. 000 82 NT! K, J ) = 0. 0 112.000 • ' " -83 13 CONTINUE 113.000

• 84 12 CCNTINUE 114.000 35 I F( SPECIE.EJ.2.0R.SPECIE.E0.3 IGO TO 1000 115.000

; 86 N = 14 116.000 87 DO 1H K=l,3 117.000

• 88 56 15 J=1,5 118.000 -

89 CALL F IXI N,J, K, PERCNT ) 119.000 • 90 NT[ K, J ) =DWTIK)*PERCNT 120.000

&

i

9 j

i

• j

91 NTI4, J1 = NTI4,JI*NT(K, J l 121.000 92 15 CONTINUE 122.000 93 14 CONTINUE 123.000 94 1000 CONTINUE 124.000

C SSPEC OAN 125.000 95 00 16 K=l,4 ; 126. OOP 96 97 98 17

00 17 J = l,5 KN( K, J 1=0.0 CONTINUE

127. 000 128.000 129.000

99 100 101

16 •CONTINUE IFISPECIE.EQ.2. OR. SPECIE. EO. 3 ) GO TO 1001 H = 17

130. 000 131.000 132.000

102 103

• 104

DO 13 K=l,3 DO 19 J=l,5 CALL PIXIN.J.K, PERCNT 1

133. 000 134.000 135.000

105 'MNK,J) = OWN(K»*PERCNT 136. 000 106 107 19

MMI4, J ) = NN ( 4 » J) +NNI K, J) CONTINUE

137.000 138.000

ioa 109

18 CONTINUE 1001 CONTINUE C SSPEC—'MCO

1 39. 000 140.000 141.000

110 111 112

N = >1 !F( S?ECIE.EQ.2.QR.SPECIE.EQ.3)N=15 K = 0

142. 000 143.000 144. 000

113 114

DO 22 J = 1 , 5 CALL FIX<N,J,K,PERCNT1

145. 000 146.000

115 NCOIJ)=DWCO*PERCNT 147. 000 116

117

22 C

CONTINUE SSPEC-OrfTN

DO 23 K=l,4

148.000 149.000 1 5U. 000

118 119 120 24

~ 00 24 J - l i 5 NTNIK,J) =0.0 CONTINUE

151.000 152.000 1 53. 000

121 122 123

23 CONTINUE IF(S°ECIE.EQ.2.0R.SPECIE.EQ.31 GO TO 1002 DO 25 K=l,4

154.000 155.000 1 56. 000

124 125

DO 26 J=l,5 NTNIK,Jl=NT(K,J)*NN(K,J)

157.000 158. 000

126 26 CONTINUE 159. 000 127 128 129

25

1002

CONTINUE GO TO 1003 N = 15

160.000 161.000 162.000

130 131 132

00 27 K=l,3 DO 23 J=l,5 CALL F I X 1 N, J , K, PERC NT I

163.000 164. 000 165. 000

133 134

NTNIK,J)=OWTN(Kl«PERCNT NTNI4,J)=NTN(4,Jl+NTNtK,J»

166.000 167. 000

135 28 CONTINUE 168. 000 136 137

27 CONTINUE 1003 CONTINUE C CALCULATING TOTALS FOR THE TREE

169.OOO 170.000 171.000

133 139

• 0 32 J=l,5 ' TOTAL I J) = NWOOC( 7 , J ) +N BARK (7, J 1+NB1 (J I+NB4TI J )• 1NC0I I)*NTNI 4,J)

172.000 173.000 174.000

140

141

32 C

CONTINUE • ••PRINTOUT***

WRITz16, 200)TRE E.SPECIE,L,D8H,CROD,CROL,BUTTL

175.000 17t. 000 177.000

142 200 FORMAT<• 1'//• ','TREE',3X,'SPECIES',3X,'LENGTH' I'WN1, 3 X,1 CROWN' ,3X,•BUTT'/• ' ,1X,•NO.•,5X,•NO.'

,3X,1DBH' , 3X, ,7X,'(MI',4X,

'CRO' , MM)',

178.000 179. 000

13X.'0IA(MI',2X, 'L.(M) ',3X,'L.(Ml•/• • ,1X, I3.6X, 11 ,7X,F4.i,3X ,F5.3, 180.000 )

13X ,F3.1 ,5X,F4.1 ,3X,F4.1/////' ' , ••••PERCENT NUTRIENT CONTENT BY •, 1•SAMPLE SPEC I F I C A T I O N ' , 7 9 ( • • • ) / ' •,T16,'*•,151•-• 1,•WOOD**SECTlON'

181.000 182.000

\

9

1*3

1 , 1 S ( ' - ' I , ' * ' , T 7 9 , ' * ' , 1 5 ( ' - , I , 'BARK**SECTI0N',15<'-• 1 ,'*•/• 12( 16X, '1 • ,7X, '3 • ,7X,« 5',7X,'7' ,7X, '9< , 7X, • 1 0' , 7X) / I W R I T E ( 6 , 2 0 1 ) ( N P E R I J I , J = i , 1 2 1 , ( P P E R ( J I , J = l , 1 2 ) , ( C A P E R ! J ) , J = 1 , 1 2 ) ,

1 < MGPER! J I , J = l ,1 2) , ( KPFRI J I , J = l ,1 2 I

183.000 184.000 185.000 186. 000 J

201 FORMAT!' •,2( 'NITROGEN -',6(2X,F6.3),6X)/2('PHOSPHORUS-', 16<2X,F6.31.6X )/2('CALCIUM - • , 6 ( 2X, F6. 3 ), 6X)/2( ' MAGNE S IUM - ' ,

] 87.000 188. 000

\

16(2X,F6.3),6X1/2('POTASSIUM - • , 6 ( 2 X , F 6 . 3 ) , 6 X ) ) 189.000 145 WRITE (6 , 202) 190.000 146 202 FORMAT!•-•,T13,2!• 8RANC H•),3X,'*',7('-'),'TWIGS',7!'-'),'*',3X, 191.000

l'*',6('-'),'FOLIAGS',7('-'),'*',3X,'CONES'i3X,'*',3('-'l,'TW!GS + '» 192.000 1•FOLIAGE',' *•/' ', T17, •>1•,5X,•1-1/4',2(3X,•(TOP)',3X,•( MIDI • , 193.000 13X, • ( LOW I • 1 ,12X ,' (TOP) • ,3X,' ( MIDI • ,3X, • (LOW) • / ) 194. 000

147 . tF(S?ECIE.EQ.2.0R.SPECIE.EQ.3l GO TO 1006 195. 000 143 WRIT£(6,2)3)(NPER(JI,J = ) 3 , 2 1 I , ( T N N ( J ) , J = 1 , 3 ) , ( P P E R ( J ) , J = 1 3 , 2 1 ) ,

1 ( TN»( J ) , J = l, 3 ) , (CAPER! J l , J = 13 ,21 ) , (TNCAI J 1 , J=1,3I , 196. 000 197.000

® 1(MGPER(JI,J=13,21 ),(TNMG!J),J=1,3),(KPER(JI,J=13,21I , 199.000 1 ( TN<( J ) , J = l , 3 ) 1 99. 000

149 203 FOR MA T ( 1 ', 'NITROGEN - • , 8 (2 X, F6. 3 ) ,1 X ,4 I 2 X, F6. 3 1 / • PHOSPHORUS-• , 1 8 ( 2X, F6. 3 1, 1X.4I2X, F6 . 3 ) / 'CALCIUM - • , 8( 2 X, F6„ 3) , IX , 4( 2 X ,F 6= 3) /

200.000 201. 000

1 • MAG'IESIUM -',9(2X,F6.3), 1 X, 4 ( 2X , F6. 3 1 / • P OT ASS I UM - •, 8 ( 2X, F 6. 3) , 202.000 U X , 4 ( 2X.F6.3I / / / / / I 203.000

@ 150 G") T l ICO5 204. 000

©

151 1006 WRI T? ( 6, 2031 (NPERU 1 , J = 13 ,14) , ( ZEPOI J l , J = l, 3) , 1 ZERO! J) , J = l , 3) , 1 ( N P 5 1 ( J ) , J = l 5 , 1 3 1 , ( P P E R I J ) , J = 1 3 , 1 4 1 , ( Z E R O ! J I , J = l , 3) , H Z = ,=<3 ( J l , J»l, 3) , (PPER ( J 1 , J = 15 ,18 ) , (CAPER! JI »J-13»14) ,

205. 000 206.000 207.000

1 ( ZERO! J l , J = l , 3 1 , ( ZERO I J l , J = l, 3) , (CAPER ( J l , J * l 5,181 , 2C8. 000 1(MGPER!J1.J=13 ,14),(ZERO!J),J = l , 3 ) , ( Z E R O ! J) iJ = l,31, 209.000 1 (MGPc R( J ) , J = 1 5 , 1 8 l , ( K P E R ( J ) , J = 1 3 , 1 4 l , ( Z E R O ( J ) , J = l , 3 l , 210.000 1 1 ZERO ( J) , J = l ,3) , (K°ER ( J) , J = 15 ,18) 211.000

. ÊXTENSION* LIMIT OF 5 CONTINUATION CARDS EXCEEDED 152 100 5 CONTINUE 212.000 153 WRITE(3,2 99)TREE,SPECIE.DBH 213.000 154 299 FORMAT(I 3,11,F5.3I 214.000

@ 155 WRIT?(3,300)INPER!J1,J=l,121 215.000 @ 156 157

WRI Tc ( 3, 300) ( PPER1J 1 , J = l , 12 1 WRITE 1 3, 300i (CAPER! J l ,J=1,121 , • vi'

216. 000 217.000

<i 159 WRIT? ( 3,300) (MG PER( J) , J -1 ,12) ' : ' ' 218.000 159 WRIT?(3,3001(KPER!J),J=l,12) > 219.000 160 300 FORMATI12F5.3I 220.000 161 I F ! S'ECI E. EO. 2. OR, SPECIE. EO. 3 1GOT03000 221. 000 162 WRI TE ( 3, 3 011 (NPERI J I, J = 13,20) 222.000 163 WRITS ( 3,3011 < P P E R U 1, J = 13,201 223. 000 164 WRI TS ( 3, 3011 (CAPER! J) , J=13,20l 224.000 165 WRITE(3i301)(MGPER!J),J=13,20) 225.000 166 WRI T? ( 3, 3011 ( K P E R U I, J = 13,20) 226. 000

* 167 301 FORMAT! 8F5.3) 227. 000 163 GOT03 001 228.000 169 3000 WRIT?(3,3 021( N P E R ! J I , J = 1 3 , 1 4 ) , ( N P E R I J l , J = 1 6 , 1 8 ) 22 9. 000 170 WRITE! 3, 302) ( P P E R U ) , J = 13,14) ,(PPER(J),J=16,18) 230.000 171 WRITE( 3, 302 ) ( C A P E R I J ) , J = 1 3 , 1 4 ) , ( C A P E R ! J I , J = 16,18) 231. 000 172 WRITE ( 3,3 02) ( MGPER! J) , J= 1 3 ,14 1 , ( MGPER ( J 1, J = 16,1 8 1 232. 000 173 WRITE ( 3, 302) (KPER! J I, J = 13,14 l ,( KPER! J) , J=16,18l 233.000

i 174 302 F0RMATI5F5.3I 234. 000 ; 175 3001 CONTINUE . 235. 000

C •**PRINTJUT OF NUTRIENT AMOUNTS*** 236.000 176 WRI T? ( 6 , 2041 237. 000 177 204 FORMAT!' •,* ***NUTRIE NT AMOUNTS IN DECAGRAMS BY SAMPLE ', 238.000

1•SPEC IFICATION' ,74( '* • I / / ' ',T15,'*',16('-'I,'WOOD**SECTI ON' , 239.000 J

f

ITS 179

lao

118( •-'1.•*•,T80,•*•,16(»-'),'8ARK**SECTI0N',18('-'),'*'/' ',2<15X, l ' U 2 364 566 768 9 10 TOTAL ' l / l 00 2000 J=l,5 WRITE ( 6, 205) CH AR I J ), ( NWOOOI K, J) ,K = 1 ,7 I ,C HAR ( J ) , ( NBARK IK, J ) ,

1K = 1,7) 205 FORMAT!• ' , Al 1,1 X ,6 (1 X, F6. 2 1 , IX, F7. 2 , 3X , A 11 ,1 X, 6( 1X, F6. 2 1 ,1X, F7. 2 I

240.000 241.000 242. 000 243.000 244.000 245. 000

N

J f 181

132 183

2000 CONTINUE WRITEI6.206)

206 FORMAT! • 0',T13,2!IX,'BRANCH'l.lX.'+'.lOt'-'l.'TWIGS'.ll!'-'),'*',

246. 000 247.000 248. 000

\

1IX,'*•,91 •-'),'FOLIAGE',10!'-'),•*•,2X,'CONES',IX,'*•,6!' - • ), 1•TWIGS*F0LIAGE',7!•-•1,'*•,5X,'TREE'/• ',T17,'>1",4X,'1-1/4',2X, 12IMT0P) (MID) (LOW)',' TOTAL '),7X,

249.000 250.000 251.000

184 185

IMTQPI ( M i l l (LOW) TOTAL TOTAL'/) DO 2001 J = l , 5 WRI TE (6,207) CHARI J > ,NP.l 1 J) , NB4T ( J I , (NT ( K , J I , K= 1, 4 1 , INN (K , J 1 , K=l, 4

252.000 253. 000 254. 000

186 I ) ,NC:> ( J 1 , (NTNIK, J 1 ,K = 1,4 1 ,TOTAL( J )

207 FORMAT!' • , Al 1, 511X , F6. 2 ) , IX , F7. 2 , 3 (IX , F6. 2 ), 1X , F7. 2 ,4! IX, F6. 2 ), II X, F7. 2, 2X.F3.2 )

255. 000 256. 000 257. 000

187 193 189

2001 CONTINUE 1 CONTINUE

WRITE(6,203I

258. 000 259. 000 260.000

190 191 192

208 FORHATI'l'l RETURN ENO

261.000 262.000 263.000

193 SUBROUTINE SNPINPER.PPERI C READS IV NUTRIENTS N AND P FROM FILE SOUAM5,ADJUSTS FOR WEIGHT OF SAMPLE

264. 000 , 265.000

194 105

C AND AVERAGES FOR THE TWO SAMPLES TAKEN FPOM EACH SPECIFICATION INTEGER NUM3ER, SSPEC REAL VDS,W0,NPPM,PPPM,Z,NPER(21) ,PPEP.(21I

266. 000 267. 000 268.000

196 197 198

. COMMON NUMBER - ', , DO 1 1=1,21 . NPERI I l = PPER I I 1 =0.0

269.000 270.000 271.000

•EXTENSION* OTHER COMPILERS MAY NOT ALLOW MULTIPLE ASSIGNMENT STATEMENTS 199 1 CONTI N'JE

C READS IN 'NUMBER' RECORDS ACCORDING TO SPECIES -272. 000 ., 273.000 •

200 C SPECIES 1 -NUMBER =21,SPECIES 2 -NUMBER = 18

00 2 1=1,NUMBER C READS IN TWO RESULTS FOR EACH SAMPLE SPECIFICATION ANO AVERAGES

274. 000 275.000 276.000

201 202 203

0 3 3 J = l ,2 REATI 1,1001SSPEC,WOS.WD.NPPM,PPPM

100 FORMAT!3X,11,3X,2F7.5.3X,2F6.2)

277. 000 278.000 279.000

204 C ADJUSTS FOR WEIGHT. OF SAMPLE USED

Z=WOS-WD ' C CHECKS TO SEE IF THERE IS DATA FOR THIS CARD

280. 000 281. 300 282.000

205 IFIZ.GT.0.01G0 TO 4 206 NPERIIl = PPER(I I=0.0

•EXTENSION* OTHER COMPILERS MAY NOT ALLOW MULTIPLE ASSIGNMENT STATEMENTS

2e3. 000 284.000

207 208 209 •

GO TO 3 4 NPHRI I ) = NPER ( I ) + ( (N PPM/10000 ) *104. 20) 11

P PERI I ) =PPER (!)••( ( P PPM/10000 1*104. 20 )/Z

285.000 286.000 2 87. 000

210

211

3 CONTINUE C AVERAGES THE TWO READINGS

NPERI I ) = NPER( I I /2

288.000 289.000 290. 000

212

213

Pi'ERI I ) = PPER 1 I 1 /2 C CALLS SUBPROGRAM TO ADJUST FOR MOISTURE CONTENT

CALL MCI SSPEC,NPERI III

291.000 292. 000 293. 000

214 215 216

CALL MCI SSPEC,PPERI III 2 • CONTI NUE

RETURN

294.000 295. 000 296. 000

0 i ' 217 ENO 297.000

t

n

o

. 218

> 219

C

c

SUBROUTINE SCAMGKICAPER,MGPER,KPERI READS M NUTRIENTS CA.MG.K FROM S0UAM6,ADJUSTS FOR WEIGHT OF SAMPLE, ANO AVERAGES FOR THE TWO SAMPLES TAKEN FROM EACH SPECIFICATION

INTEGER NUM3ER,SSPEC

298.000 299.000 300.000 301. 000 J

Qj

0

i

I

1

r» • 220

221 222

HEAL CRU,CRUSA,CAPPM,MGPPM,KRE,KPPM,CAPER(21I,MGPER121),KPERI21),Z COMMC'N NUM3ER 00 1 1=1,31

302.000 303. 000 304.000

s 0

i ! «-

i r»

22 3 • EXTENSION* • EXTENSION*

CAPER ( I I = .MGPER( I 1 = KPER (11 = 0.0 OTHER COMPILERS MAY NOT ALLOW MULTIPLE ASSIGNMENT STATEMENTS OTHER COMPILERS MAY NOT ALLOW MULTIPLE ASSIGNMENT STATEMENTS

305.000 i

1

<%

224 1 c c

CONTINUE READS IN •NUMRER1 RECORDS ACCORDING TO SPECIES -S°ECIES 1 -NUMBER=21;SPECIES 2 -NUMB ER = 18

206.000 3 07. 000 308.000 Q

i i i

f» 22 5

22 6 c

00 2 I =1 , NUMBER READS IN TWO RESULTS FOR EACH SAMPLE SPECIFICATION AND AVERAGES

DO 3 J=l,2

309.000 310. 000 311.000 ©

I ]

<»

.22 7 228 100

c

RE AO! 2 ,100ISSPEC,CRU,CRUSA,CAPPM,MGPPM,KRE FORMAT!3X,11,3X.2F6.3 ,11X,F6.1,F6.2,F6.11

ADJUSTS FOR WEIGHT OF SAMPLE USED

312.000 313.000 314.000 ©

1

f*

229

230 c

Z=CRUSA-CRU CHECKS TO SEE IF THERE IS DATA FOR THIS CARD

IF! Z.GT.O.OIGO TO 4

315.000 31 6. 000 317.000 e

231 •EXTENSION* •EXTENSION*

CADERI I)=MGPER( I )=KPER(I 1=0.0 OTOER COMPILERS MAY NOT ALLOW MULTIPLE ASSIGNMENT STATEMENTS OTHER COMPILERS MAY NOT ALLOW MULTIPLE ASSIGNMENT STATEMENTS

318. 000

:

<»

232 23 3 234

4 GO TO 3 CAPER!I)=CAPER(11 •((CAPPM/100001*100.76)/Z MGPER< I 1=MGPER(I)+<(MGPPM/10000)*100.76)/Z

319. 000 320.000 321.000 f

235 236 237 3

KPPM= 3. dl 50+0.4605* KRE KPERI I ) = KPER ( I 1 «•( (KPPM/100001*100. 76l/Z CCNTI NUE

322. 000 323.000 324. 000

233 239

C AVERAGES THE TWO READINGS C AP;R 1 I 1 =CAP ERI 11/2 MGPER( I 1 =MGPER< 1112

325.000 326.000 327. 000

' 240

241 C

KPERI I l = KPER( I 1/2 CALLS SUBPROGRAM TO ADJUST FOR MOISTURE CONTENT

CALL MCI SSPEC ,CAPER! I I >

328.000 329. 000 330. 000

•

cj» '

242 243 244 2

CALL MC! SSPEC,MG»ER 1 I 1 1 CALL MC! SSPEC ,KPERI III CONTI NUE

331.000 332. 000 333. 000

t#

245 246

RETURN E NO

334. 000 335. 000

Qi 247 248 249

SUBROUTINE MC1SSPEC,DATAI INTEGER SSPEC REAL DAT A , MCDAT Al 8 1

336.000 337.000 338. 000 y

250 C

c

COMMON /AREA2/MCDATA . RECEIVES SAMPLE SPECIFICATION AND PERCENT NUTRIENT FOR THAT SPEC. AND ADIUSTS FOR MOISTURE CONTENT USING ARRAY MCDATA

339.000 340. 000 341.000

w •

251 252 253

DATA=DATA*(1-MC0ATA(SSPECI/100. 1 RETURN END

342.000 343.000 344. 000

254 255

SUBROUTINE CALC(N,XPER,NUT,*» REAL XPERI21),NUT(6)

345.000 346. 000 •

256 257

^ 258

DO 1 1=1,4 NUT(I)=(XPER(N)+XPER(N+1))/2 N = N*1

347.000 348.000 349. 000 •

1

259 1 CONTINUE 350.000 260 NUTI5)=XP5RIN) 351.000 261 NUT(6I=XPERIN*1) 352.000 262 RETURN1 353.000 263 END 354.000

264 SUBROUTINE FI X( N, J, K, PERCNT) 255. 000 265 REAL PERCNT,NPER(21 I,PPER(21),CAPER(211,MGPER(21),KPER(21) 356.000 _266 COMMON. ,/AREAl /._NP_ERjPPJ_P-» CAPER,_MG_P_ER,KP_ER 357. 000 267 GO TO ( 1, 21 3,4,5 ( , J 358. 000 263 1 PERCNT = NPER(K*N1 359.000 269 R-TUR N 360. 000 270 2 PERCNT^PPER I K .N 1 361.000 271 RETUfN 362.000 272 3 P£RCNT = CAPERI K + N! , , 363. 000 _ 273 RETJRN 364.000 274 4 PERCNT=MGPERIK+Nl 365.000 _27_5 RETURN 366.000 276 5 PERCN T-K P ER I K +N I 367.000 277 RETURN 368. 000 278 ENID • ' 3 69. 000

tDATA 370.000

A P P E N D I X 12

PROGRAM FOR C A L C U L A T I N G WEIGHTED CONCENTRATIONS

OF N U T R I E N T S W I T H I N D I F F E R E N T COMPONENTS OF

SAMPLED T R E E S

P r o g r a m p r e p a r e d b y M r , K e n T w a i t e s

t C O M P I L c 1 . 0 0 0

c * « * p * r i G W M T O C A L C U L A T E W E I G H T E D C O N C E N T R A T I O N S F O R N S A M P L E D T R E E S 2 . OOO

1 H E A L 0 3 0 1 ? 0 1 , T T , F T , T F T . T I 0 ) ,F ( 3 I , T F ( 3 ) .WOOD! 1 0 1 , WOODT , BAP.K ( 1 0 ) 3 . 0 0 0

2 R E A L B A R K T , M E A N ( 2 0 ) , N U T C O N < 2 0 , 5 ) , N U T S U M I 6 , 5 ) , N S U M ( 6 , 5 , 2 0 1 / 6 0 0 * 0 . 0 / 4 . 0 0 0

3 P. j ;AL r 3 H 1 5. 0 0 0

_ 4 _ : _ J N T E R_ TR E E 1 2 0 ) . S P E C I E ( 2 0 ) . F L A G ( 2 0 ) . N U M B E R , T R E E l , S P E C 11 6. 0 0 0

• 5 ' I N T C G F . R c f i S S ( 2 0 ) ' 7 . 0 0 0

C R E \ 0 1 4 J c N U M B E R OF T R E E S T O B E P P . O C E S S E D f U P TO 2 0 1 P.. 0 0 0

6 R S A )( 5,< 1 I N _ 9 . 0 0 0

7 ' 9 9 FCP. M\T ( 12 I " 1 C . 0 0 0

8 0 0 I L = 1 . N 1 1 . 0 0 0

T H I S 3 - C T I J N R E A D S I N B I O M A S S D A T A F O R O N E T R E E 1 2 . 0 0 0

9 RL'AOI 3",1 OOTTREE (L I, SPEC IE( L I • O B H ( L 1 , T T , FT ,TFT 1 3 . 0 0 0 1 0 1 0 0 F C n i l I 3 , II ,1 2 X . F S . 3 , 2 8 X , 3 * 6 . 1 I 1 4 . 0 0 0

1 1 _ _ I F ( S>'=CI E (L I . E 0 . 2 . 0 R . SPEC IEIL l . E Q ^ 3 I G O T 0 2 1 5 . 0 0 0

" l 2 R c A >( > ,1 01 I ( T ( I I , P ( 11 ,1=1 , 3 1 ' - - - 1 6 . 0 0 0

1 ? 1 0 1 F O R U T ( 7 X , 2 F 5 , 1 . 6 X . 2 F 6 . 1 , 6 X , 2 F 6 . 1 ) 1 7 . 0 0 0

1 t, G ) T ) l 1 6 . 0 0 0 2 ' R C A O I 3 , 1 3 2 ) ( T F ( I ) ,1 =1 , 3 ) " '"" 1 9 . 0 0 0

1 6 1 0 2 F U R 1 A T U ) X , F 6 . 1 , 1 2 X , F 6 . 1 , 1 2 X . F 6 . 1 I 2 0 , 0 0 0

1 7 3 R E A l t 3 , 1 0 3 I ( WCM 0 ( I I , I =1 , 1 0 1 , W OUOT, ( B A R K ( I K 1 = 1 , l _ 0 l j 3 A R _ K T _ : 2 1 . 0 0 0 Q

1 3 ' 1 0 ? ' F U R I A T ( 7 X , 1 Q F 6 . 1 , F9.1 / 7 X , 1 0 F 6 . 1 , F 9 . i I " " "" ' " •" * " 2 2 . 0 0 0 " ~~~ ~ C T H I S S E C T I O N W R I T E S T H E BIOMASS D A T A 2 3 . 0 0 0 -.;/ ,:•

J 9 WRI TJ I 6 , 2 0 0 1 ' 2 4 . 0 0 0 i^Lui!^-2 0 2 0 0 "FOR 1A" f (" • 1 • , • B 1 0 M A S S D A T A ' / / ) 2 5 . 0 0 0 i « ^ s f " 2 1 WR IT.11^,231 I T R E E ( L ) , S P E C I E I L ) , D B H I L I 2 6 . 0 0 0

2 2 2 0 ! F C R I A M * ' , ' T R E E ', 1 3 , 5 X , ' S P E C I E ' , 1 1 , 5 X , ' D B H • , F 5 . 3 / / I 2 7 . 0 0 0

2 3 ' ' W I r i * ( b , 2 12 I ( WOOD ( I I , I =1 ,1 0 ) ,'WOCDT , ( B A R K ("I I V I = 1~»1 0 1 , 3 A R K T • ~ " " Z f t . 0 0 0

2 4 2 0 ? F t R nr ( • ' , O X , 1 1' , 6 X , ' 2 1 , 6 X , • 3 ' , 6 X , , 6 X , • 5* , 6 X , ' 6 1 , 6 X , '7« , 6 X , ' 8 ' 2 9 . 0 0 0

1 , S X , • T , r>X, • i 3' , 6 X , ' S U M ' / • WOOD • , 1 0 ( F 6 . 1 » ! . X I , F 9 . 1 / ' B A R K •, 2 0 . 0 0 0 •

i i - . i i =.s". r . i x i , f 9 . rvi 3 i . w

2 5 I c ( i ° E C I E ( L ) . E 0 . 2 . 0 R . S P E C I E I L l . E O . 3 I G 0 T 0 4 3 2 . 0 0 0

2 6 I T - ( 5 , I J 3 > ( T ( I I , I = 1 , ? ) , T T , ( F ( I I , I = 1 , 3 I , F T 3 3 . 0 0 0

2 7 F j v u n • ' , =IX, • 1' , 6 X , >i< , 6 X , ' 3 " , 5 X , • S U M 1 / /' T W 1 G S ' « 4 T F 6 7 1 7 1 X 1 7 3 4 . 0 0 0

l ' F O L ' » 4 ( F j , l , l X I / / / l 3 5 . 0 0 0

2 * G O T l n • 3 6 . 0 0 0

2- i 4 WRI r-: ( 6 , 2 0 " 4 i " ( T F ( T | ' , 1= 173"! , T F T 3 7 7 0 0 0 "

3 0 2 0 4 F C R I A T I 1 • , 3 X , • 1 • , 6 X, • 2 • , 6 X ,' 3 • , 5 X , • S U M • / / • T . F ' , 4 ( F 6 . 1 , 1 X 1 / / / ) 3 8 . 0 0 0

C T H I S S E C T I O N G I V E S P E R C E N T P E R S E C T I O N 3 9 . 0 0 0

3 1 5 0 0 S 1 = 1 , 4 " 4 0 . 0 0 0

3 2 MEA J( I ) = ( WC 0 0 ( 2 * 1 - 1 ) * WOOD ( 2 * 1 ) I / WOODT 4 1 . 0 0 0

3 3 M>'A J( I * 6 I = < R A R K ( 2 * 1 - 1 ) V P A R K ( 2 * I I l / B A P K T 4 2 . 0 0 0 " 3 4 6 C ' l N T I N U E 4"37~0"0"0~

3 5 M H A I l 5 ) = W r i n O ( 9 ) /WOODT 4 4 . OOC

3 6 Md-V4-( t ) = w r , 0 1 ( 1 0 1 / w n c i D T _ 4 5 . 0 0 0

3 7 " I L V I I l l l = 3 A R < ( 9 l / B A R K T " - - - - ; 4 6 . 0 0 0

3 3 M E A N l 1 2 1 =1'ARK I I 0 ) / B A R K T 4 7 . 0 0 0

C M E A N g.J't 1 3 A N O 1 4 (_QR 1, P.R 4 ) = 1 . 0 4 R . 0 0 0

3 9 '1 •;A "•)("! 3TÊ^TnT4"r="r7u : 4~<?7iOTJ"a-

C T H I S S E C T I O N C A L C U L A T E S T H E P E R C E N T PER S E C T I O N F O R TWIG,FOL,OR T + F 5 0 . 0 0 0

C I T U S P S A F L A G W H I C H , I F E Q U A L T O 1 , M E A N S T 8 E T R E E WAS S A M P L E D F O R T H E S E 5 1 . 0 0 0

' " ' " C S P C C I - I C \ f I ON 5 • I F F L A G = 0 , TREE ' ")CI.E "CNL'Y 'WAS ' SAMPLED "SO' MEANSOVO " 5 2 . 0 0 0

• E X T E N S I O N * O T H E R C O * ° I L E R S M A Y N O T A L L O W M U L T I P L E A S S I G N M E N T S T A T E M E N T S

4 0 F L A M L 1 = 1 5 3 . 0 0 0

•"41" ~ " I F ( -,.-> = C IF. ( " L 1 7 F 0 7 " 2 7 0 R . ~ S P E C " T E T L " l T r O T 3 T T ; D T T l 7 : 5 4 T 0 0 0 ,

4 2 [ R ( T f , !: 0 . 0 . 0 , 0"- • F T.» E - 0 . 0 . 0 ) GU TOR 5 5 . 0 0 0 . \ - > )

4 3 - 0 0 » 1 = 1 5 , 1 7 5 6 . 0 0 0 * 'i K j

4 4 Mr-A |( I | = T ( !-! 41 / T T ' " . " ' 5 7 . 0 0 0 " " ' - " " " ' I

4 5 9 C O N T I N U r 5 P. 0 0 0 ; |

4 6 0 0 10 1= 1 8 , 2 0 • ' " • . 5 9 . 0 0 0 ' J

http://ii-.ii

c 47 MEAN! I l = F< 1-17) /FT 60. 000 \

48 CONTINUE 61.000 49 GOTO!1 62.000 50 8 FLAJ(LI=0 63. 000 51 DO 12 1=15,20 64. 000 52 MEAN! I 1 = 0.0 • 6 5 . 000 J

> 53 12 CONTINUE • 66. 000 \

54 GOT .111 67. 000 55 7 I Ft T=T .E0 .O . 01GOT03 6 8 . 0 0 0 56 DO 13 1=15,17 6 9 . 0 0 0 57 MEAN! I l = TF( 1-14 l/TFT 70.000 58 13 CONTI NUE 7 1 . 0 0 0

C THIS SECTION WRITES THE PERCENT PER SECTION 72. 000 59 11 WRITE ( 6 , 2 051 73. 000 60 205 FORMAT!•0','PERCENT PER SECTION') 74. 000 61 WRITE( 6 , 2 0 6 1 TREE!L).SPEC IE(L),DBH!LI,FL AG(L) 75. 000 62 206 FORMAT!' ','TREE 13,5X,•SPECIE ',I1,5X,'0BH • ,F5.3,5X,•FLAG=•, 7 6 . 0 0 0

111//) 77 . 000 63 WRI TE 16, 207)(PEAN! I ), 1=1,14) 78. 000 64 207 FORMA T( ' • , 7 X , • U2' , 4 X , ' 3 £ 4 ' ,4X, ' 5C6' ,4 X, '7te' , 5X ,'9' ,5X,« 10'// 7 9 . 0 0 0

l 'WJOO ' . 6 I F 6 . 3 , lXI/ ' B A R K • , 6 ! F6.3, 1X1//'BR1 • , F6. 3 , 3X, ' RR4 ', 8 0 . 0 0 0 1 Ft. 3/1 . 8 1 . 000

65 IF! S'ECIEIL ). EQ.2.OP. SPEC IE! L ).EQ. 31G0T014 82.000 66 WRITE ( 6 , 2081 (ME AN ( I 1, 1 = 15,20) 8 3 . 0 0 0 67 208 FORiATP ' ,8X ,' 1 • , 6 X , '2 • , 6 X , '3'//'TWIGS ' , 3 ( F6. 3 ,1 X1 / ' FOL ', 6 4 . 0 0 0

1 3 ( P 6. 3 , 1 X 1 / 1 8 5 . 0 0 0 68 GUT )!. 5 e6.000 69 14 WRI TEI 6 ,209)1 ME AN!I 1,1=15,171 87. 000 70 209 FORMAT!' •, 3X, ' 1' ,6X, ' 2 ' , 6 X , ' 3'//• T + F •, 3 ( F 6 . 3 ,1 X ) / ) 88.000

C RE»D IN.OF NUTRIENT CONTENT DATA FROM PEPNUT FILES INTO ARRAY NUTCONI20,51 89. 000 71 15 REAOI1,104)TREE1,SPECI1,OBH1 9 0 . 0 0 0 72 104 FGSM\T(I 3 ,11,F 5 . 3I 91 . 0 0 0

C CHECK TO SEE IF NUTCCN TREE SAME AS BIOMASS TREE 9 2 . 0 0 0 73 IF! R E E l L I . E Q . T R E E 1 IGO TO 16 93.000 74 WRITE(6,210ITREE(L),TREE1 94.000 75 210 FCRIATI'l','TREES 3EING PROCESSED NOT S AM E ' / • BI OM ASS TREE=',I3/ 95. 000

1 ' NIR 13NT TREE= • , 13 1 9fc.000 76 RETURN 9 7 . 0 0 0 77 16 DO 17 J = l,5 - 9 8 . 0 0 0 78 READ!1,1051(NUTCON!I,J),I=1,121 9 9 . 0 0 0 79 105 FCRMATU2F5.3I -• 100 .000 80 17 CCNTI NUE 101 .000 81 NUM3£R=20 102.000 82 IFI SPECIE! LI. EQ.2-.0R.SPECIECL 1. EQ. 3 )NUMBER = 17 103. 000 83 DO 13 J=l,5 104.000 84 "REAOI1,106)(NUTCON!I,J),1=13,NUMBERI 105 .000

. 8 5 106 • FnRM\T(3F5.3) 106. 000 • 86 18. • CCNTI NUE ; • * • . ' " • • • • . , . 107.000

C THIS SECTION WRITES THE NUTCON DATA . 103 .000 87 WRI r. (6,2111 1 09 . 000 83 211 FOR •IA T 1 ' - •, ' NUTRI EN T CONTENT DATA'/) 110 .000 89 " " WRIT1(6,2121TREE1,SPECI1,03H1,FLAG(L1 1 1 1 . 0 0 0 90 212 FCRMUC ' , ' T R E E •, 13 , 5 X , « SPECIE ',I1,5X , 'DBH « , F 5. 3, 5X, • FL AG= ' , 112. 000

111//) 113.000 91 WRIT;i6,213> J 1 4 . 0 0 0 92 213 . . F'IRMA T( • • ,T ! 6, •*• , 15 ( •-• ) , ' WOOD** SECT I ON • ,15 (•-• J , ' *• , T75, •*•, 1 1 5 . 0 0 0

1151 •-' I, 'OAR K** SECT I 0 N • , 1 5 ( • - • ) , ' * • / , ' ,21 16X,'l',7Xi'3" ,7X,'5' , 116 .000 1 7 <, < 7 • ,7X , ' 9 ' , 7 X , ' 1 0 ' ,1X1/1 ' " 117. 000 ~ "

93 .DO 19 J=l,5 " 118.000 94 WRI T 1 6 , 2 1 4 ) J , 1 NUTCON! I, J ) ,1 = 1,12) : 119 .000

95 214 FORMAT!' ', 11 , • - < ,9 X, 6! 2 X, F6. 31 ,11 X, 61 2 X, F6. 3 II 120.000 96 19 CONTINUE 121.000 97 IF(SPECIEIL). EQ.2.0R. SPECIEIL I. EO. 31 GOT 02 0 122.000 93 WRIT:(6,21.51 123.000 99 215 FORMAT (•-• ,T1 3, 2( • BR ANC H • I , 3X, • * •, 7( '-• I , • TWI GS ' , 7( • - • I , •« • , 3X, 124.000

1 ' «' ,S ( '-' I ,' FCL IAGE ' ,7C-' I . • *'/' ' , T1 7 ,' >1 1 , 5X ,' 1-1 /4', 125. 000 1 2 ( 3<, • ( TOP I ' , 3X , • IMIi) 1 •, 3X, ' ( LOWI • I/) 126.000

100 00 21 J = l , 5 127.000 101 _____ ; WRI H I 6, 2161 J , ( NUTCON ( I, J I ,1=13,201 : 128.000 102 216 FORMAT!' • , 11 , • - • , 9 X, 8 (2 X, F6.31 1 ' "129.000 103 21 •'• CONTINUE 130.000 104 G IT 122 _____ . ; 131. 000 105 20 WRITE(6,217) 132.000 106 217 FC=H\T (• -• , Tl 3,2 ( • BRANCH'I,3X,'*',3('-' 1,'TWIGS + FOLIAGE *'/ 133. 000

1' '. Tl 7, •>!•, 5X ,'1-1/4', 3X, •( TOP) • ,3X, • (MIDI ' ,3X, •( LOW) •/) 134.000 107 00 2, J = l,5 " 135. 000 103 WRI TE ( 6, 213) J , ( NUTCON (I,J ) ,1 = 13,171 136. 000 109 218 • FORMAT!' ' . 11 , ' - ' , 9 X, 5 [ 2X , F6. 31 I 13 7. OOP 110 23 CONTINUE 1 38. 000'

C THIS SECTION CALCULATES THE WEIGHTED CONCENTRATIONS (WEIGHTED MEAN X CONC.I 139.000 1U __22 NUM>-P. = 20 _ _ . 140.000 112" " fF"VSP'=ClElLi.E0i'2\OR.SPECl'E(LI.EQ.3'lNUMBER = f7 141.000" 113 00 24 .1=1,5 142.000 114 00 >> 1 = 1, NUMBER ; 143. 00 0 115 NUTCUN (,I ,J)=NUTCON! I, JI * ME AN( I I 144. 000 116 25 ' . CONTINUE 145. 000 l_7 24_ _CONriN'JE . • . - 146.000 ~ : C THIS .'SECTION WRITES" THE WEIGHTEO CONCE NT RAT "IONS-; : ' 14 7. 000" 113. WRITS (6,2191 > 143. 000 1]Q 219 FORMAT!' 1','WEIGHTED-CONCENTRATIONS'I 1 49. 000 120 . WRI f i ( 6,2121 TREE1.SPECI1, DBH1 , E L AG ( L I ~ 1'5"0.000 121 . WRITE!6,2131 ]51.000 122 DO 26 J = l,5__- _ _ _• 1 52. 000 123 WRI TS ( 6, 2141 j,"( NUTCON ( I i J I »1 -1»1 2T '. : 1 53. 000 124 26 CONTINUE 1 54. 000 125 J_F( >*rS:l.E 1 L » ' E0.2.HR. SPEC IE(L I. EQ. 3) G0T027 1 55. 000 126 WRI'Tc ( 6, 2T'5T7~ ~ ~ : " i 156.000 127 00 23 J=l,5 ' .:-"' -' 157.000 129_ WRI Ti 16,216)Jj,(NUTC0NJI,J^,1 = 13,20) 158.000 129" 28 "CONTINUE ' "" "" " ' ' ] 15^.000 130 G0T029 160.000 131 27 WRl IE ( S,21 7J 161.000 132 DO 11 J = 1T5 ~~ : T62.00TJ 133 WRITE(6,2131J,(NUTCON(I,J 1,1 = 13, 171 163.000 134 30 CONTINUE 164.000

Cn,HIS'"SECri0N~SUMS~W30D7"~BSRK7""TffI"GS7""F"0L; C1R TWTCS+FOL SEPARATELY 165. 000 135 29 00 Jl 1 = 1 ,6 166.000 136 DO 32 J = l ,5 . . . ' 167. 000

TV? N j T s i m i , J l = 3.0 ' ~ : TE"8TTJ0r0 139 32 CONTINUE 169. 000 139 31 CONTINUE 170.000 !V0"" ~00*'33 J=175 " 171.000"" 14! DO 34 1 = 1 ,6 1 72.000 142 NUT S J M! 1 , J|=NUTSUM( 1, Jt+NUTCON! 1 , J I 173.000

T41 TTOT SIMT27J IÎUrSTfMI 2, J )•NUTCON! I +6 , J I T74T0D0 144 34 CONTINUE 1 75. 000 145 NUT 5 IM( ? ,.I) = NUTSUM( 3, J 1 + NUTCO N( 13 , J) 176.000 _<,_-- • - : NUT ;JM(4,J1 = NUT SUM!4,JI*NUTCON(14» JI T77.000' 147 33 CONTINUE 178.000 148 -\Ft S3ECIE(L I.E0.2.0P. SPEC IEIL I.EQ. 3IG0T035 179.000

f 149 150 151 152

DO 36 J = l,5 DO 37 1 = 15,17 NUT3JMI 5,J) = NUTSUM(5,Jl+NUTCGN!I,J) NUTSJMI6,J|=NUTSUM(6,J>*NUTC0N(I+3,JI

. 180.000 181.000 182.000 183. 000

153 37 CONTINUE 184.000

V—-—- 154 36 CONTINUE 185. 0 0 0 f 155 . GOT 13 3 1 8 6 . 0 0 0 : <

15& 157

35 • DO 3 9 J=l,5 00 4 0 1=15,17

187.000 188.000

15B NUT i i M ( 5 , J 1 = Ni)T SUM! 5 , J I + NUTCON! I , J ) 189. 0 0 0 159 40 CfNTINUE 190.000 160 39 CONTINUE 191. 0 0 0

C THIS SECTION GIVES THE TREE A DBH CLASS:CLASS ES 1-11 = 6-10",10-14",. ..,46"- 1 192.000 161 162

38 )=O.H(L> CALL CL\SSR.(n,M)

193.000 194. 0 0 0

163 CLA~,3(Lt = M 195.000 C THIS SECTION WRITES THE SUMMED WEIGHTED CONCENTRATIONS 196. 0 0 0

164 WRI TE ( 6, 2201 1 97. 0 0 0

165 220 .FORMAT!'-",'SUMMED WEIGHTED CONCENTRATIONS BY SPECIFICATIONS') 198.000 166 WRI TF. (6,221 ) TREE1 ,SPECU ,DBH1 , FLAG! L ), CLASS (L ) 199.000 -167 _221.__ F O R UTI '0' , 'TREE • , 1 3 , 5X,'SPECIE ' »11,5X,'DBH • ,F5.3 ,5X,•FLAG=' 200. 000

i l l , 3 X , ' O B H CLASS=', Ji/7) 201.000 163 I F ( JPFCI E( L) . E0.2.0R. SPEC IE(L ) . EO. 3 1 G0T04 1 202.000 169 WRITE(6,2;2) 203. 000 170 222 FGRU'TC ",4X,'waOO ",3X,'8ARK ' , 3 X , " BR 1 " , 3 X , ' BR4 ',3X, 204.000

171 1•TW13S',3X,• FOL • / 1 DO 4 2 J=1 , 5

205. 000 206.000

172 WRITE(6, 223 1 J,INUTSUM(I,J 1 , 1 = 1,6) 207. 000 173 223 FOR n i l " ",I1, •- • , 6 ( F 5 . 3 , 3 X I ) . 2Ce. 000 174 42 • CONTINUE 209.000 175 176 41

GOT 14 J

. WRI T r ( 6 , 2 2 4 ) 210. 000 211. 000

177 224 FORMAT!• " , 4 X , ' W 0 0 0 ",3X,"BARK ',3X,' BR1 ',3X,' BR4 ',3X, 212.000

• 178 1 ' T*F •/ I DO. 4 4 J = l , 5

"" 213.000 214.000

179 WRI TE ( 6, 2251J, < NUTSUM < I, J I , 1 = 1,5) 215.000 190 225 FORMAT(• ',11,'- ',5(F5.3,3XII 216.000 181 , 4 4 CONTI NUE ••• ' • 217.000

C THIS SECTION i>UTS THE ARRAY NUTSUM INTO A FINAL ARRAY INSUM1 FOR ALL TREES 218.000 182 4 3 DO ,b • J=l,5 ""219.000 ie3 0 0 4 6 .1 = 1 ,6 220. 000 194 NSJM I , J , L ) = NUTSUM( I , J) 221.000

. 185 46 CONTINUE 222.000 186 45 CONTINUE 223.000 187 1 CONTINUE 224.000

C END O F PROCESSING FOR THE N TREES "" 225. 000 C INFORMATION IS IN THE F OL LOW I NG ARRAYS : TREE! L 1 , SPECI E ( L 1 , OBH(L), ClASSILI, 226.000 C FLAG IL1, NSUM!I,J,L); L = TP EE PROCESSING ORDER; I=1-6(SPECIFICAT IONSI; 227.000 C J=l -5INJTRIENTS1; CLASS IS ONE OF 11 DBH CLASSES; FLAG=1 IF TPEE COMPLETELY 22H.O0O C SAMPLED, F L A 3 = 0 IF TREE BOLE ONLY SAMPLED 229.000

188 WRITE(t>, 226> 230.000 189 226 FORMAT('1'1• 231. 000 190 WRITE(6,2201 232. 000 191 DO 5 0 L=1,N 233. 000 192 WRITE(o,221ITREE(L),SPECIEIL) ,OtiH( L) , FL AG ( L 1 , CL AS S ( L ) 234.000 193 IF(S»ECIEIL).EQ.2.0R.SPEC IE(LI.EQ.3IG0TO51 235.000 194 WRITE(6,222) 236.000 195 DO 52 J = 1,5 ~ ' " 237.000 196 WRITE(6,2 23)J,<NSUH(I,J,L),I=1,6) 238.000 197 52 CONTINUE 239. 000

o

o

I o

•9

3

•J

r 198 GQT05 3 2 4 0 . 0 0 0 \

199 51 WRITE(6 ,224) 2 4 1 . 0 0 0 200 DO 54 J = l , 5 242 . 000 201 WR!TE(6 , 2 2 5 I J , ( N S U M ( I , J , L ) . I = 1 . 5 I 2 4 3 . 0 0 0 202 54 CONTI NUE 2 44. 000

\ 203 5 3 CONTINUE 2 4 5 . 000 ) 204 50 CONTINUE ' . 2 4 6 . 0 0 0 < 205 WRITE(6,2271 2 4 7 . 000 206 227 f n r U T I ' l ' l 2 4 8 . 0 0 0

C PUT AR R A / NSUM ON F ILE 2 4 9 . 0 0 0 207 DO 55 L=l ,N 250. 000 203 WRI TE ( 2 . 2 50) TRtE (L) .SPEC IE (LI ,OBH( L) ,F L AG ( L ) , CL AS S ( L 1 2 5 1 . 0 0 0 209 250 FORMAT(I 3 , 1 1 , F 5 . 3 , 1 1 , 111 2 5 2 . 0 0 0 210 DO 56 J = l , 5 2 5 3 . 000 211 WRI TE ( 2 , 251 ) ( NSUM( I , J , L ) , 1 =1,61 2 5 4 . 0 0 0 212 251 FOR Mi T(6 F5•3 I 2 5 5 . 0 0 0 213 56 CONTINUE 256. 000 214 55 CONTI NUE 2 5 7 . 0 0 0 215 RET H ) 25B. 000 216 E NO 2 5 9 . 0 0 0

217 SUBROUTINE CLASSR(DBH,CLASS 1 2 6 0 . 0 0 0 C SUBROUTINE TO CLASSIFY OBH INTO ONE OF 11 G R O U P S ( 6 - 1 0 " , 1 0 - 1 4 " , . . . , 4 2 - 4 6 " , 46" -1 2 6 1 . 0 0 0

218 ' REAL TBH.C/O. 0254/ 2 6 2 . 0 0 0 219 I NT;jER C L A S S , J 2 6 3 . 0 0 0 220 CLASS =1 2 6 4 . 0 0 0 221 J=6 2 6 5 . 000 22 2 IF0J3M.LT. ( J = CI IRETURN 2 6 6 . 000

; ; . 2 2 3 . on i 1 = 1 , 1 0 2 6 7 . 0 0 0 224 CLASS = 1 2 6 8 . 000

. 225 IF( J H.GE . I J * C I . ANO. DBH.LT. (( J + 4I*CI IGOT02 269.COO 22 6 J = J*4 2 7 0 . 000 22 7 1 • ' CONTI NUE 2 7 1 . 000 22 8 CLA3S=11 2 7 2 . 0 0 0 229 2 RETURN 2 7 3 . 0 0 0 230 FN ) 2 74. 000

$OATA 2 7 5 . 0 0 0

L_

A P P E N D I X 13

ROGRAM FOR C A L C U L A T I N G BIOMASS FOR 2 5 6 T R E E S

ON THE SQUAMISH SAMPLE PLOT ACCORDING TO

L O G A R I T H M I C R E G R E S S I O N EQUATIONS

SCOMPILE C BIOMASS DISTRIBUTION FOR 256 TREES FROM C NEAR SQUAMISH

1 CHARACTER»3 TRE

HIGH ELEVATION SAMPLE PLOT

2 I N T E G E R SP 3 R E A L L

_ _ a HW00D=H8ARKsHDwBl »HDWBt|aHTWJJ3«HFOL IBHTWFPSHSTRU • E X T E N S I O N * OTHER C O M P I L E R S MAV NOT ALLOW M U L T I P L E ASSIGNMENT • E X T E N S I O N * OTHER C O M P I L E R S MAY NOT ALLOW M U L T I P L E ASSIGNMENT • E X T E N S I O N * OTHER C O M P I L E R S MAY NOT ALLOW M U L T I P L E ASSIGNMENT

"H3CRO*H3TRE"0,0 STATEMENTS STATEMENTS STATEMENTS

• E X T E N S I O N * • E X T E N S I O N * • E X T E N S I O N * • E X T E N S I O N * • E X T E N S I O N * • E X T E N S I O N *

5 • E X T E N S I O N * • E X T E N S I O N * • E X T E N S I O N * • E X T E N S I O N *

_ * E X T E N S I O N « _ • E X T E N S I O N * • E X T E N S I O N * • E X T E N S I O N *

' " E X T E N S I O N * 6 7

OTHER COMPILERS MAY NOT ALLOW MULTIPLE ASSIGNMENT STATEMENTS OTHER COMPILERS MAY NOT ALLOW MULTIPLE ASSIGNMENT STATEMENTS OTHER COMPILERS MAY NOTJLLOW MULTIPLE ASSIGNMENT STATEMENTS OTHER COMPILERS MAY"NOT ALLOW-MULT I PLE ASSIGNMENT"ST A TtMENT S OTHER COMPILER3 MAY NOT ALLOW MULTIPLE ASSIGNMENT STATEMENTS OTHER COMPILERS MAY NOT ALLOW MULTIPLE ASSIGNMENT STATEMENTS

AwOOD = ABARK = AnwHl«ADwBllBATWIG3AFOLI«AT»«FO = ASTHU: OTHER COMPILERS MAY NOT ALLOW MULTIPLE ASSIGNMENT OTHER COMPILERS MAY NOT ALLOW MULTIPLE ASSIGNMENT OTHER COMPILERS MAY NOT ALLOW MULT IPLE ASSIGNMENT OTHER COMPILERS MAY NOT ALLOW MULTIPLE ASSIGNMENT

_OTHER COMPILERS MAY NOT ALLOW MULTIPLE ASSIGNMENT 0fH£R COMPILERS MAY NOT ALLOW MULTIPLE ASSIGNMENT OTHER COMPILERS MAY NOT ALLOW MULTIPLE ASSIGNMENT OTHER COMPILERS MAY NOT ALLOW MULTIPLEASSIGNMENT OTHER COMPILERS MAY NOT ALLOW MULTIPLE ASSIGNMENT

ILINEsO DO 1 1*1,256

A"SCH"O = ASTHE»O70r STATEMENTS STATEMENTS STATEMENTS STATEMENTS STATEMENTS STAT EMtNTS STATEMENTS STATEMENTS STATEMENTS

8 9

10 1 i " 12 13 TC" 15

"16"" 17 16 T»~ 20 21

"22 23 2a ~?:v 26 27 28 29 30

" J l " 32 33

"3<r 35 36

"37" 38

HE AD(6,11) TRE,SP,0,H,BUTL,CW1,CW2 11 FORMAT (A3, I I , 3<F4, 1),4X,F2,0,F2,0>

Cws(CC*UCW2)/2,01*0,3048 _ _ " " 0*6,0254*0 "

CL*H"BUTL BAa(3,H159«0«D) /a,0 • ' IFCSP.EQ . i ) GO TO 21 " ~ ~ ~ IF (SP.EQ.2) GO TO 22

C 8PI«HEML0CK,SP2=ABIFS "21""" B W 0 0 D s 2, 319 5 • 0 ,7 4 5 5 * A L"0G10 (D"«D *"H) ""

BBARK33,1089*1,0388«AL0G10(BA) BDWB1=0.6467+0.7853*ALOG10(D*0*H*CW) "D'WB«s I,90'ii6 + 0,2089*(BA»H*CW*CLJ ' BTWFOaO,8851+l.5576*ALOG10(D*0*M)*0, BTWIG=-1.2724*1,1630«ALOG10(D*CL*CL) BFOLI=0,5980*|,7174*ALOGI0(D*D»H)»0, WOODslO,0«*Bk.OOD BARK = 10,0»*B!IARK "DwB'l = 1 0", 0 •"•BDWB'l :

TwFO=iO,0«*BTWFO TWIG=IO,0*»RT*IG P O L I " 1 0 , 0 * » 3 F O L I 7 " STRU=WOOO*BARK SCROiOoBl *nWH4*TWFO ' """' S T H E = 3 T R l J t S C R ' 0 : ~ HnOOD=nWOOD+"000 .HBARKSHBAKK+HARK HDwBlsHDwBl+DWfll ' HDWB4 = H0».B4 + DWB4 HT.WIG = HTMG + TWIG

"TIF'OC'rBHFOLIVro'CI HTWFOsHUFO + TWFO

0362*D*D*H

0U47*D«D*H

5.000 6,000 7,000 8,000 9,000

10.000 11.000

12,000

13,000 14.000 15,000 16,000 17,000_

"18,000 19.000

_20 , 000_ ?:\~, ooo 22,000 23,000 24,000" 25,000 26,000

"27"; 0 00"" 28,000 29,000 30.000 31,000 32.000

"3370 0 0" 34.000 35.000

"36.000" 37,000 38,000

"3'9700"0"" 40.000 41.000 "42 , 000 43,000 44.000

"457000" 46,000

39 40 41 42

H3TRU»HSTRU*3TRU HSCR03HSCRO+3CRO HSTREsHSTRE+STRE GO TO 29

4 7 , 0 0 0 4 8 , 0 0 0 4 9 , 0 0 0

/ 43 2 2 BW00D=2,0472*0,9S26*AL0G10(D*D«H) 51,000 s 44 BBARK=3,0965+1,3268*ALOG10(BA) 52,000 45 B0*B1 = 1 , 2 6 8 8 * 0 ,0525*0 * C L 53.000 © 46 BDv.B4 = 0.6615 + 0,7597*ALOG10(D*CL) 54,000

© 47 BT»F0=0,8787+1,0376*ALOG10(D »CL ) 55,000 48 WOOD=10,0**BwOOD 56,000 49 BARK=10,0**RHARK 57,000 SO DHB1*10,0*»BDWB1 5B.OO0 51 D*HUal0.0*»BDwB4 59.000 © 52 T*FO=10,0*«BTWFO 60,000 53 TWIGaO.O 61,000 54 FOLI=0,0 62,000 o 5 5 STRUsWOOD+BARK 63.000 o 56 SCROsQ».Fil+DWB4 + TWF0 64.000 57 STRE=STRU+3CR0 65,000 o 5 8 A*0O0=AK0OD+w0OD 66,000

o 5 9 A8ARK=A8ARK+BARK 67,000 60 ADwBlsAOWfll+OWBl 66.000 © 61 AD*B4aADI»B4*DWB4 69,000

© 62 ATWIGSATKIG+TWIG 70,000 63 AFOLIaAFOLI+FOLl 71.000 o 64 ATWFOsAUFO + TWFO 72,000 65 A3TRU=ASTRU+STRU 73.000 66 ASCRO=ASCRO+SCRO 74,000 o 67 A3THESASTRE+3TRE 75.000 o 68 GO TO 29 76.000

C PRINT AUT RESULTS FOR INDIVIDUAL TREE3 77,000 o 69 29 I F d . E U . l ) GO TO 3i 78,000 70 GO TO 33 79,000 71 31 WRITEC6.32) 80,000 o 72 ~~32 F ORHAT (11', 'TREE 1X, < SP» , IX,'DBH !,?)(,'LENGTH l , lX,"CRLEN6THl, IX, 61,000

1"CRD IAl,4X,tWOOD',OX,'BARK',2X,'BIG BRAN',1X, '3 M BRAN',2X, ' TWIG', 82,000 23X, 'FOLIAGE', IX, 'TW+FOL' ,2X, 'TREE BOLE',2X,'CROWN',4X,'SUM TREE'/ 63,000 o 3T10, ' (M) ' ,T16, ' (M) ' ,T23, ' (M) ' ,T29, ' (M) ' ,T37 i ' (KG) 'VT4S, ' (KG) '",T53» 84,000 4 ' ( K G ) ' , T 6 2 , ' (KG) ' , T 6 9 , ' ( K G ) ' , T 7 7 , • ( K G ) ' , T 8 5 , ' C K G ) ' , T 9 4 , I ( K G ) ' , 85,000 ST104, ' (KG)•,T114, ' (KG) ' / / ) 86,000 o

73 33 WRITE (6,34) TRE, SP, D, H, C L , CW, WOOD"; BARK , DWB1, DWU4, T w IG, F O L I , T WFO, 8 7,000 o

1STRU,SCR0,STRE 88,000 74 34 FORMAT ( ' 0 ' , A3 ,2X , I I , 1X ,F5 ,3 ,3 (2X , F 4 , 1 ) , 3X ,7 ( F B , 1 ) , 3 (F10 ,1 ) ) 89,000 tf 75 ILINEsILINE+1 90,000 76 I F ( I L I N E , E 0 , 2 7 ) GO TO 40 91 ,000 77 UO TO 1 92.000 7B 40 WRITfcl6,3<iJ 93,000 79 ILINE=0 94,000 80 1 CONTINUE 95,000 81 T>OOD = HWOOD + AWOOD 96,000 8 2 TBARKSHBARK+ABARK 97,000 83 TOWBlaHD»Bl+ADWBl 98,000 8 4 T0*BUaHD>.B4 + ADWB4 99,000 8 5 ' TTWIGaHUIG + ATWlG 100,000 86 TFOLI=HFOLItAFOLI 101,000 87 TTxF0=HT«F0+AT«F0 1 02 , 000 """ ' " 6 8 T3TRU=HSTRU+ASTRU 103,000 8 9 TSCROsHSCRO+ASCRO 104,000 J o 9 0 TSTRE=HSIRL+ASTKf 105,000

C PRINT OUT SUMARY FOR SAMPLE PLOT 106,000

91 NRITE<6,35) HWOOD,HBARK,HDWB1,HOWB«,HTWIG,HPOLI»HTWFO,HSTRU,HSCRO, 107,000 1HSTRE»AW00D,ABARK,A0WB1, ADWB4, ATWIG, AFOU , ATWFO, ASTRU, ASCRO, A3TRE# 108,00 0 2T*000,TBARK,TDWB1,TDW8«,TTWIG,TFOLI,TTWFO,TSTRU,TSCRO,T3TRE 109,000

<2 35 F ORMATC///j'/,'BIOÂSSjFHEMJ.OC K_ TR EE SJ _T 35,7 tF 8,1 )__3__F 1 0,1)_// 11 0,000 1 "BIOMASS CF ABlE3~fREES~t,"T3~5,7{F^,Or3(F10,l)//V/ISUM OF BIOMASS F 111 ,060 20R STAND',T35,7(F8,l),3CF10,11) 112,000

93 WRITE(<>,36) ; 113.000 94 36 " "FORMAT (' 1 • > " " - - i i i , 0 0 0 95 STOP 115.000 96 END ' 116,000

SDATA 117,000

BIOMASS AND NUTRIENT DISTRIBUTION IN TWO OLD ... - Climate

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