CONTRIBUTION TO THE STUDY OF THE FLORA OF ATTICA …institucional.us.es/revistas/lagascalia/17.2/03 sarlis.pdfCONTRIBUTION TO THE STUDY OF THE FLORA OF ATTICA (GREECE) G. P. SARLIS

Lagascalia 17(2): 229-256 (1994)

CONTRIBUTION TO THE STUDY OF THE FLORAOF ATTICA (GREECE)

G. P. SARLISInstitute of Systematic Botany, Agricultural University of Athens, Iera Odos

75, 118 85 Athens, Greece(Recibido el 6 de Abril de 1992)

Resumen. Se estudia la flora de la cuenca de Atenas y de las montañas que la ro-dean. Se trata de una flora extraordinariamente diversa, compuesta por 1144 taxones,pertenecientes a 1084 especies, que representan a 490 géneros y 102 familias. Los com-ponentes principales de esta flora son especies pratenses, medicinales, aromáticas yapícolas. Muchas de estas especies se encuentran amenazadas o se han extinguido yaen la zona de estudio.

Summary. A study of the flora of the Athens basin and the surrounding mountainsis presented. The results of our work show that this flora is exeptionally diverse andconsists of 1144 taxa belonging to 1084 species, 490 genera and 102 families.Components of this flora include many pasture and forage species, along with medici-nal, aromatic and apicultural species. It is documented that many of these importantspecies are threatened or have already become extinct.

INTRODUCTION

Athens basin, where the city of Athens (capital of Greece) is built, is inAttica (Fig. 1). This basin (an area of 383 sq. Km) is the biggest of the fourplanes in Attica. At its S and SW side the basin is open to Saronikos Gulf andthe rest is surrounded by mountains, some of them with considerable altimetry.Mount Parnes coVers the N side of the basin, mount Penteliko the NW, mountHymettus the E and SE and mount Egaleo the W and SW side. The basin isdivided in Eastern and the Western parts, by a chain of hills consisted ofLycabettus, Acropolis, Philopappou and some lower hills.

The flora and the vegetation of the Athens basin and the mountains thatsurrounds it, was and it is a subject of great interest between botanists all overthe world. Many botanists visited Attica until 1940 and published their studies,

230 Contribution ro the study of the flora ofAttica (Greece)

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3 Philopappou

5 Lycabettus

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Fig. 1. Map of Attica

including Sibthorp, Spruner, Fraas, Heldreich, Holzmann, Halascy, Adamovic,Rikli, Rubel, Mattfeld, Guiol, Stoyanoff, Jordanoff, Forbes and Rechinger(PAPANIKOLAOU & SARLIS, 1991).

Geology, Soil, Climate

The geological base of the basin is consisted of Mesozoic (gray limestone,arilaceous shist, limestone mar) and Cenozoic (argilaceous and marlysediments, red loam, mar!, red earths) layers (LEPSiuS, 1893).

The soil types of the basin are mostly, limestone and marly rendzinas,regosols, brown forrestal and mediterranean, alluvial deposits mixted withregosols, and brown-red forrestal of medium depth mixted with sceletonformations (ZVORYKIN & SAUL, 1948).

Lagascalia 17(2), 1994 231

The climate in the basin includes the highest mean minimum temperatures,during May to August, and the lowest relative moisture (44% average for July),in the country. Al! these in connection with the small annual precipitation andthe enclosure from the mountains that surround the basin, establish a special(in a climatic point of view) region of Greece (MAVROMMATIS, 1980).

MATERIALS AND METHODS

For this study, plant-specimens were selected and botanical observation weremade during our visits in each one of the chosen areas, for the period between1980 and 1988. The following areas were covered: Mt. Parnes, Mt. Penteliko,Mt. Hymettus, Mt. Egaleo, Tatoi (National Park), the hills Lycabettus,Acropolis, Philopappou, the park of Daphni and selected spots in fields or nearroads.

Species were identified at the laboratory of the Institute of SystematicBotany in Agricultural University of Athens.

Results were compared with those reffered in the studies of ZERLENTIS

(1959, 1968), DIAPOULIS (1958) and SARLIS (1977, 1980). Using ah l the namesof the identified species, plus the mentioned in the previous studies, we createda computer database. This database consist of 1144 rows and 10 columns. Therows where occupied from the names of the species (according to TUTIN &

al.,1964-1980). The columns were occupied from the names of genus-species,family, order, class, areas, vegetation forms symbols, ecological forms symbols(two columns), rareness symbol and symbol of cultivation.

The Ashton-Tate's «Framework II» package was chosen because of itsfiltering convinience and its compatibility with «Dbase III plus». The databasethat was created, covered total memory of 297 Kbytes.

RESULTS

The diversity of the flora is represented in Table 1 and presentation ofanalytical data of our studies is made in the appendix.

DISCUSSION

The most important element of this study is the presence of many pastureand forage species. This is documented by the presence of 147 taxa belonging

232 Contribution to the study of (he flora of Attica (Greece)

Systematic units Fam. Gen. Spec. Subsp. vars. f. taxa Percentage

Pteridophyta 9 10 12 1 – – 12 1.05Gymnospermae 3 5 10 3 – 2 12 1.05Dicotyledones 79 384 844 80 23 – 899 78.73Monocotyledones 11 91 218 22 2 – 219 19.17Total 102 490 1084 106 25 2 1144 100.00

Table 1. Systematic units and taxa of the flora.

to Leguminosae and 88 taxa belonging to Gramineae. The fact that 26 taxabelong to genus Trifolium, 21 to genus Vicia, 14 to genus Lathyrus and 16 togenus Medicago corroborates this aspect (SARLIS, 1985).

The presence of species important in apiculture is also documented in theappendix, especially whith the many taxa belonging to Labiatae andLeguminosae (SARLIS, 1991).

An element of great importance is also the presence of many aromatic andmedicinal plants. Some of them like Chamomilla recutita, Crocus ssp., Laurusnobilis, Lavandula stoechas, Salvia ssp., Styrax officinalis, Verbena officinalis,Satureja thymbra, Mentha pulegium, Tussilago farfara, Taraxacum officinaleand Thymus capitatus are very common in the studied areas.

REFERENCES

DIAPOULIS, C. (1958) The flora of Parnes. 6: 163-158 (in Greek)LEPSIUS, R. (1893) Geologie von Attica. T.U. Berlin.MAVROMMATIS, G. (1980) Le Bioclimat de la Grece; relations entre le climat et la

vegetation naturelle. Cartes Bioclimatiques. Institut de Recherches Forestieres d'Athenes.

PAPANIKOLAOU, G., & SARLIS, G. (1991) Phytosociological Studies in the Tatoi District(Attica, Greece). Vegetatio 93: 81-90

SARLIS, G., P. (1977) Contribution to the study of the Aromatic and Medicinal Flora ofMount Parnes (Attica). Scient. Studies & Publications: 7-22. Athens (in Greek).

— (1980) The Flora of Mount Egaleo (Attica, Greece). Phyton (Austria) 20, 3-4: 261-278.

— (1985) Range management. Agric. University of Athens, Athens (in Greek).— (1991) Aromatic and Medicinal Plants. Agric. University of Athens, Athens (in

Greek).TUT1N, T.G., & al. (1964-1980) Flora Europaea. 1-5: Cambridge University Press,

Cambridge.ZERLENTIS, K. (1959) The plants of Lycabettus, Bull. Agric. Bank 109: 19-28 (in Greek).— (1968) Contribution to the study of the flora of mount Hymettus, Bull. Agric.

(Minist. Agric.) 19: 5-30. (in Greek).ZVORYKIN, A., I., & SAUL, J., P. (1948) Soil - map of Attica. Scale 1:100,000. Athens.

Lagascalia 17(2), 1994 233

APPENDIX

Floristic, classification and ecologic data. A, Acropolis; B, waste lands nearroads; D, Daphni; E, Egaleo; F, Philopappou; L, Lycabettus; P, Parnes; PN,Penteliko; T, Tatoi; Y, Hymettus. *, cultivated species. (?), species mentionedby Zerlentis (1959, 1968) and Diapoulis (1958) but not found in this study.

SYSTEMATIC UNITS

REA LIFE FORMS

PTERIDOPHYTASelaginellaceae

Selaginella denticulata (L.) Link P-L-F-A-T-Y-PN-E-D B P HreptEquisetaceae

Equisetum telmateia Ehrh. P P GrhE. palustre L.

P-PN P GrhSinopteridaceae

Cheilanthes fragrans (L. fil.) Swartz P-L-Y-E-PN-D-T P GrhHypolepidaceae

Pteridium aquilinum (L.) Kuhn T-P P GrhAdianthaceae

Adiantum capillus-veneris L. P-E-PN-T P GrhAspleniaceae

Asplenium trichomanes L. P-PN-D-T-D P HcaespA. onopteris L. P-PN P HcaespCeterach officinarum DC. P-L-F-A-T Y-PN-E-D P Hcaesp

PolypodiaceaePolypodium vulgare L. P-Y P Grh

AspidiaceaeDryopteris villari Schinz.& Thell. subsp.pallida Heywood

P-PN P GrhAthyriaceae

Cystopteris fragilis (L.) Bemth. P-PN-T-E P HcaespGYMNOSPERMAEPinaceae

Abies cephalonica Loudon P-T Ar MPh*Pinus halepensis Miller P-L-F-A-T-Y-PN-E-D Ar MPh*P. nigra Amold T Ar MPh*P. brutia Ten. T Ar MPh*P. pinea L.

T Ar MPhCupressaceae

*Cupressus sempervirens L. f. sempervirens L-T-Y-E-D-PN-B Ar MPh*C. sempervirens L. f. horizontalis (Miller) Voss. L-T-Y-PN-E-D-B Ar MPhJuniperus communis L. subsp. communis T-PN Fr NPhJ. oxycedrus L. P-T-Y-PN-E-D Fr NPhJ. oxycedrus L. subsp. macrocarpa(Sibth. & Sm.) Ball T-Y-E-D Fr NPhJ. phoenicea L.

P-T-Y-E-D-PN-B Fr NPhEphedraceae

Ephedra fragilis Desf. subsp. campylopoda(Meyer) Ascher. & Graeb. P-L-T-Y-PN-E-D P Chfrut

DICOTYLEDONESSalicaceae

*Salix alba L. P-T-PN-E-D-B As MPh

234 Contribution to the study of the flora of Attica (Greece)

*Populus alba L.*R tremula L.

CorylaceaeCarpinus orientalis MillerOstrya carpinifolia Scop.

FagaceaeQuercus coccifera L.Q. ilex L.*Q. petraea ((Mattuschka) Liebl.Q. macrolepis Kotschy*Q. frainetto Ten.Q. robur L.Q. pubescens Willd.

Ulmaceae*Ulmus minor Miller

Moraceae*Ficus carica L.

UrticaceaeUrtica dioica L.U. urens L.U. pilulifera L.U. membranacea Poir.Parietaria diffusa Mert. & KochP. cretica L.P. lusitanica L. subsp. serbica (Paricic)P.W. Ball

SantalaceaeOsyris alba L.Thesium humile VahlT. bergeri Zucc.Comandra elegans Reinhenb fil.

LoranthaceaeViscum laxum Boiss. & Reuter var. abietis

AristolochiaceaeAristolochia microstoma L.A. longa L.Asarum europaeum L.

CytinaceaeCytinus hypocistis (L.) L.C. hypocistis (L.) L. subsp. macranthus Wettst.

PolygonaceaePolygonum arenarium Waldst. & Kit.P. aviculare L.Rumex acetosella L.R. acetosa L.R. conglomeratus MurrayR. pulcher L.R. crispus L.R. bucephalophorus L.Atraphaxis billardieri Jaub. & Spach.

ChenopodiaceaeBeta vulgaris L. subsp. maritima (L.) ArcangeliChenopodium bonus-henricus L.Ch. vulvaria L.Ch. murale L.Ch. album L.

PN-T-D-BP-PN

T-PNT-P

P-L-F-A-T-Y-PN-E-D-BP-Y-PN-T-E-DT-PNP-T

PNP-T-PN

F-A-T-Y-PN-E-B

P-F-A-B-D-T-PN-EF-A-B-D-T-Y-PN-EP-PN-L-F-A-B-D-Y-E-TL-PN-E-T-BL-F-A-B-D-T-PN-E-YL-Y-E-B-TY-E-B-T

P-D-T-Y-PN-EY-E-DP-Y-E-DP(?)

Y-E-PN-D-BP-PN-Y-D-T-BP-PN

P-Y-T-PN

F-A-D-PN-E-TL-F-A-B-T-Y-EP-PN-F-A-T-EP-PN-T-DF-A-T-PN-L-DF-A-D-T-Y-PN-ED-B-T-EF-A-D-T-EY(?)

Y(?)F-A-D-T-PNY-E-D-BL-T-E-D-BL-F-A-B-D-T-Y-PN-E

Ar MPhAr MPh

Fr NPhFr NPh

Fr NPhFr NPhAr MPhAr MPhAr MPhAr MPhFr NPh

Ar MPh

Fr NPh

P HscapA ThscapA ThscapA ThscapP HscapP HscapA Thrept

P ChsuffrA ThcaespP HreptFr NPh

P EA

P GbulbP GbulbP Hrept

P GparP Gpar

A ThcaespA ThreptP HscapP HscapP HscapP HscapP HscapA ThscapA Chfrut

A ThscapA ThscapA ThscapA ThscapA Thscap

L,agascalia 17(2), 1994 235

Arthrocnemum fniticosum (L.) Moq. E-B-Y P ChsuffrSalsola kali L.

E-Y A ThscapAmaranthaceae

Amaranthus deflexus L. L-F-A-D-T-Y-PN-E A HscapA. retroflexus L. P-PN-B-T-E-B A ThscapA. albus L. B-Y-T-E A ThscapA. blitoides S. Watson B-T-E A ThscapA. lividus L.

L-D-T-Y-E A ThscapPortulacaceae

Portulaca oleracea L. L-F-A-B-D-T-Y-PN-E A ThsuccCaryophyllaceae

Minuartia mediterranea (Link) K. Maly P-PN-Y-E A ThscapM. hamata (Hausskn.) Mattf. P-E A ThscapM. globulosa (Labill.) Schinz & Thell. P-PN-Y A ThscapM. hybrida (Vill.) Schischkin subsp. hybrida Y A ThscapM. hybrida (Vill.) Schischkin P A HscapM. venta (L.) Hiero. P-Y-PN P ChsuffrM. venia (L.) Hiero. var. attica Hayek P-Y-PN P ChsuffrBuffonia stricta (Sibth. & Sm.) Gurke var.hymmetia Hal. Y(?) P ThcaespArenaria leptoclados (Reincheb.) Guss. T-Y-PN-D A ThcaespA. guicciardi Heldr. P A ThscapA. serpyllifolia L. P-PN-Y-D-T A ThcaespStellaria media (L.) Vill. P-L-F-A-B-D-T-Y-PN-E A ThscapHolosteum umbellatum L. P-PN-Y-D-T A ThscapMoenchia graeca Boiss. & Heldr. P-Y A ThscapM. mandica (L.) Bartl. subsp. caerulea(Boiss.) Clapham. P-T-D A ThscapCerastium candidissimum Correos P-PN-Y P ThcaespC. illyricum Ard. P-L-F-A-D-T-Y-PN-E A ThcaespC. brachypetalum Desp. var. luridum Boiss. P-PN-Y-D-T A ThscapC. dichotomum L. P(?) A ThscapC. glomeratum Thuill. P-PN-Y-D-T A ThcaespScleranthus annuus L. subsp. polycarpos(L.) Thell. P-D-T A ChreptParonychia kapela (Hacq.) Kerner subsp. chionaea(Boiss.) Borhidi P-D-T-PN-E P HcaespP. capitata (L.) Lam. L-PN-E P ThcaespP. macrosepala Boiss. L-Y P ThcaespHerniaria cinerea DC. Y(?) A ThreptH. hirsuta L. E-Y-T A HreptPolycarpon tetraphyllum (L.) L. P-PN-T-Y A ThcaespSpergularia rubra (L.) Presl. T-Y-E A ThcaespAgrostemma githago L. P-D-T-Y-PN-E-B A ThscapVelezia rigida L. Y-D-T-B A ThcaespSilene conica L. P-PN-Y-T A ThcaespS. nocturna L. T-Y-B A ThscapS. rigidula Sibth. & Sm. L-Y-E-B A ThscapS. spinescens Sibth. & Sm. P-PN-T-Y P HscapS. italica (L.) Pers. P-L-Y-E-B P HscapS. vulgaris (Moench) Garcke P-L-T-B P ChsuffrS. vulgaris (Moench) Garcke subsp. vulgaris Y-E-T-B P ChsuffrS. muscipula L. E-B A ThscapS. behen L. T-P-PN-Y-E-B A ThscapS. sedoides Poiret P-PN-T A Thscap


S. gallica L. P-D-T-Y-PN-ES. colorata Poiret P-L-D-T-Y-PN-E-BS. cretica L. P-T-D-BS. longipetela Vent. P-BS. gigantea L. P(?)Petrohagia armerioides P.W. Ball & Heywood P-EP. illyrica (L.) P.W. Ball & Heywood P-YP. ochroleuca Sibth. & Sm. P(?)P. thessala (Boiss.) P.W. Ball & Heywood P(?)P. velutina (Guss.) P.W. Ball & Heywood P-YSaponaria calabrica (Guss.) P.W. Ball & HeywoodP-TVaccaria pyramidata Medicus

P-T-B

Bolanthus graecus (Shreber) Barkoudah

P-YDianthus serratifolius Sibth. & Sm. P-PN-YD. deltoides L. PND. diffusus Sibth. & Sm. P-Y-D-T-PN-E

RanunculaceaeNigella arvensis L. subsp. aristata (Sibth. &Sm.) Nyman

L-YN. damascena L. P-L-F-A-D-T-Y-E-BDelphinium peregrinum L. L-D-T-Y-PN-E-BD. fissum Waldst. & Kit. P(?)Consolida tenuissima Soo. P(?)C. ambigua (L.) P.W. Ball & Heywood

PN

Anemone coronaria L. P-D-T-Y-PN-E-BA. hortensis L. T-PNA. pavoniana Lam. P-D-TA. pavoniana var. purpureoviolacea Boiss. T-YA. blanda Schott. & Kotschy

P-T

Clematis cirrhosa L. P-D-T-Y-E-BC. flammula L. P-T-PNC. vitalba L. P-D-T-BAdonis aestivalis L. L-A-D-T-Y-PN-BA. microcarpa DC. P-L-Y-BRanunculus chius DC. P-Y-BR. sardous Crantz P-L-Y-B

R. muricatus L. Y-E-T-BR. millefoliatus Vahl

P-D-T-ER. spruneranus Boiss. D-Y-ER. neapolitanus Ten. P-T-PNR. marginatus D' Urv. PN-TR. repens L. F-A-T-BR. arvensis L. P-F-A-T-BR. velutinus Ten. P-PN-TR. ficaria L. Y(?)R. ophioglossifolius Vill. P-TR. rumelicus Gris. P(?)R. padulosus Poiret

P-F-D-PN-A-TCeratocephalus falcatus (L.) Pers. P-PN-T

PaeoniaceaePaeonia mascula (L.) Miller

P(?)

BerberidaceaeBerberis cretica L. P-YLeontice leontopetalum L. P-PN-T-B

Lauraceae

P ThscapA ThscapA ThscapA ThscapP HrosP HcaespP HcaespP HscapP HscapA ThscapA HscapA ThscapP HcaespP HcaespP HcaespP Hcaesp

A ThcaespA ThcaespA ThcaespP HscapA ThscapA ThcaespP GrhP GrhP GrhP GrhP GrhFr NscandFr NscandFr NscandA ThscapA ThscapA ThscapA ThcaespA ThcaespP HscapP HcaespP HcaespA ThcaespP ThcaespA ThcaespP HcaespP GbulbA ThscapP ThscapP HcaespA Thscap

P Grh

Fr NPhP Grh

Lagascalia 17(2), 1994 237

*Laurus nobilis L.

D-T-PN-E-B Fr NPhPapaveraceae

P. rhoeas L. P-L-F-A-T-Y-PN-E-B A ThscapP. dubium L. P-PN-T-B A ThscapP. hybridum L. P-PN-Y-B A ThcaespP. argemone L. T-B A ThscapP. apulum Ten. P-PN-T-B A ThcayespGlaucium comiculatum (L.) J.H. Rudolph. Y-PN-E A ThscapG. flavum Crantz Y-PN-E-B B HscapRoemeria hybrida (L.) DC. Y(?) A ThscapHypecoum imberte Sibth. & Sm. L-Y A ThcaespCorydalis solida (L.) Swartz. subsp. solida P A GbulbFumaria macrocarpa Pan. P-L-B-D-T-Y-PN-E A ThcaespF. kralikii Jordan Y A ThscapF. densiflora DC. Y-D-T-B A ThscapE petteri Reichenb. L-F-A-D-T-Y-PN-B A ThcaespF. petteri Reichenb. subsp. thuretii (Boiss.)Pugsley P-PN-D-T-Y-E A ThcaespF. officinalis L. L-F-A-D-T-Y-PN-E-B A ThcaespE capreolata L.

P-PN-T A ThscapCapparidaceae

Capparis spinosa L. L-F-A-B-D-T-Y-PN-E P ChfrutC. ovata Desf.

T-D-B P ChfrutCruciferae

Sisymbrium irio L. L-F-A-B-D-T-Y-PN-E A ThscapS. orientale L. Y-B A ThscapErysimum pusillum Bory & Chaub. subsp.parnasii Hayek P-E-Y P HscapE. graecum Boiss. & Heldr. L-T-Y-E-B B ThscapNasturtium officinale R. Br. Y(?) P HscapMalcolmia africana (L.) R. Br. T-E-Y-B A ThscapM. maritima (L.) R. Br. Y-E A ThcaespM. graeca Boiss. & Spruner P-Y-T-E A ThscapM. flexulosa Sibth. & Sm. L-Y-E A ThscapMatthiola sinuata (L.) R. Br. E B ChsuffrM. fruticulosa (L.) Maire P-T-Y P ChsuffrM. longipetala (Vent.) DC. subsp. bicomis P.W.Ball Y(?) A ThscapM. tricuspidata (L.) R. Br. L-E A ThscapFibigia eriocarpa (DC.) Boiss. P-Y P HscapCardamine graeca L. P-T-Y-E A ThcaespC. hirsuta L. P-PN-T-Y-B A ThcaespArabis yema (L.) R. Br. P-Y-B A ThscapA. muralis Bertol. P-T-PN P HscapAubrietia deltoidea (L.) DC. P-Y-T P ChsuffrHesperis laciniata All. subsp. secundifloraBoiss. & Sprun. P-Y P HscapAlyssum saxatile L. P-L-F-A-D-T-Y-PN-E P ChsuffrA. montanum L. P-E P ChcaespA. montanum L. var. graecum Hal. Y(?) P ChcaespA. montanum L. var. hymettium Hal. T-Y P ChcaespA. desertorum Stapf. Y(?) A ChcaespA. foliosum Bory & Chaub. P-PN-Y A ThscapA. campestre L. P-PN-T A ThscapA. minutum Sclilecht P(?) A Thscap

238 Contribution to (he study of (he flora of Attica (Greece)

Clypeola jonthlaspi L.C. eriocarpa Cav.Erophila venia (L.) Chevall.Neslia paniculata (L.) Desv.Capsella bursa-pastoris (L.) MedicusCamelina rumelica Vel.Aethionema saxatile (L.) R. Br.Thlaspi perfoliatum L.Th. graecum JordanRapistrum rugosum (L.) Ah. subsp. orientale(L.) ArcangeliCalepina irregularis Thell.lberis pruitii TineoI. pruitii Tinco var. spruneri Hal.I. odorata L.Coronopus squamatus (Forskal) AshersonLepidium hirtum (L.) Sm. subsp. nebrodense(Rafin) Thell.Cardaria draba (L.) Desv.Brassica cretica Lam.B. tournefortii GouanB. nigra (L.) KochSinapis arvensis L.S. alba L.Eruca vesicaria (L.) Cav. subsp. sativa (Miller)Thell.E. vesicaria L. var. longirostris RouyHirschfeldia incana (L.) Lagreze-FossatCakile maritima Scop.Isatis tinctoria L.Biscutella didyma L.B. variegata Boiss. & ReuterRaphanus raphanistrum L.Homungia petraea (L.) Reinheb.

ResedaceaeReseda alba L.R. lutea L.

CrassulaceaeUmbilicus parviflorus (Desf.) DC.U. chloranthus Hal.U. erectus DC.U. horizontalis (Guss.) DC.U. rupestris Salisb.Sedum sediforme (Jacq.) PauS. tenuifolium (Sibth. & Sm.) Strobl.S. litoreum Guss.S. rubens L.S. hispanicum L.S. athoum DC.S. ochroleucum ChaixS. acre L.S. sartorianum Boiss.S. caespitosum (Cav.) DC.S. laconicum Boiss.

A ThscapA ThscapA ThscapA ThscapA ThscapA ThscapA ThscapA ThscapP Thscap

A ThscapA ThscapP HscapP HscapP HcaespA Thrept

P HscapP HscapP *HscapA ThscapP ThscapA ThscapA Thscap

A HscapA ThscapA ThscapA ThscapP HscapA ThscapA ThscapA ThscapA Thscap

A HscapA Hscap

P GbulbP GbulbP GbulbP GbulbP GbulbP ChscapP ChsuccA ThcaespA ThsuccA ThcaespP ChscapP ChscapP ChscapP ChscapA ThscapP Chsucc

P-PN-T-Y-EEP-YF-A-T-EP-L-F-A-B-D-T-Y-PN-ET-YP-L-Y-EP-PN-YP(?)

P-T-BP-T-PN-B

T-YP-PN-D-T-BP(?)

L-F-A-B-D-T-Y-PN-EP-D-T-Y-ET-Y-EF-A-B-D-TF-A-B-D-T-Y-PN-EF-A-B-D-T-PN

L-F-A-D-T-PN-E-BT-YF-A-D-T-Y-PN-E-BD-Y-EP(?)P-PN-L-F-A-T-Y-E-BT-PND-T-Y-PN-E-BT-Y

L-F-A-B-D-T-Y-PN-EL-F-A-B-D-T-Y-PN-E

P-Y-EP-T-Y

P-Y-TYL-F-A-D-T-PN-E-YF-A-D-T-PN-E-P-YP-Y-L-AP-PN-YP-F-A-D-T-Y-PN-EP-YP-PN-YT-YP-PN-YP-A-PN

Saxifragaceae

Lagascalia 17(2), 1994 239

Saxifraga hederacea L. P-D-Y-E P ThreptS. tridactylites L. D-Y-E P ThscapS. graeca Boiss. P P HscapS. chrysoplenifolia Boiss. P(?) P Hscap

Platanaceae*Platanus orientalis L. PN-E-T-B Ar MPh

RosaceaeRubus ulmifolius Schott. T-PN-B Fr NscandR. canescens DC. P-D-T-Y-E-B P HscapRosa sempervirens L. P-PN-E Fr NscandR. corymbifera Bork. PN-T Fr NPhR. canina L. P-D-T-Y Fr NPhR. micrantha Borer P Fr NPhAremonia agrimonoides (L.) DC. P P HscapSanguisorba minor Scop. P-F-A-D-T-PN-E-B P HscapS. minor Scop. subsp. magnolii (Spach) Briq. P-Y-PN-B P HscapS. minor Scop. subsp. muricata Briq. P P HscapSarcopoterium spinosum (L.) Spach P-L-F-A-D-T-Y-PN-E-B Fr ChfrutPotentilla micrantha Ramond P(?) P HcaespP. hirta L. P-PN P HscapP. hitra L. var. pedata Koch. P(?) P HscapGeum urbanum L. P P HscapAphanes arvensis L. P-Y-E A ThscapPyrus amygdaliformis Vill. P-F-A-D-T-Y-PN-E-B Fr NPhMalus domestica Bork T-PN Av MPh*Pyracantha coccinea M.J. Roemer PN-B Fr NPhCrataegus monogyna Jacq. P-T-PN Fr NPhC. heldreichii Boiss. P Fr NPhAmelanchier ovalis Medicus P(?) Fr NPhPrunus prostrata Labill. P-D-T-E Fr NPhP. cocomilia Ten. P(?) Ar MPhP. webbii (Spach) Vierh. P-PN Av MPhP. domestica subsp. insititia (L.) C.K. Schneider T-P Fr NPh

Leguminosae*Cercis siliquastnim L. P-D-T-Y-PN-E-B Fr MPh*Ceratonia siliqua L. D-T-PN-E-B Ar MPh*Parkinsonia aculeat'a L. E-B Av MPhAnagyris foetida L. P-Y-E P NPhCalicotome villosa (Poiret) Link P-D-T-Y-PN-E Fr NPhGenista acanthoclada DC. P-D-T-Y-PN-E Fr NPh*Spartium junceum L. P-F-D-T-Y-PN-E-B Fr NPhLupinus angustifolius L. T-Y-PN-E A Thscap*Robinia pseudacacia L. T-Y-PN-E-B Ar MPhColutea arborescens L. P-PN-T-E P NPhAstragalus sinaicus Boiss. P-PN-T-Y-E A HscapA. hellenicus Boiss. P-PN-E P ThscapA. boeticus L. D-T-PN A HscapA. graecus Boiss. & Spniner D-T-PN-E A HscapA. monspessulans L. P-T-PN P ChfrutA. hamosus L. F-A A ThrosA. spruneri Boiss. P-L-D-T-Y-PN-E P ChfrutA. angustifolius Lam. P P ChfrutA. parnasii Boiss. P(?) P ChfrutA. depressus L. P(?) P HrosBiserrula pelecinus L. P-PN-L-B A Thscap

240 Contribution to (he study of (he flora of Attica (Greece)

Psoralea bituminosa L. P-PN-L-F-A-B-D-T-Y-E P HscapVicia cracca L. D-T-Y P HscandV. villosa Roth. subsp. microphylla (D' Uvr.)P.W. Ball P-L-D-T-Y-PN-E-B A ThscandV. villosa Roth. subsp. varia (Host) Cord Y(?) A ThscandV. villosa Roth. subsp. eriocarpa (Hausskn.)P.W. Ball P-T-Y-B A ThscandV. bithynica (L.) L. P-PN-B P ThscandV. pinetorum Boiss. & Spruner P(?) P ThscapV. cretica Boiss. & Heldr. P-B A ThscapV. cretica Boiss. & Heldr. var. spruneri Boiss. P-Y A ThscapV. hirsuta (L.) S.F. Gray P-D-T-Y-PN-B A ThscandV. tenuissima (Bieb.) Sing. & Thell. E-B A ThscandV. pubescens (DC.) Link P-D-T-Y-PN-B A ThscandV. sibthorpii Boiss. P-T-Y-B A ThscandV. sativa L. subsp. amphicarpa (Dorthes)Ascherson & Graebner L-B-D-T A ThscandV. pannonica Crantz. subsp. striata (Bieb.)Nyman PN-T-B A ThscapV. barbazitae Ten. & Guss. P A ThcaespV. peregrina L. B-D-T-Y-E A ThscandV. lathyroides L. P-PN-Y-B A ThscandV. hybrida L. var. spruria Strobl. Y-B A ThscapV. hybrida L. P-PN-F-A-B A ThscandV. lutea L. D-T-PN-B A ThscandV. melanops Sibth. & Sm. P A ThscandLens nigriscans (Bieb.) Gordon P-B A ThscandL. ervoides (Brigh) Grande P(?) A ThscandLathyrus niger (L.) Bemh. subsp. jordanii(Ten.) Arcanqeli PN P GrhL. pratensis L. P-D-T-PN-B P HscapL. saxatilis (Vent) Vis. P-Y-E-B A ThscapL. cicera L. P-T-Y-E-B A ThscandL. sphaericus Retz. P-PN-Y-E-B A ThscapL. aphaca L. P-L-D-T-Y-PN-E-B A ThscandL. setifolius L. P-L-Y-B A ThscandL. grandiflorus Sibth. & Sm. P-F-A-B P GrhL. nissolia L. P(?) A ThscandL. laxiflorus Desf. P-PN P GrhL. digitatus (Bieb.) Fiori P(?) P GrhL. venetus Wohl. P(?) P GrhL. hirsutus L. P-B A ThscandL. amphicarpos L. P(?) A ThscapPisum sativum L. D-T-PN-B A ThscandP. sativum L. subsp. elatius (Bieb.) Asherson& Graebner P A ThscandOnonis omithopodioides L. L-Y-E-B A ThscapO. reclinata L. D-T-Y-PN-E-B A ThscandO. pubescens L. P-D-T-E-B A ThscapO. viscosa L. subsp. breviflora (DC.) Nyman P-Y A ThcaespO. viscosa L. subsp. sieberi (Besser ex DC.) Sirj. Y-E A ThcaespO. pusilla L. P(?) P HscapO. spinosa L. subsp. antiquorum (L.) Arcangeli Y-B P ChfrutMelilotus officinalis (L.) Pallas L-F-A-B-D-T-PN B HcaespM. indica L. L-B-D-T A Thscap

Lagascalia 17(2), 1994 241

M. italica (L.) Lam. P-Y-E A ThscapM. neapolitana Ten. P-L-Y-E A ThcaespTrigonella graeca (Boiss. & Spruner) Boiss. P-L-Y A ThscapT. balansae Boiss. & Reuter P-PN-D-B A ThscapT. sprunerana Boiss. L-D-T-Y-PN-E A ThscapT. coerulescens (Bieb.) Hal. L-Y-E-B A ThscapT. gladiata Steven P-D-T-E-B A ThscapT. monspeliaca L. P-PN A ThscapT. coerulea Ser. L(?) A ThscapT. spicata Sibth. & Sm. L-D-B A ThscapMedicago lupulina L. P-F-A-D-T-PN-B B HcaespM. sativa L. subsp. sativa P-F-A-B-D-T-PN P ChscapM. sativa L. subsp. falcata (L.) Arcangeli P-D-T-PN-B P HscapM. rugosa Desr. P-D-B A Thscap*M. arborea L. L-F-A-D-T-PN-B Fr ChsuffrM. orbicularis (L.) Bartal. P-F-A-Y-E-B A ThcaespM. scutellata (L.) Miller P-PN-D-B A ThscapM. marina L. T-Y-E P ChsuffrM. truncatula Gaertner P(?) A ThcaespM. rigidula (L.) Al! P-L-F-A-Y-E-B A ThscapM. littoralis Rohde T-Y-E A ThcaespM. arabica (L.) Hudson L-Y-E-B A ThcaespM. disciformis DC. P-L-T-Y-PN-B A ThcaespM. polymorpha L. P-T-Y A ThcaespM. coronata (L.) Baila!. P-L-T-Y-B A ThcaespM. minima (L.) Bartal. P-Y-E-B A ThcaespTrifolium physodes Stev. P-Y-B P ThscapT. repens L. P-D-T-PN-B A HcaespT. nigrescens Viv. P-L-D-T-PN-B A ThcaespT. tomentosum L. P-L-F-A-D-T-Y-PN-E-B A ThcaespT. fragiferum L. P-D-B P HreptT. resupinatum L. P-PN A ThcaespT. uniflorum L. P-Y P HreptT. spumosum L. Y(?) A ThcaespT. suffocatum L. P-Y A ThcaespT. aureum Pollich P-Y B ThcaespT. speciosum Willd. Y(?) A ThcaespT. scabrum L. P-Y-B A ThcaespT. arvense L. P-Y-E-B A ThcaespT. stellatum L. P-L-F-A-B-D-T-Y-PN-E A ThcaespT. pratense L. D-T-PN-B P HcaespT. boissieri Guss. P-L A ThscapT. angustifolium L. P-F-A-D-T-Y-PN-E-B A ThcaespT. subterraneum L. P(?) A ThreptT. lappaceum L. P(?) A ThcaespT. hirtum Al!. P(?) A ThscapT. cherlieri L. P-PN-D-B A ThreptT. dalmaticum Vis. P A ThscapT. clypeatum L. P(?) A ThscapT. diffusum Ehrh. P A ThscapT. latinum Sebastian P(?) A ThscapT. leucanthum Bieb. P(?) A ThscapDorycnium hirsutum (L.) Ser. P-D-T-PN-B P ChsuffrD. pentaphyllum Scop. subsp. herbaceum (Vill.)Rouy P-L-Y-PN-E P Chsuffr

242 Contribution ro (he study of (he flora of Attica (Greece)

D. rectum (L.) Ser. P-PN-B P ChsuffrLotus comiculatus L. P-B-D-T P ChcaespL. peregrinus L. Y-E A ThscapL. edulis L. Y-B A ThcaespL. omithopodioides L. P-Y-B A ThcaespHymenocarpos circinnatus (L.) Savi P-PN A ThscapAnthyllis hermanniae L. P-L-B-D-T-Y-PN-E Fr NPhA. montana L. subsp. hispanica (Degen &Herrier) Cullen PN P ChsuffrA. vulvenaria L. P-L-B P HscapA. vulvenaria L. var. spruneri Boiss. Y(?) P HscapA. tetraphylla L. P-L-Y A ThcaespCoronilla minima L. PN Fr NPhC. emerus L. subsp. emeroides (Boiss. &Spruner) Hayek P-PN-D-T-Y-E-B Fr NPhHippocrepis unisiliquosa L. P-PN-Y-E-B A ThcaespH. ciliata Willd. P-PN-L-Y-B A ThcaespH. comosa L. P P ThscapScorpiurus muricatus L. L(?) A ThscapOnobrychis aequidentata (Sibth. & Sm.) D' Urv. P-Y-B P ThscapO. ebenoides Boiss. & Spruner P-L-D-T-Y-PN-E P HscapO. caput-galli (L.) Lam. P-PN-L-Y-E A ThscapO. viciifolia Scop. PN A HscapHedysarum spinosissimum L. Y-B A ThscapEbenus sibthorpii DC. P P Chsuffr

OxalidaceaeOxalis corniculata L. L-F-A-B-D-T-Y-PN-E A ThreptO. pes-caprae L. L-F-A-B-D-T-Y-PN-E P Hros

GeraniaceaeGeranium rotundifolium L. D-Y-E-B A ThscapG. sanguineum L. PN-B P HscapG. molle L. P-L-F-A-D-T-Y-PN-E-B A ThscapG. lucidum L. P-D-T-Y-PN-E A ThscapG. purpureum Vill. P-PN-L-Y-T A ThscapErodium laciniatum (Cav.) Willd. T-Y-E A ThscapE. malacoides (L.) L Her. L-T-D-Y-E-B A ThscapE. gruinum (L.) L' Her. P-E-B A ThscapE. ciconium (L.) L' Her. P-PN-T A ThscapE. cicutarium (L.) L' Her. P-L-F-A-B-D-T-Y-PN-E A ThscapE. moschatum (L.) L' Her. T-PN-B A ThscapE. acaule (L.) Becherer & Thell. P-PN-F-A-D-B A Thscap

ZygophyllaceaeTribulus terrestris L. P-L-F-A-B-D-T-Y-PN-E A Thcaesp

LinaceaeLinum bienne Miller P-E P HscapL. trigynum L. P-Y-E A ThscapL. strictum L. P-PN-L A ThscapL. strictum L. subsp. corymbulosum (Reinheb.)Rouy P(?) A ThscapL. strictum L. var. spicatum (Lam.) Pers. Y(?) A ThscapL. leucanthemum Boiss. & Spruner P-PN-Y P HscapL. nodiflorum L. L A ThscapL. pubescens Banks & Solander P(?) A Thscap

EuphorbiaceaeChrozophora tinctoria (L.) A. Juss. Y-E-B A Thscap

Lagascalia 17(2), 1994 243

Mercurialis anima L.M. perennis L.Euphorbia dendroides L.E. acanthothamnos Heldr. & Sart.E. helioscopia L.E. myrsinites L.E. exigua L.E. falcata L.E. characias L. subsp. wulfenii A.R. Sm.E. characias L. var. sibthorpiiE. peplus L.E. apios L.E. taurinensis Ah.E. rigida Bieb.E. heldreichii Orph.E. deflexa Sibth. & Sm.E. oblongata Griseb.

RutaceaeRuta graveolens L.R. chalepensis L.

Simarubaceae*Ailanthus altissima (Miller) Swing.

Meliaceae*Melia azedarach L.

PolygalaceaePolygala venulosa Sibth. & Sm.P. nicaensis Risso

CoriariaceaeCoriaria myrtifolia L.

AnacardiaceaeRhus coriaria L.Cotinus coggygria Scop.Pistacia terebinthus L.P. lentiscus L.*Schinus molle L.

SapindaceaeKoelreuteria paniculata Laxm.

RhamnaceaeRhamnus lycioides L. subsp. graecus (Boiss. &Reuter) TutinR. alatemus L.Frangula rupestris Schur.

MalvaceaeMalva aegyptia L.M. cretica Cav.M. sylvestris L.M. pusilla Sm.Althaea officinalis L.A. cannabina L.Alcea rosea L.A. pallida (Willd.) Waldst. & Kit.

ThymelaeaceaeThymelaea hirsuta (L.) Endl.Th. tartonaira (L.) All.Daphne gnidium L.

P-F-A-D-Y-E-BF-A-B

P-F-A-D-T-Y-PN-E-BL-F-A-D-T-Y-PN-E-BP-PN-F-A-D-T-Y-EP-F-A-T-E-BT-PN-BL-TP-Y(?)F-A-D-Y-E-BP-T-YY(?)P-YP(?)P-YP-PN-D-B

L-B-D-T-Y-E

E-B

Y-B

E-L-Y-BP-Y-PN

P(?)

P-PN-TP-T-PNP-A-D-T-PN-EP-L-F-A-D-T-Y-PN-E-BT-Y-E

A

P-PN-D-Y-E

P(?)

Y(?)B-D-T-Y-EP-L-F-A-B-D-T-Y-PN-EF-A-BP-D-T-BT-PNT-E-BT-B

P-L-D-T-Y-PN-E-BP-L-D-T-Y-PN-E-BP-T-PN

A ThscapP GscapFr NPhFr ChfrutA ThscapP ChscapA ThscapA ThscapP ChsuffrFr NPhA ThcaespP GbulpA ThscapP ChsuffrP HcaespP HcaespP Gscap

P ChfrutP Chsuffr

Ar MPh

Ar MPh

P ThscapP Thscap

Fr NPh

Fr NPhFr NPhFr NPhFr NPhFr NPh

Ar MPh

Fr NPhFr NPhFr NPh

A ThscapA ThscapP HscapA ThscapP HscapP HscapP HscapP Hscap

Fr NPhFr NPhP Chsuffr


GuttiferaeHypericum empetrifolium Willd. P-D-T-Y-PN-E P ChsuffrH. hirsutum L. D-T-B P HscapH. perfoliatum L. P-T-Y-PN-E-B P HscapH. tetrapterum Fries PN P HscapH. triquetrifolium. Turra P-L-D-T-Y-PN-E-B P HscapH. perforatum L.

P-PN-D-B P HscapViolaceae

Viola odorata L. P-PN P HrosV. hymmetia Boiss. & Heldr. P-Y-PN A ThscapV. alba Besser subsp. thessala Hayek P P HrosV. reichenbachiana Jordan P(?) P HscapV. kitaibeliana Schultes P(?) A Thscap

CistaceaeCistus incanus L. B-D-PN Fr NPhC. incanus L. subsp. creticus (L.) Heywood B-P-L-D-T-PN-E Fr NPhC. parviflorus Lam. B-D-Y-E Fr NPhC. monspeliensis L. D-T-Y-PN-E Fr NPhC. salvifolius L. B-L-D-T-Y-PN-E - Fr NPhHalimium umbellatum (L.) Spach T-PN Fr NPhTuberaria guttata (L.) Fourr. P-PN-Y-E-B A ThcaespHelianthemum lavandulifolium Miller P-Y-E Fr ChsuffrH. hymettium Boiss. & Heldr. P-PN-D-Y-E P ChscapH. salicifolium (L.) Miller Y-PN-T A ThscapH. apenninum (L.) Miller Y Fr ChsuffrFumana arabica (L.) Spach P-L-F-A-D-T-Y-E P ChsuffrF. thymifolia (L.) Spach P-L-D-T-Y-PN-E P ChsuffrF. ericoides Pau.

L-T P ChsuffrTamaricaceae

Tamarix gallica L. PN-E-B Ar MPhT. parviflora DC.

Y-B As MPhCucurbitaceae

Ecballium elaterium (L.) A. Richard P-L-F-A-T-B-D-Y-PN-E P HscapBryonia cretica L. subsp. dioica (Jacq.) Tutin Y-E-B P Hscand

CactaceaeOpuntia ficus-indica (L.) Miller E-B Fr NPh

LythraceaeLythrum hyssopifolia L. Y-E A Thrept

MyrtaceaeMyrtus communis L. B-P-T-PN Fr NPh*Eucalyptus robustus Sm. E-B As MPh*E. globulus Labil.

E-B As MPhPunicaceae

*Punica grartatum L. E-B Fr NPhOnagraceae

Epilobium hirsutum L. P P HscapE. lanceolatum Sebastiani & Mauri P P Hscap

TheligonaceaeTheligonum cynocrambe L. P(?) A Thscap

Araliaceae*Hedera helix L.

P-F-A-D-T-Y-PN-E-B Fr NscandUmbelliferae

Eryngium maritimum L. D-E P HscapE. campestre L. L-F-A-D-T-Y-PN-E-B P HscapE. creticum Lam. P-B P Hscap

Lagascalia 17(2), 1994 245

Lagoecia cuminoides L.Anthriscus tenerimmus Boiss. & SprunerScandix australis L.S. australis L. subsp. grandiflora (L.) Thell.S. pecten-veneris L.S. olusatrum L.S. perfoliatum L.Scaligeria cretica (Miller) Boiss.Smymium rotundifolium MillerS. perfoliatum L.Pimpinella cretica PoiretP. peregrina L.Conium maculatum L.Huetia cynapioides (Guss.) P.W. Ball subsp.macrocarpa (Boiss. & Spruner) P.W. BallJohrenia distans (Griseb.) Hal.Foeniculum vulgare MillerBupleurum fruticosum L.B. trichopodium Boiss. & SprunerB. lancifolium Homem.Carum multiflorum (Sibth. & Sm.) Boiss.Ammi majus L.Ferulago nodosa (L.) Boiss.Tordylium apulum L.Ferula communis L. subsp. glauca (L.) Rouy& CamusTorilis leptophylla (L.) Reincheb.T. nodosa (L.) GaertnerT. tenella (Delile) Reinheb.Thapsia garganica L.Caucalis platycarpos L.Oenanthe silaefolia Bieb.Orlaya kochii HeywoodOpopanax hispidus (Friv.) Griseb.Peucedanum vittijugum Boiss.Malabaila involucrata Boiss. & SprunerDaucus guttatus Sibth. & Sm.D. carota L. subsp. maximus (Desf.) BallD. carota L.D. involucratus Sibth. & Sm.Pseudorlaya pumila (L.) Grande

EricaceaeErica arborea L.E. manipuliflora Salisb.Arbutus unedo L.A. andrachne L.

PrimulaceaeCyclamen hederifolium AitonC. graecum LinkC. persicum MillerAnagallis arvensis L.A. foemina Miller

PlumbaginaceaeLimonium sinuatum (L.) MillerL. echioides (L.) Miller

P-L-F-A-Y-PN-E-BP-PN-YP-PN-L-D-Y-BP-D-T-BL-F-B-Y-EE-BE-BP-YP-PN-TP-Y-TL-YP-D-BP-T-B

Y(?)P-YPN-E-BP-E-T-PNP-L-T-Y-PNP-PN-D-BP-Y-D-BP-PN-E-BP-E-Y-BL-F-A-PN-E-B

P-TY(?)P-PN-L-F-A-BYL-F-A-D-T-Y-PN-E-BT-BP(?)Y(?)P-PN-BP(?)P-YD-T-Y-PN-E-BE-BPN-T-BP-YPN-Y

P-D-T-EP-D-T-PN-EP-D-T-PN-EP-D-T-PN-E

P-T-PNP-L-F-A-D-T-Y-PN-EF-A-TF-A-D-T-Y-PN-E-BP-L-D-T-Y-E-B

YY

A ThscapA ThscapA ThscapA ThscapA ThscapA ThscapA ThscapB HcaespB HscapB ThscapA ThscapB HscapP Hscap

P GbulbP ThcaespP HscapP ThscapA ThcaespA ThcaespB HscapA ThcaespP HscapA Thscap

P HscapA ThscapA ThscapA ThscapP HscapA ThscapP HscapA ThscapP HscapP ThscapB ThscapA ThscapB HscapB HscapA ThscapA Thscap

Fr NPhFr NPhFr NPhFr NPh

P GbulbP GbulbP GbulbA ThcaespA Thcaesp

P HrosA Thros


Armeria undulata (Bory) Boiss. P-PN-Y P HscapStyracaceae

Styrax officinalis L.

P-T Fr NPhOleaceae

Fraxinus ornus L. P-PN-T Ar MPh*Olea europaea L. var. europaea E-B Ar MPhO. europaea L. var. sylvestris Brot. P-F-A-D-T-Y-PN-E Fr NPhPhillyrea latifolia L. E Fr NPhP. media L.

P-D-T-Y Fr NPhGentianaceae

Blackstonia perfoliata (L.) Hudson P A ThscapCentaurium erythraea Rafn subsp. erythraea P-Y-B A ThscapC. pulchellum (Schwartz) Druce P A Thscap

Apocynaceae*Nerium oleander L. P-D-T-Y-PN-E-B Fr NscandVinca minor L.

P-PN-T P NscandRubiaceae

Asperula arvensis L. P-D-T-Y-E-B A ThscapA. lutea Sibth. & Sm. P-L P HcaespA. rigida Sibth. & Sm. P(?) P HcaespA. boissieri Heldr. P(?) P HcaespCrucianella latifolia L. P-Y A ThscapC. graeca Boiss. P-PN-Y A ThscapGalium verticillatum Danth P-PN-Y P ThscapG. tricomutum Dandy P-Y A ThscapG. incurvum Sibth. & Sm. E-Y A ThscapG. aparine L. L-B-D-T-Y-PN-E A ThscandG. murale (L.) Ah. P-B-D-T-Y-E A ThscapG. monachinii Boiss. & Heldr. Y A ThscapG. setaceum Lam. Y A ThscapG. capitatum Bory & Chaub. P-Y A ThscapG. rotundifolium L. P P ChcaespValantia hispida L. P-PN-Y A ThscapRubia tenuifolia D' Urv. P-Y P NscandR. peregrina L. P-D P HscandR. tinctorum L. P-B-D P HscandPutoria calabrica (L. fil.) DC. P-B Fr NPh

ConvolvulaceaeCuscuta epithymum (L.) L. B-D-Y-PN P ThparConvolvulus boissieri Steudel E P HscapC. boissieri Steudel. subsp. compactus (Boiss.)Stace P(?) P HscapC. dorycnium L. E-B P HscapC. arvensis L. P-L-F-A-B-D-T-Y-PN-E P HscandC. althaeoides L. P-D-T-Y-PN-B P HscandC. althaeoides L. subsp. althaeoides E-B P HscandC. althaeoides L. subsp. tenuissimus (Sibth.& Sm.) Stace P-L-F-A-B-D-T-PN-E-Y P HscandC. cantabrica L.

L-B P HcaespBoraginaceae

Heliotropium europaeum L. L-F-A-D-T-Y-E-B A ThscapH. hirsutissimum Grauer P-B A ThscapBuglossoides tenuifiora (L. fil.) I.M. Johnston Y A ThscapB. arvensis (L.) I.M. Johnston P-Y-E A ThscapB. arvensis (L.) I.M. Johnston subsp. arvensis D-T-PN A Thscap

Lagascalia 17(2), 1994 247

B. arvensis (L.) I.M. Johnston subsp.gasparrinii(Heldr. ex Guss.) R. Fernandes P A ThscapOnosma frutescens Lam. P-PN-L-Y-E P ChsuffrO. erecta Sibth. & Sm. P-Y P ChsuffrO. erecta Sibth. & Sm. var. pubiflorum Ha!. Y(?) P ChsuffrO. montana Sibth. & Sm. P-Y P HcaespO. graeca Boiss. P-Y B HscapCerinthe minor L. P-Y-E-B A ThscapC. major L. L-B A ThscapC. retorta Sibth. & Sm. Y A ThcaespAlkanna tinctoria (L.) Tausch P-L-F-A-D-T-PN-E-B P HrosEchium angustifolium Miller D-T-PN P HscapE. italicum L. P-D-T-PN-E-B P HscapE. plantagineum L. P-L-F-A-D-T-Y-E-B A HscapE. parviflorum Moench L-D-Y-E A ThcaespE. arenarium Guss. D-Y-E B HscapAnchusa undulata L. D-T-PN-E-B B HscapA. undulata L. subsp. hybrida (Ten.) Coutinho P-PN B HscapA. sartorii Heldr. Y-PN B HscapA. variegata (L.) Lehm. F-A-Y-E-B A ThscapA. azurea Miller P(?) B HscapA. officinalis L. L P HscapMyosotis incrassata Guss. L-Y A ThscapM. ramosissima Rochel subsp. ramosissima Y A ThscapM. sylvatica Hoffm. P B HscapM. arvensis (L.) Hill. P-PN B ThscapM. discolor Pers. P(?) A ThscapCynoglossum columnae Ten. P-D-T-Y-PN-E A HscapC. creticum Miller E-B B HscapSymphytum bulbosum (L.) Schimp. P P HscapNonea obtusifolia (Willd.) DC. P(?) A ThscapAsperugo procumbens L. P-Y-B A Thscap

VerbenaceaeVitex angus-castus L. P-D-T-Y-PN-E-B P NPhVerbena officinalis L. P-D-T-PN-E-B P Hscap

LabiataeAjuga iva (L.) Schreber E-T-B P ChscapA. orientalis L. P P HscapA. chamaepitys (L.) Schreber subsp. chia(Schreber) Arcangeli P-Y-B P HscapTeucrium divaricatum Sieber P-L-Y-E-T P ChsuffrT. polium L. P-L-F-A-B-D-T-Y-PN-E P ChsuffrT. chamaedrys L. P-PN P ChsuffrScutellaria rubicunda Hornem subsp. rupestrisI.B.K. Richardson P(?) P HscapPrasium majus L. P-F-A-PN-E P ChsuffrMarrubium vulgare L. L-F-A-T-Y-E-B P ChsuffrSideritis romana L. P-PN A ThscapS. syriaca L. P Fr ChsuffrPhlomis fruticosa L. P-L-F-A-D-T-Y-PN-E-B P ChsuffrLamium garganicum L. subsp. pictum (Boiss.& Heldr.) P.W. Ball L-F-A-D-T-Y-PN-E P HscapL. garganicum L. P P HcaespL. amplexicaule L. P-L-F-A-B-D-T-Y-PN-E A ThscapBallota acetabulosa (L.) Bentham P-L-F-A-B-D-T-Y-PN-E P Chsuffr

P-T-Y-PN-E

HscapP-T-E

HscapHscap

T-B

A ThscapE

HscapE

HscapP-D-PN

HscapP-T

HscapP-F-A-D-T-Y-PN-E

ChsuffrY-E-B

ChsuffrP-PN-L

A Thscap

P(?)P-L-F-A-D-T-Y-PN-E-BP-L-F-A-D-T-Y-PN-E-BP-PNP-BP-BD-T-BD-T-Y-PN-EP-B-D-T-Y-PN-EP-PN-YP-L-YP-Y-PN-E-BD-PN-E-BD-Y-BPND-T-PNP-F-A-B-D-T-Y-PN-E-LP-PN-L-Y-EP-Y-ET-PNB-P-PN-L-F-A-T-D-Y-E

P-F-A-B-TE-BF-A-B-D-T-Y-ED-T-BB-PN-E-DF-A-B-D-E

P-L-F-A-D-T-Y-PN-E-BPN-TP-E

P-Y-BP-YP-BP-BP-YPN-EP-D-T-Y-PN-E-BP-PN-E-BP-Y-E

Fr ChsuffrChsuffrChsuffrChsuffrHscapHscapChsuffrHscapChfrutChsuffrChsuffrHscap

Fr NPhChsuffr

Fr NPhFr Chsuffr

ChsuffrChsuffrHrosHscap

A Hscap

HscapA ThscapA ThscapA ThscapA ThscapFr NPh

HscapHscapHscapHscapHscap

A ThscapHcaesp

A ThcaespHscap

A ThscapHscap

A ThscapA Thscap


Stachys cretica L. subsp. salvifolia (Ten.)Rech. fil.S. spruneri Boiss.S. acutifolia Bory & Chaub.S. spinulosa Sibth. & Sm.Gleochoma hederacea L.Melissa officinalis L.Nepeta sibthorpii BenthamN. catana L.Satureja thymbra L.Calamintha nepeta (L.) SaviAcinos rotundifolius Pers.A. alpinus (L.) Moenh. subsp. majoranifolius(Miller) P.W. BallMicromeria nervosa (Desf.) BenthamM. juliana (L) BenthamM. graeca (L.) BenthamPrunella vulgaris L.P. laciniata (L.) L.Thymbra spicata L.Origanum heracleoticum L.Thymus capitatus (L.) Hoffman & LinkT. atticus Cel.T. ocheus Heldr. & Sart.Mentha pulegium L.*Rosmarinus officinalis L.Lavandula stoechas L.L. angustifolia MillerSalvia officinalis L.S. triloba L. fil.S. pomifera L.S. verbenaca L.S. pratensis L.S. viridis L.

SolanaceaeHyoscyamus albus L.H. niger L.Solanum nigrum L.S. citrullifolium A. BraunDatura stramonium L.Nicotiana glauca R.C. Graham

SchrophulariaceaeVerbascum undulatum Lam.V. thapsus L. subsp. giganteum (Willd.) NymanV. sinuatum L.V. graecum Heldr. & Sart.V. boissieri (Heldr. & Sart. ex Boiss.) O. Kuntz.Linaria simplex (Willd) DC.Kickxia commutata (Bemh) Fritsch.K. elatine (L.) Dumort.Scrophularia lucida L.S. peregrina L.S. heterophylla Willd.Misopates orontium (L.) Rafin.Veronica glauca Sibth. & Sm.

Lagascalia 17(2), 1994 249

V. arvensis L.V. polita FriesV. cymbalaria Bodard.V. anagalloides Guss.Parentucellia latifolia (L.) Caruel.Bellardia trixago (L.) Ah.

GlobulariaceaeGlobularia alypum L.

Acanthaceae*Acanthus mollis L.A. spinosus L.

OrobanchaceaeOrobanche crenata ForskalO. pubescens D' Urv.O. vesicolor D' Un'.O. elatior SuttonO. ramosa L. subsp. mutelii (F.W. Schultz)CoutinhoO. ramosa L. subsp. nana (Reuter) CoutinhoO. minor Sm.O. gracilis Sm.O. alba StephanO. fuliginosa Reuter

PlantaginaceaePlantago major L.P. major L. subsp. intermedia (DC.) ArcangeliP. coronopus L.P. lanceolata L.P. lagopus L.P. albicans L.P. bellardii All.P. bellardii Ah. subsp. deflexa (Pilger) Rech. fil.P. arenaria Waldst. & Kit.

CaprifoliaceaeLonicera implexa AitonL. etrusca G. SantiL. periclymenum L.

ValerianaceaeValerianella echinata (L.) DC.V. hirsutissima LinkV. eriocarpa Desr.V. coronata (L.) DC.V. turgida (Steven) BetckeV. dioscoridis Sibth. & Sm.Centranthus ruber (L.) DC.C. ruber (L.) DC. subsp. sibthorpii (Heldr.& Sart.) HayekC. calcitrapae DC.

DipsacaceaeCephalaria ambrosioides (Sibth. & Sm.)Roemer SchultesKnautia intergrifolia (L.) Bertol.Scabiosa hymettia Boiss. & SprunerS. webbiana Don.S. atropurpurea L.

P-F-A-D-T-PN-E-BD-T-BL-D-T-Y-PN-E-BP-BP-BY-E-D-B

L-D-T-PN-E

D-TP-Y-L-PN-B

F-A-D-EF-Y-EP-APN

P-Y

P-PN-BP-PN-YY-D-BY-B

T-BD-EF-A-B-D-T-Y-PN-EP-PN-L-F-A-B-D-T-Y-EP-L-F-A-B-D-T-Y-PN-EP-PN-Y-EP-D-T-Y-PN-E-BT-BP-D-Y-PN-E

P-D-T-Y-PN-EP-D-T-Y-PN-EY(?)

P-Y-BP-BP(?)P-Y-B

P-YP-PN-Y-E

L-BP-PN-Y-E

P(?)P-L-F-A-D-T-Y-EP-PN-D-T-Y-E

P(?)

A ThscapA ThscapA ThscapP HHA ThscapA Thscap

Fr Chfrut

P HscapP Hscap

A ThparA ThparA ThparA Thpar

A ThparA ThparA ThparA ThparB GparA Thpar

P HrosP HrosB HrosP HrosA ThrosP ThcaespA HrosA ThrosA Thros

Fr NscandFr NscandFr Nscand

A ThscapA ThscapA ThscapA ThscapA ThscapP ThscapP Hscap

P HscapA Thscap

A ThscapA ThscapA ThscapB HcaespP Hcaesp


Pterocephalus papposus (L.) CoulterTremastelma palaestinum (L.) Janchen

CampanulaceaeCampanula celsii A. DC.C. rupestris Sibth. & Sm.C. drabifolia Sibth. & Sm.C. drabifolia Sibth. & Sm. subsp. drabifoliaAsyneuma limonifolium (L.) JanchenLegousia speculum -veneris (L.) FischL. falcata (Ten.) FritschL. hybrida (L.) Delarbe

CompositaeBellis annua L.B. perennis L.B. hybrida Ten.Conyza canadensis (L.) Cronq.Logfia gallica (L.) Cosson & Germ.Filago arvensis L.Evax pygmaea (L.) Brot.Helichrysum stoechas (L.) Moench subsp.barrelieri (Ten.) NymanPhagnalon graecum Boiss. & Heldr.Dittrichia viscosa (L.) W. GreuterD. graveolens (L.) W. GreuterInula candida (L.) Cass.I. candida (L.) Cass. var. methaneaI. candida (L.) Cass. var. parnesiaI. verbascifolia (Willd.) Hausskn. subsp.methanea (Hausskn.) TutinPulicaria odora (L.) Reichenb.Pallenis spinosa (L.) Cass.P. spinosa (L.) subsp. microcephala (Ha!.)Rech. fil.Xanthium strumarium L.X. spinosum L.AMhemis orientalis (L.) DegenA. orientalis (L.) Degen var. montana HayekA. orientalis (L.) Degen var. cronia HayekA. tomentosa L.A. tomentosa L. subsp. tomentosaA. tenuiloba R. Femandes subsp. cronia(Boiss. & Heldr.) R. FemandesA. arvensis L.A. chia L.A. tinctoria L.A. cotula L.Achillea ligustica All.Chamomilla recutita (L.) RauschertChrysanthemum segetum L.C. coronarium L.Leucanthemum vulgare Lam.Doronicum orientale Hoffm.Senecio vulgaris L.S. vemalis Waldst. & Kit.S. macedonicus Griseb.

P-YP-PN-Y

P-PN-L-Y-EP-A-D-T-Y-PN-EP-D-T-Y-EA-TP-B

P-Y-BP(?)

P-L-A-D-T-PN-E-BP-B-T-PN

L-F-A-T-B-Y-PN-EY(?)P-PN-YP-Y-B

P-L-B-D-T-Y-PN-EL-F-A-D-T-Y-PN-E-BL-D-T-Y-PN-E-BD-T-Y-PN-E-BP-Y-BP(?)P(?)

P-L-YP-E-BL-F-A-D-T-Y-E-B

D-TT-BD-T-Y-PN-E-BP-D-T-PN

D-T-PN-EP-T-Y

P-PNY-BP-PN-L-D-T-E-BD-T-PN-E-BP-BP-PN-D-TP-L-F-A-B-D-T-Y-PN-EF-A-B-D-T-PN-EL-F-A-B-D-T-PN-ET-PNP-YP-L-F-A-B-D-T-Y-PN-EP-Y-BP(?)

A ThscapA Thscap

B ThscapB HcaespA ThscapA ThscapP HscapA ThcaespA ThcaespA Thscap

A HrosP HrosA HrosA ThscapA ThscapA ThscapA Thscap

P ChsuffrP ChsuffrP ChsuffrA ThscapFr ChsuffrFr ChsuffrFr Chsuffr

P HscapP HscapB Hscap

B HscapA ThscapA ThscapP ThscapP ThscapP ThscapA ThscapA Thscap

A ThscapA ThscapA ThscapB HscapA ThscapP HscapA ThscapA ThscapA ThscapP HscapP GrhA ThscapA ThscapA Thscap

251Lagascalia 17(2), 1994

Calendula arvensis L. Y-P-L-F-A-B-D-T-PN-E A ThscapC. arvensis L. var. aegyptiaca Hayek Y(?) A ThscapCarlina corymbosa L. subsp. graeca (Boiss.)Nyman B-D-T-Y-PN-E B HscapAtractylis gummifera L. P-PN-B P HrosA. cancellata L. P-L-F-A-T-Y-E-B A ThscapEchinops graecus Miller L-D-T-Y-PN-B P HscapE. sphaerocephalus subsp. albidus (Boiss.& Spruner) Kozuhanov P-Y-E P HscapE. microcephalus Sibth. & Sm. D-T-PN-E-L P HscapE. spinosissimus Turra subsp. bithynicusKozuhanov Y(?) P HscapXeranthemum inapertum (L.) Miller Y-B A ThscapJurinea mollis (L.) Reincheb. P-L-Y P HscapCarduus tmoleus Boiss. subsp. armatus(Boiss. & Heldr.) Franco P-PN P ThscapC. pycnocephalus L. P-F-A-D-T-PN-E-B A ThscapCirsium palustre (L.) Scop. PN B HscapC. creticum (Lam.) D' Urv. P-L-F-A-D-T-Y-PN-E A ThscapC. arvense (L.) Scop. P-T-PN P HscapC. lanceolatum (L.) Scop. P A ThscapC. vulgare (Savi) Ten. F-A-Y A ThscapPtilostemon chamaepeuce (L.) Less. E P ChsuffrP. chamaepeuce Ten. var. muticum Hayek Y(?) P ChfrutCynara cardunculus L. T-B P HscapPicnomon acarna (L.) Cass. P-F-A-PN-E-B A ThscapNotobasis syriaca (L.) Cass. L-E A ThscapOnopordon illyricum L. P-Y-PN-E-B B HscapSilybum marianum (L.) Gaertner F-A-B-D-T-E A ThscapCentaurea raphanina Sibth. & Sm. subsp. mixta(DC.) Runemark P-L-F-A-D-T-Y-PN-E P HrosC. attica Nyman P-PN P HscapC. attica Nyman subsp. attica P-F-A-D-T-Y-PN-E P HscapC. spinosa L. subsp. spinosa D-T-PN-E P ChsuffrC. orphanidea Heldr. & Sart. PN-B P HscapC. solstitialis L. P-L-Y-E-B A ThscapC. affinis Frisv. P(?) P HscapC. iberica Trev. P B HscapC. pindicola Griseb. P-Y P HscapC. cyanus L. P A ThscapCrupina crupinastrum (Moris) Vis. P-L-Y-E-B A ThscapCarthamnus lanatus L. P-B-D-T-PN-E A ThscapScolymus hispanicus L. E-B B HscapCnicus benedictus L. P-B A ThscapCichorium intybus L. P-L-F-A-D-T-Y-PN-E-B B HscapC. spinosum L. L-Y P HscapTolpis umbellata Bertol. Y-B A ThscapRhagadiolus stellatus (L.) Gaertner P-F-A-Y-E-B A ThscapHypochoeris achyrophorus L. PN-Y-P-L A ThscapH. cretensis Bory & Chaub. Y-B P HscapH. radicata L. P-B P HrosPicris pauciflora Willd. P-PN-L-Y-E-B A ThscapScorzonera crocifolia Sibth. & Sm. Y-E-B P HscapS. mollis M.B. P-Y P HscapS. lanata (L.) Hoffm. P-L-F-A-Y-E P Hros


S. cana Hoffm. P-Y P HscapLeontodon crispus Vill. L-Y-PN P HrosL. tuberosus L. P-Y P HrosTragopogon porrifolius L. P-L-F-A-D-T-Y-PN-E-B B HscapReichardia picroides (L.) Roth F-A-Y-E-L-B P HscapSonchus oleraceus L. P-L-F-A-B-D-T-Y-PN-E A ThscapS. tenerrimus L. Y-B A ThscapSteptorhamphus tuberosus (Jacq.) Grossh. P-PN-Y-E P HscapLagoseris sancta (L.) K. Maly subsp. bifida(Vis.)Thell. P-L A ThscapLactuca serriola L. P-L-F-B-D-T-PN-E-Y B HscapL. virosa L. F-A B HscapL. viminea (L.) J. & C. Presl. P-Y B HscapL. saligna L. Y-B A ThscapTaraxacum megalorrhizon (Forskal) Hand.-Mazz. P-Y-PN-E-B P HrosT. officinale Weber P-D-T-Y-PN-E P HrosT. laevigatum DC. P-PN-Y P HrosLapsana communis L. D-T-PN A ThscapUrospermum picroides (L.) Scop. P-Y-L-B A ThscapChondrilla ramosissima Sibth. & Sm. L-F-A-Y-E-B P HscapCrepis foetida L. P-PN-B A ThscapC. foetida L. subsp. foetida D-E-T-B A ThscapC. foetida L. subsp. commutata (Sprengel)Babcock Y(?) A ThscapC. rubra L. P-PN A ThscapC. dioscoridis L. P-Y A ThscapC. neglecta L. P-F-A-B A ThrosC. neglecta L. subsp. fuliginosa (Sibth. & Sm.)Vierh. Y(?) A ThrosC. bulbosa L. P-PN-Y-B P GbulbC. fraasii Schultz Bip. P-L-Y P HrosC. vesicaria L. subsp. vesicaria P-D-T-Y-PN P ThscapC. micrantha Czerep. P-E A ThscapC. incana Sibth. & Sm. P- A ThrosC. pannosum Boiss. P-PN A ThscapTussilago fmfara L.

P-D-B P HrosMONOCOTYLEDONESPotamogetonaceae

Zostera noltii Hornem. Y P HHLiliaceae

Merendera attica Boiss. & Spruner P-Y-E P GbulbColchicum pusillum Sieber. E P GbulbC. bivonae Guss. P-D-T-Y P GbulbC. cupanii Guss. P-PN-D-Y P GbulbC. neapolitanum (Ten.) Ten. Y P GbulbAsphodelus fistulosus L. L-F-A-D-T-Y-PN-B P HscapA. aestivus Brot. P-Y-E-L-F-A-D-T-PN-B P GrhAsphodeline lutea (L.) Reinheb. P-PN-T-Y P GrhGagea peduncularis (Presl) Pascher Y-E P GbulbG. arvensis (Pers.) Dumort. Y-E P GbulbG. pusilla (F.W. Schmidt) Schultes &Schultes fil. P-D-T P GbulbG. lutea (L.) Ker. -Gawler B-T-PN P GbulbG. reticulata (Pallas) Schultes & Schultes fil. P-T-PN P GbulbG. graeca Boiss. & Spruner P-L-F-A-D-T-Y-PN-E P Gbulb

Lagascalia 17(2), 1994 253

P-PN-Y-EP(?)P-Y-BP-BY(?)

D-Y-E-B

P-PN-T

F-A-D

G. amblyopetala Boiss. & Heldr.G. bohemica (Tauschner) Schultes & Schultes filLillium heldreichii Freyn*L. candidum L.Fritillaria graeca Boiss. & GaertnerF. obliqua Ker.-GrawlerOmithogalum collinum Guss.O. sibthorpii W. GreuterO. comosum L.O. orthophyllum Ten.O. atticum Boiss. & Heldr.O. narborense L.O. umbellatum L.O. nanum Sibth. & Sm.O. nutans L.Bellevalia ciliata (Cyr.) NeesUrginea maritima (L.) Baker*Hyacinthus orientalis L.Scilla autumnalis L.Strangweia spicata (Sibth. & Sm.) Boiss.Muscari comosum (L.) MillerM. commutatum Guss.M. pulchellum Heldr. & Sart.M. weisii Freyn var. sartorianum Heldr.Allium roseum L.A. neapolitanum Cyr.A. stamineum Boiss.A. scorodoprasum L. subsp. rotundum (L.) StearnA. sphaerocephalon L. subsp. arvense (Guss.)ArcangeliA. guttatum Steven subsp. sardum (Moris) StearnA. chamaespathum Boiss.A. subhirsutum L.A. ampeloprasum L.A. pallens L. subsp. pallens (L.) Regl.Asparagus acutifolius L.A. aphyllus L.Ruscus aculeatus L.Smilax aspera L.Tulipa orphanidea Boiss.T. patens Roemer & SchultesT. beotica Boiss. & Heldr.

AmaryllidaceaeStembergia lutea (L.) Ker-Gawler subsp. sicula(Tinco) D.A. WebbS. colchiciflora Waldst. & Kit.Narcissus tazetta L.N. serotinus L.Galanthus nivalis L.

Agavaceae*Agave americana L.

DioscoreaceaeTamus communis L.

IridaceaeIris pumila L.

GbulbGbulbGbulbGbulbGbulbGbulbGbulbGbulbGbulbGbulbGbulbGbulbGbulbGbulbGbulbGbulbGbulbGbulbGbulbGbulbGbulbGbulbGbulbGbulbGbulbGbulbGbulbGbulb

GbulbGbulbGbulbGbulbGbulbGbulb

Fr ChscandFr ChscandFr ChfrutFr Nscand

GbulbGbulbGbulb

GbulbGbulbGbulbGbulbGbulb

Ar Orad

P Grh

P Grh

P-T-PNB-T-PNP-T-Y-PN-E-BP-T-BPNF-A-D-T-PNP-F-A-D-T-Y-PN-ED-TP-PN-T-L-F-A-Y-EP(?)P-A-EP-PN-T-EP-F-A-PN

D-Y-E-BPNP-D-Y-EY-EP-L-F-A-D-T-Y-PN-E-BF-A-D-T-Y-PN-E

D-T-Y-PN-EF-A-D-T-Y-PN-E-BP-T-Y-EPN

T-PNP-PN-T-YL-YP-PN-T-D-YF-A-D-Y-E-BP-YP-L-F-A-D-T-Y-PN-E-BL-D-T-Y-PN-E-BP-F-D-T-Y-PN-EP-F-A-D-T-Y-PN-EP-PNP-YP(?)


I. pumila L. subsp. atticaI. sintenissi Jka.Gynandriris sisyrinchium (L.) Par!.Romulea linaresii Pan.R. columnae Sebastian & Maur.R. ramiflora Ten.Crocus creweii Hooker fil.C. sieberi Gay.C. cartwrightianus HerbertC. laevigatus Bory & Chaub.C. cancellatus HerbertC. olivieri GayHermodactylus tuberosus (L.) MillerGladiolus segetum Ker-GawlerG. illyricus KochG. italicus Miller

JuncaceaeJuncus compressus Jacq.J. maritimus Lam.J. subnodulosus SchrankJ. articulatus L.J. acutus L.Luzula nodulosa E.H.F. MeyerL. forsteri (Smith) DC.

GramineaeFestuca pratensis Hudson subsp. pratensisF. arundinacea SchreberF. rupicola Heuffel subsp. rupicolaLolium perenne L.L. rigidum Gaudin subsp. lepturoides (Boiss.)Senen & MauricioL. temulentum L.Desmazeria rigida (L.) TutinD. rigida (L.) Tutin subsp. rigidaVulpia ciliata DumortPoa annua L.P. bulbosa L.P. bulbosa L. var. timoleontis HayekP. pratensis L.P. trivialis L. subsp. sylvicolaDactylis glomerata L.D. glomerata L. subsp. hispanica (Roth.) NymanCynosurus echinatus L.C. elegans Desf.Lamarckia aurea (L.) Moench.Psilurus incurvus (Gouan) Schinz & Thell.Briza media L.B. maxima L.B. minor L.B. humilis Bieb.Eragrostis megastachya (Koeler) LinkMelica minuta L.M. ciliata L.Bromus sterilis L.B. fasciculatus C. Presl.

P-Y-EP-YP-BP-T-YY-E

Y(?)P-F-A-T-PN-ET-Y-PN-EP-D-T-Y-PN-EP-D-T-Y-PN-EP(?)

P-T-PN-Y-E-BP-T-PNPN

E-BEP(?)P(?)PNP-T-Y-E

T-PNP-T-PNP(?)L-F-A-T-B-D-Y-E

Y(?)L-Y-E-BP-T-Y-E-LF-AP-PN-LP-L-F-A-B-D-T-Y-PN-EP-D-T-Y-PN-E-BP(?)P-Y-BP-Y-BP-L-F-A-T-B-D-Y-E-PNT-PN-BP-L-F-A-B-D-T-Y-PN-EP-PN-T-Y-BL-DY-BP-PN-L-B-D-TP-F-A-D-T-Y-PN-E-BP-F-A-T-PN-BP-PN-Y-EP-L-Y-BP-T-PN-YP-L-T-PNP-L-F-A-B-D-T-Y-PN-EP-T-PN-Y

P GrhP GrhP GbulbP GbulbP GbulbP GbulbP GbulbP GbulbP GbulbP GbulbP GbulbP GbulbP GbulbP GbulbP GbulbP Gbulb

P GrhP GrhP GrhP HcaespP GrhP HcaespP Hcaesp

P HcaespP HcaespP HcaespP Hcaesp

A ThcaespA ThcaespA ThcaespA ThcaespA ThcaespA HcaespP HcaespP HcaespP HcaespP HcaespP HcaespP HcaespP ThcaespA ThcaespA ThscapA ThcaespA ThcaespA ThcaespA ThcaespA ThscapA ThcaespP HcaespP HcaespA ThcaespA Thscap

Lagascalia 17(2), 1994 255

B. tectorum L.B. madritensis L.B. rubens L.B. intermedius Guss.B. rigidus Roth.B. squarrosus L.B. scoparius L.Brachypodium pinnatum (L.) Beauv.B. retusum (Pers.) Beauv.B. sylvaticum (Hudson) Rehm. & Schultz.Hordeum crinitum (Schreb.) Desf.H. bulbosum L.H. murinum L.

Elymus pyracanthus (Gordon) MelderisE. repens (L.) GouldAegilops triuncialis L.A. geniculata RothA. triaristata Willd.A. comosa Sibth. & Sm.A. comosa subsp. heldreichi Eig.

*Triticum turgidum L.Helictotrichon convolutum HenrardAvena clauda Durieu

• A. barbata Pott ex LinkA. fatua L.A. sterilis L.*A. sativa L.Arrhenatherum elatior (L.) Beauv.Lophochloa cristata (L.) Hyl.Deschampsia flexulosa (L.) Trin.Lagurus ovatus L.Aira elegantissima Schur.Anthoxanthum odoratum L.Holcus lanatus L.Polypogon monspeliensis (L.) Desf.P. maritimus Willd.Phleum graecum Boiss. & Heldr.P. arenarium L.P. subulatum (Savi) Ascherson & GraebnerP. echinatum Host.Phalaris arundinacea L.P. minor Retz.Piptatherum miliaceum (L.) CossonP. coerulescens (Desf.) Beauv.Milium vernale Bieb.Stipa fontanesii Par!.S. capensis Thunb.S. bromoides (L.) DorfllerS. tortilis Desf.Arundo donax L.Phragmites australis (Cav.) Trin.Cynodon dactylon (L.) Pers.Digitaria sanguinalis (L.) Scop.

P-Y-BP-PN-L-F-A-T-Y-E-BP-L-B-D-T-Y-PN-EL-F-A-D-T-Y-PN-E-BP-PN-T-L

P-T-BT-Y-D-BY(?)P-DY-EP-YP-L-F-A-B-D-PN-E-T

P-PNL-Y-EB-T-PNP-L-F-A-T-PN-EY-E-B

APN-BP-PN-YP-T-YP-F-A-D-T-Y-PN-EF-A-B-T-PNP-L-F-A-B-T-Y-PN-ED-TYP-L-F-A-B-D-Y-EL-TP-L-F-A-B-D-T-Y-PN-EP-F-A-D-T-Y-PN-E-BP-D-T-BD-T-PN-BL-YD-T-BP-Y-BP-PN-Y-BP-P-BP-B-TP-L-B-YP-L-F-A-B-D-T-Y-PN-EL-F-B-T-Y-PN-EP-YP-L-T-Y-PN-E-BD-TP-D-Y-EP-L-Y-EY-E-D-BPN-BP-L-F-A-B-D-T-Y-PN-EP-Y-E

A ThscapA ThcaespA ThcaespA ThcaespA ThcaespA ThcaespA ThcaespP HcaespP HcaespP HcaespA ThcaespP HcaespA ThcaespA ThcaespP GrhP GrhA ThcaespA ThcaespA ThcaespA Thcaesp

ThcaespA ThscapP HcaespA ThscapA ThcaespA ThcaespA ThcaespA ThcaespP HcaespA ThcaespP HcaespA ThcaespA ThcaespP HcaespP GrhA ThcaespA ThcaespA ThscapA ThcaespA ThscapA ThscapA ThcaespA ThcaespP HcaespP HcaespA ThscapP HcaespA HcaespP HcaespA ThcaespP GrhP GrhP GrhA Thcaesp

H. murinum L. subsp. leporinum (Link) Arcangeli Y(?)

256 Contribution ro (he study of (he flora of Attica (Greece)

Sorghum halepense (L.) Pers. P-B-PN P HcaespEchinochloa crus-galli (L.) Beauv. Y-E-B A ThcaespPaspalum paspalodes (Michx) Scribner B P GrhSetaria verticillata (L.) Beauv. P-D-Y-PN-B A ThcaespS. viridis (L.) Beauv. F-A-D-T-Y-PN-E-B A ThcaespHyparrhenia hirta (L.) Stapf L P Grh

AraceaeArum italicum Mill. F-A-D-T-PN-E P GrhA. maculatum L. T-PN P GrhBiarum spruneri Boiss. P-D-Y-E-B P GrhArisarum vulgare Targ.-Tozz. P-D-E P GrhA. vulgare Targ.-Tozz. subsp. vulgare F-A P GrhDracunculus vulgaris Schott P-L-D-T-Y-PN-E P Grh

CyperaceaeCyperus rotundus L. B-T-PN-Y P GrhScripus cernuus Vahl P-T P HcaespS. setaceus L. P A ThscapCarex distachya Desf. P-T-Y-E-PN-D P HcaespC. divisa Hudson P-B P HcaespC. distans L. P-Y P HcaespC. flacca Schreber subsp. flacca P(?) P GrhC. illegitima Cesati Y(?) P HcaespC. hallerana Asso Y P HcaespC. macrolepis DC. Y P HcaespC. hispida Willd. T-Y-PN-E P Hcaesp

OrchidaceaeOrchis quadripunctata Cyr. P-L-A-D-T-Y-E P GbulbO. laxiflora Lam. P-D-T-Y-PN-E P GbulbO. laxiflora Lam. subsp. palustris (Jacq.)Bonnier & Layens PN P GbulbO. papilionacea L. P-PN-Y-B P GbulbO. simia Lam. Y(?) P GbulbO. provincialis Balbis subsp. provincialis P-Y P GbulbO. provincialis Balbis subsp. pauciflorus P-Y P GbulbO. sulphurea (Link) Franco subsp.pseudosambucina (Ten.) Y(?) P GbulbAnacamptis pyramidalis (L.) Richard E-P-PN-Y P GbulbNeotinea maculata (Desf) Steam. E P GbulbSerapias parviflora Pan. P-Y-E P GbulbS. lingua L. P-D-T-Y-PN-E P GbulbS. vomeracea (Burm) Briq. Y(?) P GbulbS. vomeracea (Burm.) Briq. subsp. laxifloraGolz & Reinchard L P GbulbOphrys apifera Hudson E-PN-B P GbulbO. heldreichii Schlechter E P GbulbO. lutea (Gouan) Cav. P-D-T-Y-PN-E P GbulbO. spruneri Nyman P-Y P GbulbO. reinholdii Spruner Y P GbulbO. scopolax Cav. subsp. oestifera (Bieb.) Soo P-PN-Y P GbulbO. fusca Link P-D-T-PN-E P GbulbO. bombyliflora Link D-T-PN P GbulbAceras anthropophorum (L.) Aiton fil. P-Y P GbulbBarlia robertiana (Loisel) W. Greuter P-Y P GbulbCephalanthera damasonium (Miller) Druce P-Y P GrhLimodorum abortivum (L.) Swartz P-Y P Grh

CONTRIBUTION TO THE STUDY OF THE FLORA OF ATTICA …institucional.us.es/revistas/lagascalia/17.2/03 sarlis.pdfCONTRIBUTION TO THE STUDY OF THE FLORA OF ATTICA (GREECE) G. P. SARLIS

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