1 Volume 8 10 January 2007 Issue 1 TORTS Newsletter of the Troop of Reputed Tortricid Systematists MODIFICATIONS TO TODD GILLIGAN’S WEBSITE - TORTRICID.NET The following message was received from Todd Gilligan regarding his website - Tortricid.net. “Just a quick note to mention that I have finished a complete redesign of my website! The new site has been uploaded and should be fully functional. You can still access it using either of these URLs: http://www.tortricidae. com or http://www.torticid.net. Some of the new features include a complete redesign including new graphics, menus, fonts, etc.; the ability to search the photo database for any genus/species or wildcard text; the ability to display a complete list of thumbnail photos from the database; and new type specimen photos, including the complete type collection at ANIC (finally!) Let me know if you have any comments/questions/changes you would like to see. Thank you for your time!” If you have not yet visited Todd’s updated website, please do so - I’m sure you will be impressed. It is an outstanding example of the type of information and images that can be made available globally on the web. Also on the site is a review of our initiative to develop a resource for molecular analyses. TORTS NEWSLETTER DISTRIBUTED AS PDF Since 2005 the newsletter has been distributed primarily via e-mail as a PDF. Several members still receive a hard-copy via regular mail owing to problems receiving attachments. If you have been receiving a hard copy and prefer to receive a PDF, please let me know. Alternatively, if you have been receiving a PDF and would prefer a hard copy, let me know. Please check your e-mail address in this issue for accuracy. If you have corrections, please provide them to me at jbrown@sel. barc.usda.gov. _____________________________________ BOOK REVIEW The following book review will appear in the spring issue of the Proceedings of the Entomological Society of Washington. However, I am so eager to let everyone know about the book and to encourage everyone to order a copy, that I am providing this sneak preview here. Olethreutinae Moths of Australia, Monographs on Australian Lepidoptera, Volume 10, by Marianne Horak, with contributions by Furumi Komai. 528 pages, 984 black-and-white photographs; 7" x 10"; ISBN 0-643-09093-2. AU $160.00 (ca. $120.00 US)
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1
Volume 8 10 January 2007 Issue 1
TORTSNewsletter of the
Troop of Reputed Tortricid Systematists
MODIFICATIONS TO TODDGILLIGAN’S WEBSITE -
TORTRICID.NET
The following message was receivedfrom Todd Gilligan regarding his website -Tortricid.net.
“Just a quick note to mention that I havefinished a complete redesign of my website!The new site has been uploaded and should befully functional. You can still access it usingeither of these URLs: http://www.tortricidae.com or http://www.torticid.net. Some of thenew features include a complete redesignincluding new graphics, menus, fonts, etc.; theability to search the photo database for anygenus/species or wildcard text; the ability todisplay a complete list of thumbnail photosfrom the database; and new type specimenphotos, including the complete type collectionat ANIC (finally!)
Let me know if you have anycomments/questions/changes you would like tosee. Thank you for your time!”
If you have not yet visited Todd’supdated website, please do so - I’m sure youwill be impressed. It is an outstanding exampleof the type of information and images that canbe made available globally on the web. Also onthe site is a review of our initiative to develop aresource for molecular analyses.
TORTS NEWSLETTERDISTRIBUTED AS PDF
Since 2005 the newsletter has beendistributed primarily via e-mail as a PDF.Several members still receive a hard-copy viaregular mail owing to problems receivingattachments. If you have been receiving a hardcopy and prefer to receive a PDF, please let meknow. Alternatively, if you have been receivinga PDF and would prefer a hard copy, let meknow. Please check your e-mail address in thisissue for accuracy. If you have corrections,please provide them to me at [email protected]._____________________________________
BOOK REVIEW
The following book review will appearin the spring issue of the Proceedings of theEntomological Society of Washington.However, I am so eager to let everyone knowabout the book and to encourage everyone toorder a copy, that I am providing this sneakpreview here.
Olethreutinae Moths of Australia,Monographs on Australian Lepidoptera,Volume 10, by Marianne Horak, withcontributions by Furumi Komai. 528 pages, 984black-and-white photographs; 7" x 10"; ISBN0-643-09093-2. AU $160.00 (ca. $120.00 US)
For over two decades, Marianne Horakhas been one of the world’s leading experts intortricid systematics, with seminal contributionsto van der Geest and Evenhuis’ Tortricid Pests,their Biology, Natural Enemies and Control(Horak 1991, Horak and Brown 1991), toNeilsen, Edwards, and Rangsi’s Checklist ofAustralian Lepidoptera (Horak et al. 1996), andto Kristensen’s Handbook of Zoology (Horak1998). Her systematic work is thorough, hermorphological investigations are meticulous,and her knowledge of the world fauna isunparalleled. Her recently published monographon Australian Olethreutinae, the subject of thisbook review, is the crowning achievement in ahighly productive career focused primarily onTortricidae.
Most of us tortricid systematistsimpatiently apply our craft to taxa and regionsof various sizes, inevitably leaving a trail ofpapers, each of which addresses a differentspecies group, genus, country, or geographicregion, but rarely fully treating an entire faunaor larger taxon in a single tome. Horak hasshown the patience and perseverance tocomplete a thorough taxonomic study of theentire subfamily Olethreutinae for the entirecontinent of Australia. In this volume shereviews the 90 olethreutine genera and 249described species (and provides comments onan additional 200 undescribed species)occurring in Australia. Because much of thefauna is derived from that of the OrientalRegion, biogeographically and phylogenetically,the treatment has significant impact on theclassification of the Olethreutinae of a
geographic area much broader than Australiaalone. Each genus is described in detail,diagnosed and defined by synapomorphies, anddiscussed in the context of biogeography andphylogenetic relationships; in addition, all theconstituent species of the Australian fauna arelisted.
The text of this monograph is presentedin 8 major sections, not including the standardfront material (i.e., abstract, introduction,materials and methods, and acknowledgments):1. Phylogeny of the Olethreutinae, 2.Morphology, 3. Biology, 4. Diversity andDistribution, 5. Australian OlethreutinaeGenera, 6. References, 7. Appendices, and 8.Index.
The first section, Phylogeny of theOlethreutinae, presents a detailed list of 126morphological characters for 73 of the includedgenera plus two outgroups, along with theresults of a cladistic analysis using WinClada.As Horak concludes, the results portrayed in theconsensus tree should be viewed as preliminaryand somewhat inconclusive since severalwidely accepted groupings based onsynapomorphies are not recovered in theanalysis. It is likely that additional charactersare required to more accurately identifyrelationships among the genera. Nonetheless,the data matrix represents an important firststep in the development of a phylogeny for theAustralian and Oriental members of thesubfamily upon which a stable classificationcan begin to be built.
The Morphology section is thorough andrich in details. The 20 pages of wing venationprovide an outstanding visual summary of thevariation in shape, venation, and malesecondary structures found in the Australian and
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Oriental Olethreutinae. The Biology sectionpresents a brief overview of tortricid lifehistories and larval food plants. And theDiversity and Distribution section likewisepresents a brief analysis of different olethreutinetribes within Australia and comparisons withother well-documented faunas.
Section 5, Australian OlethreutinaeGenera, is the meat of the monograph, withover 400 pages of diagnoses, descriptions, andillustrations of the included taxa. For each tribethere are “status remarks,” “evidence formonophyly,” “diagnosis,” “distribution,”“biology,” and “remarks.” The text in thissection reveals the depth of Horak’s knowledgeof the fauna. Included are the kinds of detailsand factoids that experts accumulate over acareer of working on a taxon but seldom findthe right medium in which to convey them. Ofparticular interest to me are the myriad host-plant records based on Horak’s personalexperience, the card file at CSIRO (compiledprimarily by the late I. F. B. Common), and thework of her colleagues in Australia andelsewhere. In this section she describes 12 newgenera and 16 new species along withproposing 121 new combinations (41 for theAustralian fauna and 80 for non-Australianspecies). For the tribe Grapholitini, Horakenlisted the assistance of the highly regardedJapanese tortricid taxonomist Furumi Komai,and the results are outstanding.
The final three sections, References,Appendices (morphological character matrixand host plants by tortricid genus), and Index,are thorough as one would expect in aprofessional taxonomic treatment. While thelast two are listed in the Table of Contents assections “7” and “8,” respectively, their title
pages in the text lack the numbering systemused for other major sections - an extremelytrivial format oversight that does not detractfrom the presentation.
Like other CSIRO publications, thephysical aspects of the book are very good -strong binding, high quality paper, and nowasted space. Fonts are highly legible, headingsare clear and well defined, and illustrations arenicely organized, distributed throughout the textrather than clustered at the back or middle.
This book is in a vein similar toDiakonoff’s (1973) classic South AsiaticOlethreutini, and it is destined to become thenew reference for all systematic work onOlethreutinae of Indo-Australia and southernAsia. But it has broader geographic implicationsas well, because many of the treated genera aredistributed on various continents around theglobe: e.g., Megalota, with its Oriental-southerncontinental distribution; Crocidosema, with itsnearly cosmopolitan distribution; and Spilonota,with its Palaearctic-Oriental-Australian range.This book is undoubtedly one of the mostsignificant and original contributions to tortricidsystematics to appear in the last few decades,and its impact likely will be felt by generationsof tortricid systematists. Anyone working ontortricid moths at the global level in any context(e.g., biogeography, biodiversity, taxonomy,host-plants, agriculture) absolutely will have toconsult this volume.
Horak is to be congratulated for hertenacity in finishing this overwhelming projectand creating a masterpiece in the process. Shehas set the bar high for her fellow tortricidtaxonomists and Lepidoptera systematists ingeneral.
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Literature Cited
Diakonoff, A. 1973. The South AsiaticOlethreutini (Lepidoptera: Tortricidae).Zoologische Monographieen van hetRijksmuseum van Natuurlijke Historie1. 699 pp.
Horak, M. 1991. 1.1 Morphology, pp. 1-22. Invan der Geest, L. P. S. & H. H.Evenhuis, eds., Tortricid Pests, TheirBiology, Natural Enemies and Control.Elsevier Science Publishers B. V.,Amsterdam.
________. 1998. Tortricoidea, pp. 199-215. InKristensen, N., ed., Lepidoptera, Mothsand Butterflies. Volume 1: Evolution,Systematics, and Biogeography.Handbook of Zoology 4 (35),Arthropoda: Insecta. Walter de Gruyter,Berlin & New York.
Horak, M. and R. L. Brown. 1991. 1.2Taxonomy and phylogeny, pp. 23-48. Invan der Geest, L. P. S. & H. H.Evenhuis, eds., Tortricid Pests, TheirBiology, Natural Enemies and Control.Elsevier Science Publishers B. V.,Amsterdam.
Horak, M., I. F. B. Common & F. Komai. 1996.Tortricidae, pp. 123-136. In Neilsen, E.S., E. D. Edwards, and T. V. Rangsi,eds. Checklist of AustralianLepidoptera. Monographs on AustralianLepidoptera 4. CSIRO, Canberra,Australia.
In the USA and Canada, the book can beordered from Antipodes Books & Beyond Ltd.;phone: (301) 602-9519; also, you can google“Antipodes Books” and query for “Horak.”
UPDATES TO THE WORLDCATALOGUE OF INSECTS,VOLUME 5, TORTRICIDAE
(LEPIDOPTERA)
In an effort to stay up-to-date withcurrent taxonomic treatments in Tortricidae andto continue to improve our developingelectronic catalog of the family, I provide a listof new species, new combinations, and newsynonymies proposed prior to 2006, mostly in2005. A bibliography also is provided insupport of the changes. If you are aware ofother additions, taxonomic changes, and/orcorrections to the catalog, please bring them tomy attention. Thanks much for your continuedsupport.
SPECIES AND GENERA OFTORTRICIDAE DESCRIBED PRIOR TO
similis Razowski & Pelz, 2005 (Exoletuncus),SHILAP Revta. Lepid. 33: 330. TL: Ecuador(Zamora-Chinchipe Province, 22 km E Loja,P.N. Podocarpus, San Francisco RangerStation). Holotype (%): SMFL.
longibarba Razowski & Pelz, 2005(Heppnerographa), Entomol. Zeit. 115: 170.TL: Ecuador (Morona-Santiago Province,Macas, Proano-Alshi, 5 km S Alshi). Holotype(&): SMFM.
monotana Razowski & Pelz, 2005(Heppnerographa), Entomol. Zeit. 115: 166.TL: Ecuador (Morona-Santiago Province,Macas, Proano-Alshi, 5 km S Alshi). Holotype(%): SMFM.
podocarpi Razowski & Pelz, 2005(Heppnerographa), Entomol. Zeit. 115: 166.TL: Ecuador (Zamora-Chinchipe Province, 22km E Loja, P. N. Podocarpus, San FranciscoRanger Stn.). Holotype (%): SMFM.
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us i t i ca Razowski & Pelz , 2005(Heppnerographa), Entomol. Zeit. 115: 166.TL: Ecuador (Loja Province, 10 km SE Loja, P.N. Podocarpus, Cajanuma Ranger Stn.).Holotype (%): SMFM.
algoana Felder & Rogenhofer to Algoforma(Razowski 2005)atalodes Meyrick to Exoletuncus (Razowski &Pelz 2005)blackmorei Obraztsov as synonym of effractana(Karsholt et al. 2005)blanditana Kuznetzov to Rhopobota(Kuznetzov 1999)chrysodetis Meyrick to Oregocerata (Razowskiand Brown 2005)Clepsimorpha as synonym of Neocalyptis(Razowski 2005)Clepsiphyes as synonym of Neocalyptis(Razowski 2005)dentiuncanae; misspelling of dentiuncana(Brown 2005)Durrantia was correctly listed as a synonym ofPhtheochroa (Brown 2005), but not recognized
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as a homonym of Durrantia (Oecophoridae) effractana Hübner as valid species (Karsholt etal. 2005)emaciatana Walsingham to Pelochrista (Wright2005)excavana Haworth, 1811, not Donovan 1794(Karsholt et al. 2005)fuscana Sheldon as synonym of effractana(Karsholt et al. 2005)indicata Diakonoff to insertae sedis (Dugdale2005)keredjana Razowski to species status(Razowski 2005)kochiana Goeze (not kochiella) as a validspecies (Karsholt et al. 2005)munda Diakonoff to Eucoenogenes (Pinkaew etal. 2005)nielseni Kawabe as synonym of blanditana(Zhang et al. 2005)pamirana Razowski to species status (Razowski2005)perproquinqua Heinrich as synonym ofemaciatana (Wright 2005)plumicornis Rothschild (in Homona) - holotypeat BMNH, not a tortricid (unpublished)praeconia Meyrick to Rubrograptis (Razowski2005)rhyparographta; misspelling of rhyparograpta(Brown 2005)stettinensis Leraut as synonym of effractana(Karsholt et al. 2005)subelectana Kawabe to Phiaris (Bae 2005)
LITERATURE IN SUPPORTOF THE CHANGES
Bae, Y.-S. 2005. Systematic study of the genusPhiaris Hübner (Lepidoptera, Tor-tricidae) from Korea and Japan, part III.
Tinea 18 (supplement 3): 124–138.Brown, J. W. 2005. World catalogue of insects.
Volume 5: Tortricidae (Lepidoptera).Apollo Books. 741 pp.
Byun, B.-K. & S.-C. Yan. 2005. Description ofa new species, records of fivepreviously unrecorded species, and re-discovery of a lost species in the genusAncylis Hübner (Lepidoptera: Tortrici-dae) from China. Zootaxa 1103: 17-26.
Dugdale, J. S. 2005. Three new species ofT r a c h o l e n a C o m m o n , 1 9 6 5(Lepidoptera: Tortricidae: Tortricinae:Schoetnotenini) from New Caledoniaassociated with Araucariaceae. Zootaxa870: 1-16.
Ferris, C. D. 2005. A new species of EucosmaHübner from the western United States(Lepidoptera: Tortricidae: Eucosmini).Zootaxa 806: 1-8.
Karisch, T. 2005. Ein neuer wickler aus Angola(Lepidoptera, Tortricidae, Olethreu-tinae). Lambillonea 105: 500-503.
Karisch, T. 2005. Uebersicht ueber die GattungDracontogena Diakonoff 1970 (Lepid-optera: Tortricidae: Olethreutinae:Grapholitini). [Overview of the genusDracontogena Diakonoff 1970(Lepidoptera: Tortricidae: Olethreu-tinae: Grapholitini).] Linzer BiologischeBeiträge 37(1): 457-476.
Karsholt, O., L. Aarvik, D. Agassiz, P. Huemer& K. Tuck. 2005. Acleris effractana(Hübner, 1799) - a Holarctic trotricid.Nota Lepidopterologica 28: 93-102.
Kuznetzov, V. I. 2005. Cochylimorpha raz-owskiana nom. n. for C. pallens (Kuz-netzov, 1966) (Lepidoptera: Cochyl-idae). Zoosystematica Rossica 14(1)
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2005: 178.Nedoshivina, S. V. and V. V. Zolotuhin 2005.
A new subspecies of Pelatea klugiana(Freyer, 1836) from the Middle VolgaRegion of Russia with notes on itsmorphology and life history (Tortric-idae). Nota Lepidopterologica 28: 3-9.
Oku, T. 2005. Some olethreutine moths(Lepidoptera: Tortricidae) from Japanconfused with or allied to other knownspecies. Tinea 18 (supplement 3): 96-114.
Pinkaew, N., A. Chandrapatya & R. Brown.2005. Two new species and a newrecord of Eucoenogenes Meyrick(Lepidoptera: Tortricidae) from Thai-land with a discussion of charactersdefining the genus. Proceedings of theEntomological Society of Washington107: 869-882.
Pooni, H. S. & H. S. Rose. 2005. Anathamnaneospermatophaga sp. nov. (Enarmoni-ini: Olethreutinae: Tortricidae) fromwestern Himalaya, India. Entomon 30:237-241.
Razowski, J. 2005. Notes and descriptions ofprimitive Tortricini from tropicalAfrica, with a list of Asian taxa(Lepidoptera: Tortricidae). SHILAP Re-vista de Lepidopterologia 33: 423-436.
Razowski, J. 2005. Review of NeocalyptisDiakonoff, 1941 (Lepidoptera: Tort-ricidae) with description of five newspecies. Polskie Pismo Entomol-ogiczne 74: 137-148.
Razowski, J. 2005. Descriptions andrectifications in Asian Cochylini(Lepidoptera: Tortricidae). PolskiePismo Entomologiczne 74: 431-438.
Razowski, J. 2005. Tortricidae (Lepidoptera)from South Africa. 1: Tortricini andCochylini. Polskie Pismo Entomol-ogiczne 74:495-508.
Razowski, J. & J. Brown. 2005. Review ofOregocerata Razowski (Lepidoptera:Tortricidae: Euliini), with descriptionsof four new species. Proceedings of theEntomological Society of Washington107: 903-913.
Razowski, J. & Pelz, V. 2005. Remarks onNeotropical Chlidanotini with a newspecies of Macrochlidia Brown, 1990and seven new species of Heppner-ographa Razowski, 1987 from Ecuador(Lepidoptera: Tortricidae). Entomol-ogische Zeitschrift (Stuttgart) 115(4):165-171.
Razowski, J. & V. Pelz. 2005. ExoletuncusRazowski, 1988 from Ecuador, withdescription of seven species(Lepidoptera: Tortricidae, Euliini).SHILAP Revista de Lepidopterologia33: 327-339.
Razowski J., Pelz V. 2005. LobogenesisRazowski, 1990 (Lepidoptera: Tor-tricidae: Euliini) from Ecuador, withdescription of ten new species. PolskiePismo Entomologiczne 74: 439-453.
Razowski, J. and V. Pelz. 2005. New species ofGorytvesica Razowski, 1997 andTranstillaspis Razowski 1987 (Lepid-optera: Tortricidae: Euliini) from Ecua-dor. Acta Zoologica Cracoviensia 48(B): 57-94.
Rose, H. S. and Pooni, H.S. 2005. Taxonomicstudies on the family Tortricidae(Tortricoidea: Lepidoptera) fromnorthwestern India - tribe Eucosmini
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(Olethreutinae). Zoos' Print Journal 20(2):1751-1765. [haven’t seen]Wang, X.-P. & H.-H. Li. 2005. A new genus
and species of Tortricinae (Lepidoptera:Tortricidae) from China. EntomologicaFennica 16: 263-265.
Wright, D. J. 2005. Some Eucosmini(Tortricidae) associated with Eucosmaemaciatana (Walsingham) andEucosma totana Kearfott; four newspecies, a new combination, and a newsynonymy. Journal of the Lepid-opterists’ Society 59: 121-133.
Wu, C. & X. Chen. 2005. First record of thegenus Lepteucosma from China, withdescription of one new species(Lepidoptera: Tortricidae). Acta Zool-ogica Cracoviensia 49(B): 79-81.
Yu, H. & H.-H. Li. 2005. First record of thegenus Rhopaltriplasia from China, withdescription of two new species(Lepidoptera: Tortricidae: Olethreu-tinae). Zootaxa 1082: 29-35.
Zhang, A.-H., H.-H. Li & S. Wang. 2005.Study on the genus Rhopobota Ledererfrom China (Lepidoptera: Tortricidae:Olethreutinae). Entomologica Fennica16: 273-286.
_____________________________________
ZOOTAXA STILL LOOKINGFOR NEW LEPIDOPTERA
SUBJECT EDITORS
Zootaxa, a rapid international journal foranimal taxonomists, is still seeking one or morequalified Lepidoptera Section editors. Ifinterested, please contact Dr. Zhang at“[email protected]”
MIXED REVIEWS OFTORTRICID CATALOGUE
I have seen a number of publishedreviews of the tortricid catalogue, and most ofthem point out a number of minor deficienciesand/or shortcomings in the product. Fortunately,most are the types of errors that can becorrected in the electronic version currentlybeing developed. Although there appear to bevery few omitted taxa and few misspellings oftaxa, there are a number of misspellings ofauthors’ names, and an inordinate number ofeither misspelled or incorrect type localities. Ifand when you encounter these types of errors,please forward them me so they can becorrected. Thanks.
Another criticism of the catalogue is thetotal loss of phylogenetic information byorganizing taxa alphabetically. Perhaps generashould have been grouped together at the triballevel, with those that are conspicuously similarplaced together, following classifications suchas those proposed by Razowski in hiscatalogues of various geographic regions -something to consider for the updated edition in25-30 years!
If you have comments and/or recom-mendations, please feel free to send them to me.It would be interesting to begin a dialogregarding the “essentials” and how to bestconvey the included information. It is possiblethat we will be able to incorporate these typesof suggestions into the current database, and inturn develop different ways of sorting the datato maximize the ease of usage and still retainthe maximum phylogenetic content. Thanks inadvance for your comments.