The vascular flora of forest and shrubland communities of Krousia and Belles Mts (NE Greece)

Page 1

343PHYTOLOGIA BALCANICA 14 (3): 343 – 365, Sofia, 2008

The vascular flora of forest and shrubland communities of Krousia and Belles Mts (NE Greece)

Georgios Fotiadis & Nikolaos Athanasiadis

Institute of Forest Botany–Geobotany, Faculty of Forestry & Natural Environment, Aristotle University of Thessaloniki, Greece, e-mail: [email protected]

Received: March 24, 2008 ▷ Accepted: July 11, 2008

Abstract. Extended parts of Krousia (Mt Disoro and Mt Mavrovouni) and Belles Mts (NE Greece) are protected by international and national conventions and laws (Ramsar Convention, Natura 2000 Network). The flora of Krousia Mts has been partly investigated, while very few floristic records exist for Mt Belles. The investigated area includes pure and mixed stands of Salix, Platanus, Alnus, Paliurus, Carpinus, Ostrya, Quercus, Castanea, Tilia, Fagus, and Abies. The present paper reports the flora of forest and shrubland habitats of the investigated area. The floristic catalogue is based on the plant species recorded in 431 relevés, as well as on a supplementary floristic inventory. For each taxon information is given about its spatial distribution, constancy and the habitat type from which it was collected. The floristic catalogue reports a total of 483 taxa. Furthermore, 439 taxa were recorded in Belles (405 new records) and 429 (five new records) in Krousia Mts (367 in Disoro and 373 in Mavrovouni).

Key words: Belles, floristic catalogue, Krousia, NE Greece, vegetation types

Introduction

Floristic information on Krousia Mts can be found in several publications (Gamisans & Hebrard 1980; Strid 1986; Strid & Tan 1991; Athanasiadis & al. 1993; Theodoropoulos & al. 1995; Babalonas 1995; Fotiadis & Athanasiadis 2003). On the contrary, Mt Belles could be regarded as floristically unexplored, since only few published data are available (Zagan-iaris 1938, 1939,1940; Strid 1986; Strid & Tan 1991). In addition to this, some floristic data can be re-trieved from the maps given in Flora Hellenica (Strid & Tan 1997, 2002). However, the data presented by the above-mentioned works about these two moun-tains are only partly informative regarding the type of habitat from which the plant specimens were col-lected.

Phytosociological studies can elucidate the vegeta-tion structure and provide ecological data. Thus a flo-ristic inventory based on phytosociological research is

provided, with important data about the species dis-tribution in different vegetation units and habitats (Spribille & al. 2001), which can further improve the management of e.g. important medicinal or endan-gered plant species. Floristic inventories of this type, especially those concerning forests and shrublands, are rather limited in Greece (e.g. Eleftheriadou 1992; Tsiripidis & Athanasiadis 2003).

This paper is aiming to present: 1) the forests and shrublands flora of Krousia and Belles region and 2) the ecological and environmental preference of each taxon. On this basis we have approached the plant va-riety and floristic diversification of different types of vegetation and, including ecological data, attempt-ed to evaluate the habitats. These investigations offer important knowledge in managing: from the protec-tion of different types of vegetation with special flo-ra composition to protection of species by means of their habitats, as well as better exploitation of forest products.

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344 Fotiadis, G. & Athanasiadis, N. • The vascular flora of Krousia and Belles Mts (NE Greece)

Investigated area

Krousia and Belles Mts are located in Central Mac-edonia (NE Greece). Krousia, which extends into the Greek territory, comprises two small mountains (Disoro and Mavrovouni), while Belles runs in east-west direction along the border of Greece, Bulgaria and F.Y.R.O.M. Krousia are located south of the val-ley that binds lake Kerkini and lake Doirani. Its high-est peak Mavrovouni reaches 1131 m. Mt Belles is lo-cated north of the same valley and its highest peak is 2031 m (Fig. 1). The northern slopes of Mavrovou-ni and the southeastern slopes of Belles are protected by the Ramsar Convention (the artificial lake of Ker-kini). Furthermore, extended parts of Krousia and Belles Mts have been included in the Greek Natura 2000 Network (GR 1230002, GR 1260001).

Krousia and Belles belong to the Serbomacedo-nian Massif. Petrographically, the area is dominated by metamorphic rocks (e.g. amphibolites, gneisses and schist). Late-tectonic and post-tectonic sedi-ments of the Pleistocene have accumulated as lacus-trine and continental deposits (IGME 1983). The climate of the area belongs to the Cfa climatic type according to Köppen’s classification and is charac-terized as humid, with long and very hot summers, mild winters, and lack of dry season (Flocas 1992). The bioclimate of the investigated area can be re-

garded as humid to subhumid, with cold to harsh winters (Mavrommatis 1980).

The forests and shrublands of the investigated area were classified into four main units representing syn-taxa in the rank of order (Populetalia albae, Pruneta-lia spinosae, Quercetalia pubescentis and Fagetalia syl-vaticae; Fotiadis 2004). Populetalia albae includes all azonal forests dominated by Platanus orientalis, Al-nus glutinosa and Salix alba, while Prunetalia spinosae includes all shrublands dominated by Paliurus spina-christi. Quercetalia pubescentis includes all shrublands dominated by Quercus coccifera and Carpinus orien-talis, as well as forests dominated mainly by Quercus pubescens, Q. frainetto, Q. petraea subsp. medwediewii and other deciduous species (Carpinus orientalis, Os-trya carpinifolia and Castanea sativa). Finally, Fageta-lia sylvaticae includes Fagus sylvatica, Tilia tomentosa and Abies borisii-regis forests. Shrublands, which be-long to Prunetalia spinosae and Quercetalia pubescen-tis, occupy 172.51 ha, while forests of Populetalia al-bae, Quercetalia pubescentis and Fagetalia sylvaticae occupy 550.04 ha (Table 1).

Material and methods

The majority of plant specimens were collected dur-ing phytosociological investigation in the Krousia Mts

and the Greek part of Mt Belles. Field work was con-ducted from May to August of 1999, 2000, 2001 and 2002 and was supplemented by collections in spring and autumn. Some extra collec-tions were made during the summer of 2003.

Collecting localities fell within the coordinates 40°58' to 41°27' N and 22°44' to 23°22' E (Fig. 1). An in-ventory of 431 phytosocio-logical relevés was evaluat-ed floristically (70 relevés in Populetalia albae, 35 relevés in Prunetalia spinosae, 242 relevés in Quercetalia pu-bescentis, and 84 relevés in Fagetalia sylvaticae). Plant Fig. 1. Map of Krousia and Greek Belles Mts, with the collecting localities.

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345Phytol. Balcan. 14(3) • Sofia • 2008

specimens were collected inside forest and shrub-land communities, whereas edges and clearings were avoided. Plant specimens outside the relevés were col-lected simultaneously. Voucher specimens are depos-ited in the herbarium of the Labolatory of Forest Bot-any / Geobotany (TAUf). Some extra floristic data of 35 relevés (Gamisans & Hebrard 1980; Theodoropou-los & al. 1995), all concerning Quercetalia pubescentis, were also included.

The entire area of investigation was subdivided for reasons of convenience into three parts: Disoro, Mav-rovouni and Belles (Fig. 1). The coordinates that con-fine the three subareas as well as their altitudinal range are given in Table 2.

Families, genera and species are arranged alphabet-ically within the four major groups of vascular plants, viz. Pteridophyta, Gymnospermae, Dicotyledoneae and Monocotyledoneae. The nomenclature follows Strid & Tan (1997, 2002), Greuter & al. (1984, 1986, 1989, 1993), Strid (1986), Strid & Tan (1991), Tutin & al. (1968-1980, 1993), and selected taxonomic literature (in the latter case the sources are cited). The authors of plant names are those provided by Brummitt & Powell (1992).

Categorizations of species constancy per subarea and phytosociological unit follow those suggested by Tsiripidis & Athanasiadis (2003):

1) for the species constancy per subarea, the terms ‘rare’, ‘scattered’ and ‘common’ apply to the species oc-currence of <10 %, 10-20 % and >20 % of the relevés per subarea, respectively, and

2) for the constancy per phytosociological unit the terms ‘very rare’, ‘rare’, ‘scattered’, ‘common’ and ‘very common’ apply to the species occurrence of <5 %, 5-10 %, 10-20 %, 20-40 %, and >40 % of the relevés per order, respectively.

Table 1. Area of different land uses and vegetation types.

Type of land uses and vegetation types Cover area (ha)

Agricultural land 404.16

Forest communities 550.04

Grassland 208.73

Quarry 14.73

Reforestation 35.55

Sand 1.91

Shrubland communities 172.51

Wetland 3.49

Total area 1391.1028

Table 2. Coordinates and altitudes of the collecting localities in the three subareas of Krousia and Greek Belles Mts.

Name of subarea

Number of collecting localities

Latitude (N) Longitude (E) Altitude (× 10 m)

min max min max min max

Belle

s

Greek Belles 257 41°13'07" 41°23'04" 22°44'58" 23°17'59" 9 150

Krou

sia

Disoro 93 41°04'61" 41°12'55" 22°48'41" 23°03'05" 9 75

Mavrovouni 116 40°58'67" 41°10'25" 22°55'26" 23°15'18" 5 104

Taxa not previously reported for Krousia or Belles (Turrill 1918, 1919, 1920, 1929; Ζaganiaris 1938, 1939, 1940; Rechinger 1939; Μoulopoulos 1965; Gamisans & Hebrard 1980; Papanicolaou & Zacharof 1980; Strid 1986; Strid & Tan 1991; Zieliński 1991; Αthanasiadis & al. 1993; Christensen 1994; Theodor-opoulos & al. 1995; Babalonas 1995; Fotiadis & Atha-nasiadis 2003) are given as new for the investigated subareas.

The floristic catalogue (Table 8) includes the fol-lowing:

found taxa (1st column); their habitats (2nd col-umn); constancy of taxa in subareas (3rd column); constancy of taxa in the defined orders (4th column); subareas where taxa are recorded for the first time (5th column); subareas where species recorded earlier by other researchers (6th column).

In the table of the floristic catalogue the following abbreviations are used:– subareas: 1 – Belles; 2 – Disoro; 3 – Mavrovouni (see

Table 2);– taxa in subareas: r –rare; s – scattered; c – common;– phytosociological orders: A – Populetalia albae

(Alnus, Platanus and Salix stands); B – Pruneta-lia spinosae (stands with Paliurus); C – Quercetalia pubescentis (Quercus, Ostrya and Castanea shrub-lands and forests); D – Fagetalia sylvaticae (Fagus, Tilia and Abies forests);

– taxa in a phytosociological order: vr – very rare; r – rare; s – scattered; c –common; vc – very common;

– authors abbreviations: G – Gamisans & Hebrard (1980); T: Theodoropoulos & al. (1995); At: Athana-siadis & al. (1993); M: Babalonas (1995); F: Fotiadis & Athanasiadis (2003); S: Strid (1986); S & K: Strid and Tan (1992).

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346 Fotiadis, G. & Athanasiadis, N. • The vascular flora of Krousia and Belles Mts (NE Greece)

Results and discussion

A total of 483 taxa are reported in the floristic catalogue (Tables 3, 8) with 439 (405 new records) of these occur-ing in Mt Belles, 367 (65 new records) in Mt Disoro and 373 (21 new records) in Mt Mavrovouni. Furthermore, 429 taxa are found all over the area of Krousia and five of them are designated as new records.

The majority of registered taxa, belongs to dicot-yledones (391), followed by monocotyledones (75) (Table 4). Likewise, the majority of families belongs to dicotyledones (59), followed by monocotyledones (7), gymnosperms (4), and pteridophytes (2).

The majority of species appear in almost all vege-tation units, both forests and shrublands (Tables 5, 8), although their constancy values may vary. Howev-er, each of the four vegetation units bears a differ-ent number of species found solely in it (Table 6). In the case of azonal forests (Populetalia albae), the occurence of these species is related to special eco-logical conditions (water springs and rivers, or even wastelands) (Prieditis 1997). Likewise, the flora of Tilia, Fagus and Abies forests (Fagetalia sylvaticae) is diversified by a number of species, the occurrence of which could be related (Bergmeier & Dimopou-los 2001) to the lack of any considerable human im-pact, the climatic conditions (higher precipitation and lower temperatures), and the dense canopy.

Although only two species are found solely in Pru-netalia spinosae (Paliurus spina-christi shrublands), the shrublands of this order are the richest from the phytodiversity point of view, considering the low number of conducted relevés (Table 6). The species richness could be attributed to the fact that they are transitional between pastures and forests, thus lying inside an ecotone characterized by rich phytodiversi-ty (Vrachnakis & al. 2005).

An altitudinal decline has been observed spe-cies richness found between the low-altitude ther-mophilous shrublands (eg. Paliurus spina-chris-ti) and forests (eg. Quercus pubescens, Q. frainetto) and high-altitude forests (eg. Tilia tomentosa, Fagus sylvatica) (Tables 7, 8). It could be explained by the fact that the former have an open canopy (Kyriazo-poulos & al. 2006) subjected to human activity (by grazing, etc.), with has an impact on a significant number of species (e.g. Asperugo procumbens, Jug-lans regia; Mucina 1997).

Table 3. Taxa recorded in the three subareas of Krousia and Greek Belles Mts.

Taxa recorded

New taxa for the area

Belles 439 405

Disoro 367 65

Mavrovouni 373 21

Krousia (Disoro and Mavrovouni) 429 5

Total area 483 5

Table 4. Floristic composition in the three subareas of Krousia and Greek Belles Mts.

families species subspeciestaxa

(species + subspecies)

Pteridophyta 2 8 3 11

Gymnospermae 4 4 2 6

Dicotylidonae 59 309 82 391

Monocotylidonae 7 60 15 75

total 72 381 102 483

Table 5. Taxa per habitat (shrublands and forests).

Habitat Number of taxa (%)

only in forests 36.03

only in shrublands 4.14

in forests and shrublands 59.83

Table 6. Taxa per phytosociological order.

Phytosociological orderNumber of taxa found only in one

order

Total number of taxa

Number of taxa per

relevé (%)

Populetalia albae 36 333 30.11

Prunetalia spinosae 2 237 42.86

Quercetalia pubescentis 52 408 9.6

Fagetalia sylvaticae 19 248 17.44

Table 7. Taxa per habitat (low and high altitudes).

Habitat Number of taxa (%)

taxa found in shrublands and forests of low altitudes 33.51

taxa found almost in all habitats 30.00

taxa found in forests of middle and high altitudes 29.46

taxa found in forests of high altitudes 7.03

Page 5

347Phytol. Balcan. 14(3) • Sofia • 2008 Ta

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6• Phytol. Balcan. 14(3) • 2008

Page 6

348 Fotiadis, G. & Athanasiadis, N. • The vascular flora of Krousia and Belles Mts (NE Greece)Ta

ble 8

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2: r

, 3: s

A: r,

C, D

: s1

2 (A

t), 3

(At)

Bora

gina

ceae

Aspe

rugo

pro

cum

bens

L.

Relat

ed to

graz

ing

1: r

A: vr

1Li

thos

perm

um p

urpu

roca

erul

eum

L.

Relat

ed to

graz

ing

1, 2,

3: r

B: r,

C, D

: vr

12

(At),

3 (A

t)M

yoso

tis a

rven

sis (L

.) H

ill su

bsp.

arv

ensis

Rare

, in th

erm

ophi

lous

fore

sts an

d sh

rubl

ands

1,

2, 3:

rA:

vr, B

: s, C

: r, D

: vr

12

(F),

3 (F

)M

. incr

assa

ta G

uss.

Relat

ed to

graz

ing

2: s,

3: r

C: r

2 (A

t), 3

(At,

M)

M. r

amos

issim

a Ro

chel

subs

p. ra

mos

issim

aRa

re, in

ther

mop

hilo

us fo

rests

and

shru

blan

ds

1: s,

2, 3

: cA:

s, B

: vc,

C: c,

D: v

r1

2 (A

t, T)

, 3 (A

t, M

, T)

M. s

ylvat

ica su

bsp.

cyan

ea (H

ayek

) Ves

terg

r.1:

r, 2

: c, 3

: sA:

s, B

: c, C

: r, D

: vr

1, 2,

3Th

e rec

ord

of M

. sylv

atica

subs

p. su

barv

en-

sis G

rau

by T

heod

orop

oulo

s & al

. (19

95) f

rom

Kr

ousia

(2 an

d 3)

may

refe

r to

the s

ubsp

. cy-

anea

. Pu

lmon

aria

mol

lis H

orne

m.

1: r

A, C

: vr

1Sy

mph

ytum

bul

bosu

m S

chim

p.M

ostly

in Q

uerc

us fr

aine

tto fo

rests

1, 2,

3: r

A: s,

C: r

, D: v

r1

2 (A

t), 3

(At,

M, T

)S.

otto

man

um F

riv.

1, 2,

3: r

A: s,

B: v

r, C,

D: r

1, 2

3 (F

)Ca

mpa

nula

ceae

Cam

panu

la li

ngul

ata

Wal

dst.

& K

it.

1, 2,

3: r

B, C

, D: v

r1

2 (A

t), 3

(At,

M)

Page 7

349Phytol. Balcan. 14(3) • Sofia • 2008 Ta

ble 8

. C

ontin

uatio

n.ta

xaha

bita

tco

nsta

ncy

in su

bare

asco

nsta

ncy

in or

ders

new

reco

rdre

cord

ed b

y ot

her a

utho

rsC.

per

sicifo

lia L

. M

ostly

in fo

rests

1: s,

2, 3

: cA:

vr, B

: r, C

, D: c

1 (S

&K)

, 2 (A

t, T)

, 3 (A

t, M

, T, F

)C.

spar

sa su

bsp.

spha

erot

hrix

(Gris

eb.)

Hay

ekM

ostly

in fo

rests

1: s,

2, 3

: cA:

vr, B

: s, C

, D: c

12

(At,

T), 3

(At,

T)C.

trac

heliu

m su

bsp.

atho

a (B

oiss

. & H

eldr.)

H

ayek

M

ostly

in fo

rests

1: s,

2: r

, 3: s

A: vr

, C: s

, D: v

c1

2 (A

t), 3

(At,

M, T

)

Cann

abac

eae

Hum

ulus

lupu

lus L

.Na

turil

ized

mos

tly in

Aln

us st

ands

1: r,

2: s

A: c,

B, D

: vr

12

(At)

Capr

ifolia

ceae

Loni

cera

capr

ifoliu

m L

.1,

2, 3:

rA:

r, C

, D: v

r1

2 (A

t), 3

(At,

T)L.

etru

sca

Sant

i 1:

r, 2

: s, 3

: cA,

B: v

r, C:

c, D

: vr

12

(At,

G), 3

(At,

M, T

)Sa

mbu

cus n

igra

L.

1, 2,

3: r

A: c,

B, C

: vr,

D: r

1, 2

3 (M

)Ca

ryop

hylla

ceae

Agro

stem

ma

gith

ago L

. sub

sp. g

ithag

oIn

Que

rcus

frai

netto

fore

sts2,

3: r

C: vr

32

(F)

Aren

aria

lept

ocla

dos (

Rchb

.) Gu

ss.

Loca

lly co

mm

on1,

2, 3:

rA,

B: s

, C: r

, D: v

r1,

23

(M)

C. b

rach

ypeta

lum

subs

p. te

nore

anum

(Ser

.) So

óRa

re, in

ther

mop

hilo

us fo

rests

and

shru

blan

ds

1: r,

2: s

, 3: r

A, B

, C: s

, D: v

r1,

23

(M)

C. fo

ntan

um su

bsp.

vul

gare

(Har

tm.)

Greu

ter &

Bu

rdet

1: r

D: vr

1 (S

)

C. gl

omer

atum

Thu

ill.

Rare

, in th

erm

ophi

lous

fore

sts an

d sh

rubl

ands

1,

2, 3:

rB:

s, C

: vr

1, 2

3 (M

)C.

pum

illum

subs

p. gl

utin

osum

(Frie

s) C

orb.

Rare

, in th

erm

ophi

lous

fore

sts an

d sh

rubl

ands

1,

2: r

B, C

, D: v

r1

2 (F

)C.

sem

idec

andr

um L

.Ra

re, in

ther

mop

hilo

us fo

rests

and

shru

blan

ds

1, 2,

3: r

A: r,

B: s

, C: v

r1,

23

(F)

Dian

thus

cory

mbo

sus S

m.

Rare

, in th

erm

ophi

lous

fore

sts an

d sh

rubl

ands

1,

2: r

B, C

: vr

12

(At)

D. cr

uent

us G

riseb

.Ra

re, in

ther

mop

hilo

us fo

rests

and

shru

blan

ds

1, 2,

3: r

B, C

: vr

12

(At),

3 (A

t, T)

D. gi

gant

eus d

’Urv

.1:

rC:

vr1

(S)

Moe

hrin

gia

trine

rvia

(L.)

Clai

rv.

Mos

tly in

Fag

us fo

rests

1, 3:

rC:

vr, D

: s1

3 (A

t, M

, T)

Moe

nchi

a m

antic

a (L

.) Ba

rtl.

Very

com

mon

in Q

uerc

us fo

rests

1:

r, 2

: c, 3

: sA:

vr, B

, C: c

, D: v

r1

2 (A

t), 3

(At,

M)

Petro

rhag

ia il

lyric

a su

bsp.

hay

nald

iana

(Jan

ka)

P.W. B

all &

Hey

wood

1, 2,

3: r

A: r,

B: s

, C: v

r1

2 (A

t), 3

(At,

T)

Sapo

naria

offic

inal

is L.

In w

et p

lace

s1,

2, 3:

rA:

r1,

23

(M)

Silen

e atro

purp

urea

(Gris

eb.)

Greu

ter &

Bur

det

Mos

tly in

Que

rcus

fore

sts

1, 2:

s, 3

: 3A:

vr, C

: s, D

: c1

2 (A

t), 3

(At,

T)S.

com

pacta

Fisc

h.

In C

asta

nea

stand

s1:

rC,

D: v

rS.

coni

ca L

.M

ostly

in sh

rubl

ands

1, 2,

3: r

A: vr

, B: r

, C: v

r1,

32

(F)

S. co

rona

ria (L

.) Cl

airv

.M

ostly

in Q

uerc

us fo

rests

1:

r, 2

: c, 3

: sA:

vr, C

, D: s

12

(At,

T), 3

(At,

M, T

)S.

italic

a (L

.) Pe

rs. s

ubsp

. ita

lica

One o

f the

mos

t com

mon

spec

ies i

n Qu

ercu

s fo

rests

1, 2,

3: c

A, B

: r, C

: vc,

D: s

12

(At,

G, T

), 3

(At,

M, T

)

S. vi

ridifl

ora

L.M

ostly

in fo

rests

1, 2,

3: r

A, C

, D: r

12

(At,

G), 3

(At,

M, T

)S.

vulga

ris su

bsp.

bos

niac

a (G

. Bec

k) G

reut

er &

al.

Very

com

mon

in fo

rests

1: c,

2: s

, 3: c

A, C

, D: c

12

(At,

G, T

), 3

(At,

M, T

)St

ellar

ia a

quat

ica (L

.) Sc

op.

In w

et p

lace

s3:

rA:

vr3

(At)

S. m

edia

(L.)

Vill.

It en

ters

into

ther

mop

hilo

us fo

rests

and

shru

blan

ds1:

c, 2

, 3: s

A, B

: c, C

: s, D

: r1,

23

(M)

Page 8

350 Fotiadis, G. & Athanasiadis, N. • The vascular flora of Krousia and Belles Mts (NE Greece)Ta

ble 8

. C

ontin

uatio

n.ta

xaha

bita

tco

nsta

ncy

in su

bare

asco

nsta

ncy

in or

ders

new

reco

rdre

cord

ed b

y ot

her a

utho

rsCe

lastr

acea

eEu

onym

us eu

ropa

eus L

. 1,

2, 3:

rA:

r, B

, C, D

: vr

12

(At),

3 (A

t)E.

latif

oliu

s Mill

. M

ostly

in fo

rests

of h

igh

altit

udes

1, 3:

rC:

vr, D

: vr

13

(F)

Cista

ceae

Cistu

s cre

ticus

L. s

ubsp

. cre

ticus

1:

r, 2

: c, 3

: rA,

B: v

r, C:

s1

2 (A

t, T)

, 3 (A

t)Co

mpo

sitae

Achi

llea

clype

olat

a Sm

. M

ostly

in fo

rests

and

shru

blan

ds o

f low

altit

udes

1: s,

2: c

, 3: s

A: r,

B, C

: c1

2 (A

t), 3

(At,

T)A.

gran

difo

lia F

riv.

Mos

tly in

fore

sts1:

rA:

r, C

, D: v

r1

A. m

illefo

lium

L. s

ubsp

. mill

efoliu

m1,

2, 3:

rA,

B: v

r, C:

r1

2 (A

t, T)

, 3 (A

t, M

)An

them

is ar

vens

is L.

subs

p. ar

vens

isM

ostly

in sh

rubl

ands

and

open

woo

dlan

ds1:

c, 2

: r, 3

: sA:

r, B

: vc,

C: c,

D: r

12

(At),

3 (A

t)A.

tinc

toria

subs

p. a

ustra

lis R

. Fer

nánd

ezM

ostly

in sh

rubl

ands

and

open

woo

dlan

ds1:

s, 2

, 3: c

A: vr

, B, C

: c, D

: vr

12

(At),

3 (A

t, T)

Arct

ium

min

us B

ernh

.In

wet

pla

ces

1: r,

2: s

, 3: r

A: c,

C: r

1, 2

3 (M

)Ar

tem

isia

vulga

ris L

.In

wet

and

dum

p pl

aces

1, 2,

3: r

A: c,

B: v

r1,

23

(M)

Belli

s per

enni

s L.

Mos

tly in

Pal

iuru

s shr

ubla

nds

1, 2,

3: r

A: vr

, B: r

1, 3

2 (F

)Ce

ntau

rea

cf a

ffini

s Friv

. sub

sp. a

ffini

s1:

r, 2

: c, 3

: rA:

vr, B

: r, C

: s1

(S&

T), 2

(At,

F), 3

(At,

T, F)

Chon

drill

a ju

ncea

L.

Rare

, in th

erm

ophi

lous

fore

sts an

d sh

rubl

ands

1, 2:

rC:

vr1

2 (A

t)Cr

epis

sanc

ta (L

.) Ba

bc.

Mos

tly in

Pal

iuru

s shr

ubla

nds

1, 2,

3: r

A: vr

, B: c

, C: v

r1,

23

(F)

Doro

nicu

m h

unga

ricum

Rch

b. f.

3: r

C: vr

3 (A

t, T)

D. or

ienta

le H

offm

.1,

2, 3:

rA,

C, D

: vr

12

(At),

3 (A

t)He

dypn

ois c

retic

a (L

.) Du

m.C

ours

. M

ostly

in P

aliu

rus s

hrub

land

s1,

3: r

B: s,

C: v

r1

3 (F

)Hi

erac

ium

bau

hini

Bes

ser

1, 2,

3: r

A: vr

, B, C

: r1

2 (A

t), 3

(At,

T)H

. cym

osum

L.

1, 2,

3: r

C, D

: vr

1 (S

), 2

(At),

3 (A

t, T,

F)H

. mac

rant

hum

(Ten

.) Te

n. su

bsp.

mac

rant

hum

Mos

tly in

ther

mop

hilo

us fo

rests

and

shru

blan

ds1:

r, 2

: s, 3

: rB:

vr, C

: r, D

: vr

12

(At),

3 (A

t)H

. mur

orum

L.

1, 3:

rC:

vr, D

: r1

3 (A

t, M

)H

. pilo

sello

ides

Vill

. sub

sp. p

ilose

lloid

es1:

r, 2

: s, 3

: rB,

C: r

, D: v

r1

2 (A

t), 3

(At,

M, T

)H

. rac

emos

um W

illd.

1,

2: r,

3: s

A, C

, D: r

12

(At,

T), 3

(At,

M, T

)H

. rec

hing

erio

rum

Zah

n M

ostly

in F

agus

fore

sts1:

s, 2

, 3: r

C: vr

, D: c

12

(At),

3 (A

t)H

. um

bella

tum

L.

1, 2,

3: r

C, D

: vr

12

(At),

3 (A

t, M

, T)

Hypo

chae

ris cr

eten

sis (L

.) Bo

ry &

Cha

ub. –

Co

rrec

t spe

lling

(vs.

‘Hyp

ocho

eris’

) acc

ordi

ng to

Gr

eute

r & al

. (19

93: 5

57)

Mos

tly in

Pal

iuru

s shr

ubla

nds

1: r,

2: s

, 3: r

A: r,

B: c

, C: r

12

(F),

3 (F

)

Lactu

ca se

rrio

la L

.1,

2, 3:

sA:

s, B

: c, C

: r, D

: vr

12

(At),

3 (A

t, M

, T)

Lactu

ca v

imin

ea (L

.) J. &

C. P

resl

subs

p. v

imin

ea

1: r,

2: c

, 3: s

C: s,

D: v

r1

2 (A

t), 3

(At,

M, T

)La

psan

a co

mm

unis

L. su

bsp.

com

mun

isCo

mm

on in

fore

sts1,

2: c,

3: s

A: c,

B: v

r, C,

D: c

12

(At,

T), 3

(At,

T)Le

onto

don

cicho

riace

us (T

en.)

Sang

uin.

Ve

ry co

mm

on in

Que

rcus

frai

netto

fore

sts

1: r,

2, 3

: cA:

r, B

: s, C

: c, D

: vr

12

(At,

T), 3

(At,

M, T

)Le

onto

don

crisp

us V

ill. s

ubsp

. cris

pus

In th

erm

ophi

lous

fore

sts an

d sh

rubl

ands

1, 2,

3: r

A: vr

, B, C

: r1

2 (A

t), 3

(At,

M, T

)Le

onto

don

hisp

idus

L. s

ubsp

. hisp

idus

In

ther

mop

hilo

us fo

rests

and

shru

blan

ds2:

s, 3

: rB:

s, C

: vr

2 (F

), 3

(F)

Page 9

351Phytol. Balcan. 14(3) • Sofia • 2008 Ta

ble 8

. C

ontin

uatio

n.ta

xaha

bita

tco

nsta

ncy

in su

bare

asco

nsta

ncy

in or

ders

new

reco

rdre

cord

ed b

y ot

her a

utho

rsM

atric

aria

recu

tita

L. -

For c

orre

ct n

omec

latur

e of

the c

ham

omile

, see

Jeffr

ey (1

979:

350

)1,

2, 3:

rA:

r, B

: c, C

: vr

12

(At),

3 (A

t, M

)

Myc

elis m

ural

is (L

.) Du

mor

t. Ve

ry co

mm

on in

Aln

us an

d Fa

gus f

ores

ts1:

c, 2

: r, 3

: sA:

c, B

, C: r

, D: v

r1

2 (A

t), 3

(At,

M, T

)Pr

enan

thes

pur

pure

a L.

It wa

s fou

nd in

one

loca

lity,

insid

e Tili

a fo

rest

1: r

D: vr

1Rh

agad

iolu

s stel

latu

s (L.

) Gae

rtn.

1: s,

2, 3

: rA:

c, B

: vr,

C: vr

12

(F),

3 (F

)Si

lybum

mar

ianu

m (L

.) Ga

ertn

.Ra

re, in

ther

mop

hilo

us sh

rubl

ands

1,

2, 3:

rA:

s, B

: c1,

23

(F)

Tara

xacu

m of

ficin

ale a

ggr.

Scat

tere

d in

azon

al fo

rests

and

Paliu

rus

shru

blan

ds1,

2, 3:

rA,

B: s

, C: v

r1,

23

(M)

Teph

rose

ris in

tegrif

olia

subs

p. a

uche

ri (D

C.) N

ord.

1,

3: r

A: vr

31

(S)

Trag

opog

on d

ubiu

s Sco

p.Ra

re, in

ther

mop

hilo

us fo

rests

and

shru

blan

ds1:

r, 2

: s, 3

: rB,

C: v

r1,

23

(M)

Conv

olvu

lace

aeCa

lyste

gia

sepi

um (L

.) R.

Br.

1: r

A, B

, C, D

: vr

1Co

nvol

vulu

s can

tabr

ica L

.Ra

re, in

ther

mop

hilo

us fo

rests

and

shru

blan

ds2,

3: r

A, C

: vr

2 (A

t), 3

(At,

M)

Corn

acea

eCo

rnus

mas

L.

Very

com

mon

in sh

rubl

ands

and

fore

sts1:

s, 2

, 3: c

A, B

: s, C

, D: c

12

(At,

G, T

), 3

(At,

M, T

)C.

sang

uine

a (C

.A. M

ey.)

Jáv.

subs

p. au

stral

isM

ostly

in P

lata

nus s

tand

s 1,

2, 3:

rA:

vc, C

: vr

12

(At),

3 (A

t, M

)Cr

assu

lace

aeSe

dum

cepa

ea L

. On

rock

s, in

side f

ores

ts an

d sh

rubl

ands

1, 2,

3: r

A: vr

, C: r

, D: s

12

(At,

G), 3

(At,

M, T

)S.

hisp

anicu

m L

. On

rock

s, in

side f

ores

ts an

d sh

rubl

ands

1, 2,

3: r

C: vr

, D: s

1, 2

3 (F

)Um

bilic

us lu

teus

(Hud

s.) W

ebb

& B

erth

el.M

ostly

in T

ilia

stand

s1,

2: r

D: vr

1, 2

U. ru

pestr

is (S

alisb

.) D

andy

On ro

cks,

insid

e for

ests

and

shru

blan

ds1,

2, 3:

rA,

B, C

, D: v

r1,

23

(M, F

)Cr

ucife

rae

Allia

ria p

etio

lata

(M. B

ieb.

) Cav

ara &

Gra

nde

Mos

tly in

azon

al fo

rests

1, 3:

rA:

s, B

, C, D

: vr

13

(At,

M, T

)Al

yssu

m m

ural

e Wal

dst.

& K

it.

1, 2,

3: r

A, B

: vr,

C: r,

D: v

r2

1 (S

), 3

(M)

Alys

sum

turk

esta

nicu

m R

egel

& S

chm

alh.

Rare

, in th

erm

ophi

lous

fore

sts an

d sh

rubl

ands

1, 2,

3: r

A: vr

, B: r

, C, D

: vr

1, 2

3 (F

)Ar

abid

opsis

thal

iana

(L.)

Hey

nh.

1, 2,

3: r

A, B

, C: r

, D: v

r1

2 (A

t), 3

(At,

M)

Arab

is la

xa Sm

. M

ostly

in fo

rests

1, 3:

rA:

r, C

: vr,

D: r

13

(F)

A. sa

gitta

ta (B

erto

l.) D

C.

Mos

tly in

fore

sts1,

3: r

A, C

, D: v

r1

3 (A

t, M

, T)

A. tu

rrita

L.

Mos

tly in

fore

sts, e

spec

ially

of h

igh

altit

udes

1, 2:

r, 3

: sA,

C: r

, D: c

1, 2

3 (M

)Be

rtero

a or

bicu

lata

DC.

1, 2:

rA,

C: v

r1,

2Ca

lepin

a irr

egul

aris

(Ass

o) T

hell.

Scat

tere

d in

azon

al fo

rests

and

Paliu

rus

shru

blan

ds1,

2, 3:

rA,

B: s

1, 2

3 (M

)

Caps

ella

burs

a-pa

storis

(L.)

Med

ik.

Rare

, in th

erm

ophi

lous

fore

sts an

d sh

rubl

ands

1: s,

2: r

, 3: s

A: c,

B: v

c, C:

vr1

2 (F

), 3

(F)

Card

amin

e acr

is Gr

iseb.

In

wet

pla

ces

1, 3:

rΑ:

vr1,

3C.

bul

bifer

a (L

.) Cr

antz

Mos

tly in

Fag

us fo

rests

1:

r, 3

: sA:

r, C

: vr,

D: c

13

(At,

M, T

)C.

grae

ca L

.M

ostly

in fo

rests

1, 2:

c, 3

: sA:

vc, B

: r, C

, D: c

12

(At,

T), 3

(At,

M, T

)C.

hirs

uta

L.

Mos

tly in

fore

sts an

d sh

rubl

ands

of l

ow al

titud

es1:

s, 2

: c, 3

: sA,

B, C

: s, D

: vr

12

(At),

3 (A

t, M

, T)

Page 10

352 Fotiadis, G. & Athanasiadis, N. • The vascular flora of Krousia and Belles Mts (NE Greece)Ta

ble 8

. C

ontin

uatio

n.ta

xaha

bita

tco

nsta

ncy

in su

bare

asco

nsta

ncy

in or

ders

new

reco

rdre

cord

ed b

y ot

her a

utho

rsC.

impa

tiens

subs

p. p

ectin

ata

(Pal

l. ex

DC.

) Tra

utv.

1, 2:

rA,

C: v

r1,

2Dr

aba

mur

alis

L.Ve

ry co

mm

on sp

ecie

s of P

aliu

rus s

hrub

land

s1:

r, 2

: s, 3

: cA:

vr, B

: vc,

C: r

1, 2

3 (M

)Er

ysim

um cu

spid

atum

(M. B

ieb.

) DC.

Ra

re, in

ther

mop

hilo

us fo

rests

and

shru

blan

ds1,

3: r

A, C

, D: v

r1

(S),

3 (M

)Er

ysim

um cr

assis

tylu

m C

. Pre

sl1,

2: r

A: vr

, B: r

, C, D

: vr

1, 2

Luna

ria a

nnua

subs

p. p

achy

rhiz

a (B

orbá

s) H

ayek

M

ostly

in az

onal

and

Fagu

s for

ests

1: s,

2: r

A: s,

C: v

r, D:

s1,

2Na

sturti

um of

ficin

ale R

. Br.

A ra

re sp

ecie

s of P

aliu

rus s

hrub

land

s2:

rB:

vr2

(F)

Rorr

ipa

thra

cica

(Gris

eb.)

Frits

chM

ostly

in th

erm

ophi

lous

fore

sts an

d sh

rubl

ands

1: r,

2: s

, 3: r

A: vr

, B: s

, C: r

12

(At,

T), 3

(At,

M, T

)Si

sym

briu

m of

ficin

ale (

L.) S

cop.

Mos

tly in

ther

mop

hilo

us fo

rests

and

shru

blan

ds,

espe

cially

in P

aliu

rus s

hrub

land

s1:

s, 2

, 3: r

A: s,

B: v

c, C:

vr1

2 (F

), 3

(F)

Tees

dalia

coro

nopi

folia

(Ber

gere

t) Th

ell.

1, 2,

3: r

B: r,

C: v

r1,

23

(M)

Thla

spi p

erfo

liatu

m L

. sub

sp. p

erfo

liatu

mRa

re, in

ther

mop

hilo

us fo

rests

and

shru

blan

ds1,

2, 3:

rA,

B, C

: vr

1, 2,

3Di

psac

acea

eKn

autia

am

bigu

a (F

riv.)

Boiss

. & O

rph.

1, 2,

3: r

C, D

: r1

2 (S

&K)

, 3 (T

)K.

arv

ensis

(L.)

Coul

t. Ve

ry co

mm

on in

Pal

iuru

s shr

ubla

nds

1, 2,

3: r

A: r,

B: v

c, C:

r1

2 (A

t), 3

(At,

M, T

)K.

integ

rifol

ia (L

.) Be

rtol. s

ubsp

. integ

rifol

ia

1: r

C: vr

1K.

mac

edon

ica G

riseb

. 1,

2: r,

3: s

A: vr

, B: s

, C: r

12

(At),

3 (A

t, M

, T)

Erica

ceae

Vacc

iniu

m m

yrtil

lus L

.Ra

re, in

Fag

us fo

rests

1: r

1 (S

)Eu

phor

biac

eae

Euph

orbi

a am

ygda

loid

es L

. sub

sp. a

myg

dalo

ides

Com

mon

in fo

rests

, loca

lly ve

ry co

mm

on1:

c, 2

: s, 3

: cA:

vc, C

: c, D

: vc

12

(At,

T), 3

(At,

M, T

)E.

bar

relie

ri Sa

vi1,

2: r

C: vr

12

(At)

E. cy

paris

sias L

.3:

rC:

vr3

(At)

E. ep

ithym

oide

s L.

1, 2,

3: r

A: r,

C: v

r1,

23

(F)

E. h

elios

copi

a L.

Mos

tly in

Pal

iuru

s shr

ubla

nds

1, 2,

3: r

A: s,

B: c

, C: v

r1,

23

(M)

E. li

ngul

ata

Heu

ff.2,

3: r

C: vr

23

(F)

E. p

laty

phyl

los L

. A

rare

spec

ies o

f azo

nal f

ores

ts1,

2: r

A: r

12

(F)

E. ta

urin

ensis

All.

1, 2,

3: r

A: vr

, B, C

: r1

2 (A

t), 3

(At,

T)Fa

gace

aeCa

stane

a sa

tiva

Mill

.Ta

xon

does

not

form

pur

e sta

nds e

xcep

t in

east

part

of G

reek

Bell

es (s

mal

l thi

cket

s in

Fagu

s an

d Qu

ercu

s for

ests)

. In

Krou

sia fo

und

only

as

juve

nile

1, 2,

3: r

A: s,

C, D

: vr

1, 2

3 (M

)

Fagu

s sylv

atica

L. s

ubsp

. sylv

atica

Taxo

n fo

rmed

exte

nsiv

e for

ests,

mai

nly i

n Be

lles

1: c,

2: r

, 3: c

A: vr

, C: r

, D: v

c1

2 (A

t), 3

(At,

T)Qu

ercu

s cer

ris L

. 1,

2: r

C, D

: vr

12

(At,

G)Q.

cocc

ifera

L.

Taxo

n fo

rmed

exte

nsiv

e shr

ubla

nds i

n so

uthe

ast

part

of B

elles

and

in ea

st - s

outh

east

part

of

Mav

rovo

uni

1: s,

3: c

A, B

: r, C

: c1

3 (A

t, M

)

Q. f

rain

etto

Ten

.Ta

xon

form

ed ex

tens

ive f

ores

ts, m

ainl

y in

Krou

sia1:

r, 2

, 3: c

A: vr

, B: r

, C: c

, D: v

r1

2 (A

t, G,

T),

3 (A

t, M

, T)

Page 11

353Phytol. Balcan. 14(3) • Sofia • 2008 Ta

ble 8

. C

ontin

uatio

n.ta

xaha

bita

tco

nsta

ncy

in su

bare

asco

nsta

ncy

in or

ders

new

reco

rdre

cord

ed b

y ot

her a

utho

rsQ.

pet

raea

subs

p. m

edwe

diew

ii (A

. Cam

us)

Men

itsky

Taxo

n fo

rmed

exte

nsiv

e for

ests

in K

rous

ia an

d Be

lles

1: c,

2: s

, 3: c

A: r,

C: v

c, D:

vc1

2 (A

t, T)

, 3 (A

t, M

, T)

Q. p

ubes

cens

Will

d.Ta

xon

form

ed ex

tens

ive f

ores

ts, m

ainl

y in

Krou

sia1:

r, 2

, 3: c

A: s,

B, C

: c1

2 (A

t, G)

, 3 (A

t, M

, T)

Q. ro

bur s

ubsp

. ped

uncu

liflo

ra (K

. Koc

h) M

enits

ky1,

2, 3:

rB,

C: v

r1

2 (A

t), 3

(At)

Gent

iana

ceae

Cent

auriu

m er

ythr

aea

Rafn

subs

p. er

ythr

aea

1, 2:

rB:

r, C

: vr

12

(At)

Gera

niac

eae

Gera

nium

colu

mbi

num

L.

Mos

tly in

ther

mop

hilo

us fo

rests

and

shru

blan

ds1,

2, 3:

rA:

r, B

: s, C

: r, D

: vr

12

(At),

3 (A

t, M

, T)

G. lu

cidum

L.

Very

com

mon

in fo

rests

and

shru

blan

ds o

f low

al

titud

es1,

2, 3:

cA,

B, C

: vc,

D: r

12

(At,

T), 3

(At,

M, T

)

G. m

acro

rrhi

zum

L.

In T

ilia

stand

s1,

3: r

D: vr

31

(S)

G. m

olle

subs

p. b

rutiu

m (G

asp.

) Gra

ebn.

3: r

C: vr

3 (F

)G.

mol

le L.

subs

p. m

olle

1, 2,

3: r

A: s,

B: v

c, C:

r1

2 (A

t), 3

(At,

F)G.

pus

illum

Bur

m. f.

1, 2:

rA:

s, B

: s, C

: vr

12

(F)

G. ro

berti

anum

L. s

ubsp

. rob

ertia

num

Mos

tly in

wet

and

dum

p pl

aces

1: s,

2, 3

: rA:

c, B

: vr,

C, D

: s1

2 (F

), 3

(F)

G. ro

tund

ifoliu

m L

.M

ostly

in fo

rests

and

shru

blan

ds o

f low

altit

udes

1: s,

2: r

, 3: s

A, B

: c, C

: r, D

: vr

12

(At),

3 (A

t, M

)G.

sang

uine

um L

.3:

rC:

vr3

(At,

M)

Gutti

ferae

Hype

ricum

mon

tbre

tii Sp

ach

Rare

, in in

to th

erm

ophi

lus Q

uerc

us fo

rests

1: s,

2: c

, 3: s

A: vr

, C: c

, D: r

12

(At),

3 (A

t, T)

H. o

lym

picu

m L

.Ra

re, in

ther

mop

hilo

us fo

rests

and

shru

blan

ds2,

3: r

C: vr

2 (A

t), 3

(At,

M)

H. p

erfo

ratu

m L

.Co

mm

on in

ther

mop

hilu

s Que

rcus

fore

sts1:

r, 2

, 3: c

A, B

: s, C

: c, D

: r1

2 (A

t, T)

, 3 (A

t, M

, T)

Jugla

ndac

eae

Jugla

ns re

gia

L.Na

tura

lized

in B

elles

and

Krou

sia m

ount

ains

. Its

occu

rren

ce is

relat

ed w

ith h

uman

activ

ities

1: r,

2: s

A: c,

C, D

: vr

1, 2

Labi

atae

Ajug

a ge

neve

nsis

L.

1, 2,

3: r

A, C

, D: v

r1

2 (A

t), 3

(At,

M, T

)A.

rept

ans L

. In

good

qua

lity s

tand

s 1,

2, 3:

rA:

r, B

, C: v

r, D:

r1

2 (A

t), 3

(At,

M)

Glec

hom

a hi

rsut

a W

alds

t. &

Kit.

1,

2, 3:

rA,

C, D

: vr

12

(At,

T), 3

(At,

M, T

)La

miu

m a

mpl

exica

ule L

. sub

sp. a

mpl

exica

ule

Rare

, in th

erm

ophi

lous

fore

sts an

d sh

rubl

ands

2: r

C: vr

2 (A

t, M

)L.

mac

ulat

um L

. Co

mm

on in

fore

sts an

d sh

rubl

ands

1, 2,

3: c

A: vc

, B: c

, C: s

, D: c

12

(At),

3 (A

t, M

, T)

Lyco

pus e

urop

aeus

L.

In w

et p

lace

s 1,

2, 3:

rA:

s1,

32

(F)

Meli

ttis m

elisso

phyl

lum

subs

p. a

lbid

a (G

uss.)

P.W

.Bal

l Lo

cally

com

mon

in fo

rests

1, 2:

r, 3

: sA:

s, C

: r, D

: vr

12

(At),

3 (A

t, M

, T)

Phlo

mis

sam

ia L

. In

Que

rcus

frai

netto

fore

sts

3: r

C: vr

3 (F

)Pr

unell

a la

cinia

ta (L

.) L.

1,

2, 3:

rA:

vr, B

: r, C

, D: v

r1

2 (A

t), 3

(At,

M)

P. vu

lgaris

L.

1: r,

2: s

, 3: r

A: c,

C, D

: vr

12

(At),

3 (A

t)Sa

lvia

glut

inos

a L.

In

good

qua

lity s

tand

s 1:

rA,

D: v

r1

Satu

reja

vul

garis

(L.)

Frits

ch

Very

com

mon

spec

ies o

f Que

rcus

fore

sts1,

2, 3:

cA,

B: s

, C: v

c, D:

c1

2 (A

t, G)

, 3 (A

t, M

, T)

Page 12

354 Fotiadis, G. & Athanasiadis, N. • The vascular flora of Krousia and Belles Mts (NE Greece)Ta

ble 8

. C

ontin

uatio

n.ta

xaha

bita

tco

nsta

ncy

in su

bare

asco

nsta

ncy

in or

ders

new

reco

rdre

cord

ed b

y ot

her a

utho

rsSc

utell

aria

colu

mna

e All.

subs

p. co

lum

nae

Mos

tly in

fore

sts1,

2: s,

3: c

A: s,

C: s

, D: c

12

(At,

T), 3

(At,

M, T

)St

achy

s alo

pecu

rus (

L.) B

enth

.1:

rC:

vr1

S. ge

rman

ica L

. sub

sp. g

erm

anica

Ra

re, in

ther

mop

hilo

us fo

rests

and

shru

blan

ds

1, 2,

3: r

B, C

: vr

12

(At),

3 (A

t, M

)S.

palu

stris

L.

3: r

C: vr

3 (A

t, T)

S. pl

umos

a Gr

iseb.

Ra

re, in

good

qua

lity s

tand

s1,

2, 3:

rC:

vr1

(S),

2 (A

t), 3

(At,

M, T

)S.

recta

subs

p. su

bcre

nata

(Vis.

) Briq

. 1

: rC:

vr1

S. sy

lvatic

a L.

Ra

re, in

good

qua

lity s

tand

s1,

2: r

A: s,

C: v

r1

2 (A

t)Te

ucriu

m ca

pita

tum

L.

Rare

, in th

erm

ophi

lous

fore

sts an

d sh

rubl

ands

1,

2, 3:

rB:

r, C

: vr

1, 2

3 (M

)T.

cham

aedr

ys L

. sub

sp. c

ham

aedr

ys

Com

mon

in fo

rests

and

shru

blan

ds1:

s, 2

, 3: c

A: r,

B: c

, C: v

c, D:

vr1

(S),

2 (A

t, G,

F),

3 (A

t, M

, T, F

)Th

ymus

long

icaul

is C.

Pre

sl Ra

re, in

ther

mop

hilo

us fo

rests

and

shru

blan

ds

1, 3:

rB,

C: v

.r.1

3 (F

)T.

sibt

horp

ii Be

nth.

Co

mm

on in

fore

sts an

d sh

rubl

ands

1, 2,

3: c

A: s,

B: c

, C: v

c, D:

vr1

(S),

2 (A

t), 3

(At,

M, T

)Le

gum

inos

aeAs

traga

lus g

lycyp

hyllo

s L.

Mos

tly in

fore

sts1:

s, 2

, 3: r

A: s,

C, D

: r1

2 (A

t), 3

(At,

M, T

)A.

glyc

yphy

lloid

es D

C.

One l

ocal

ity in

Pla

tanu

s sta

nd

2: r

A: vr

2 (F

)A.

onob

rych

is L.

Rare

, in th

erm

ophi

lus f

ores

ts an

d sh

rubl

ands

1, 3:

rA,

B, C

: vr

13

(At)

Cham

aecy

tisus

supi

nus (

L.) L

ink

Mos

tly in

fore

sts1,

2, 3:

rA,

C: v

r, D:

r1

2 (A

t), 3

(At,

T)C.

trifl

orus

(Lam

.) Sk

alick

á M

ostly

in fo

rests

1, 2,

3: r

C: r,

D: v

r1

2 (A

t), 3

(At,

T)Co

lute

a ar

bore

scen

s L.

3: r

C: vr

3 (A

t, F)

Dory

cniu

m h

erba

ceum

Vill

. sub

sp. h

erba

ceum

M

ostly

in Q

uerc

us p

ubes

cens

fore

sts1:

r, 2

: s, 3

: rC:

r1

2 (A

t, S)

, 3 (A

t, M

, T, S

)Ge

nista

carin

alis

Grise

b.

Mos

tly in

Que

rcus

and

Fagu

s for

ests

1, 2:

r, 3

: sC:

r, D

: vr

1 (S

), 2

(At),

3 (A

t, M

, T)

G. ti

ncto

ria L

. 1:

rC,

D: v

r1

Hipp

ocre

pis e

mer

us su

bsp.

emer

oide

s (Bo

iss. &

Sp

rune

r) L

asse

n M

ostly

in Q

uerc

us sh

rubl

ands

1,

2, 3:

rA,

C: r

12

(At),

3 (A

t, M

)

Lath

yrus

aph

aca

L.

1, 2,

3: r

B: vr

, C: r

12

(At),

3 (A

t, T)

L. la

xiflo

rus (

Desf.

) Kun

tze

Mos

tly in

Que

rcus

and

Fagu

s for

ests

1, 2,

3: c

C: c,

D: v

c1

2 (A

t, S)

, 3 (A

t, M

, T, S

)L.

nig

er (L

.) Be

rnh.

subs

p. ni

ger

Mos

tly in

Que

rcus

and

Fagu

s for

ests

1, 2:

s, 3

: cC:

r, D

: vr

12

(At,

G), 3

(At,

M, T

)L.

niss

olia

L.

1: r,

2: s

, 3: r

B, C

: r1

2 (A

t), 3

(At,

T)L.

pra

tens

is L.

In

Cas

tane

a an

d Fa

gus f

ores

ts1:

rC,

D: v

r1

(S)

L. sp

haer

icus R

etz.

1: r,

2: c

, 3: s

A: vr

, B, C

: s1

2 (A

t), 3

(At,

M, T

)L.

vene

tus (

Mill

.) W

ohlf.

M

ostly

in Q

uerc

us p

etra

ea su

bsp.

med

wedi

ewii

and

Tilia

fore

sts

1: c,

2: r

, 3: s

C: s,

D: c

12

(At,

T), 3

(At,

T)

Lens

nig

rican

s (M

. Bie

b.) G

ordo

n 1,

2, 3:

rC,

D: v

r1

2 (A

t), 3

(At)

Lotu

s cor

nicu

latu

s L.

Rare

, in th

erm

ophi

lus f

ores

ts an

d sh

rubl

ands

1, 2,

3: r

B: r,

C: v

r1

2 (A

t), 3

(At,

M, T

)Lu

pinu

s ang

ustif

oliu

s L.

Rare

, in th

erm

ophi

lus f

ores

ts an

d sh

rubl

ands

1, 2:

rC:

vr1

2 (A

t)M

edica

go a

rabi

ca (L

.) Hu

ds.

Very

com

mon

in P

aliu

rus s

hrub

land

s1,

2, 3:

rA:

s, B

: vc,

C: vr

1, 2

3 (F

)M

. lupu

lina

L.

1, 2:

r, 3

: sA:

vr, B

: r, C

: vr

12

(At),

3 (A

t, T)

M. m

inim

a (L

.) Ba

rtal

. 1:

r, 2

: s, 3

: rA:

r, C

: s1,

23

(M)

Page 13

355Phytol. Balcan. 14(3) • Sofia • 2008 Ta

ble 8

. C

ontin

uatio

n.ta

xaha

bita

tco

nsta

ncy

in su

bare

asco

nsta

ncy

in or

ders

new

reco

rdre

cord

ed b

y ot

her a

utho

rsM

. mon

speli

aca

(L.)

Trau

tv.

Rare

, in th

erm

ophi

lus f

ores

ts an

d sh

rubl

ands

1, 2:

rA:

vr, B

: r, C

: vr

1, 2

M. r

igid

ula

(L.)

All.

1, 2,

3: r

B, C

: vr

12

(At),

3 (A

t, M

)M

elilo

tus o

fficin

alis

(L.)

Pall.

1, 2:

rA:

r, C

: vr

12

(F)

Orni

thop

us co

mpr

essu

s L.

Mos

tly in

ther

mop

hilu

s shr

ubla

nds a

nd fo

rests

1: r,

2, 3

: sA:

vr, B

, C: s

, D: v

r1

2 (A

t), 3

(At,

M, T

)Ro

bini

a ps

eudo

acac

ia L

. Do

ubtfu

lly n

atur

alize

d. It

s occ

urre

nce i

s co

nnec

ted

with

hum

an im

pact

1,

2: r

A: vr

1, 2

Spar

tium

junc

eum

L.

Foun

d on

ly in

the S

E pa

rt o

f Bell

es, in

Que

rcus

sh

rubl

ands

1:

rC:

vr1

Trifo

lium

alp

estre

L.

Mos

tly in

Que

rcus

fore

sts

1: s,

2: r

, 3: s

A, B

: vr,

C: s,

D: r

1 (S

), 2

(At),

3 (A

t, M

, T)

T. a

ngus

tifol

ium

L.

Mos

tly in

ther

mop

hilu

s shr

ubla

nds a

nd fo

rests

1: r,

2, 3

: sB,

C: s

12

(At),

3 (A

t)T.

arv

ense

L.

Mos

tly in

ther

mop

hilu

s shr

ubla

nds a

nd fo

rests

1: r,

2, 3

: sA:

vr, C

: s, D

: r1

2 (A

t), 3

(At,

M)

T. ca

mpe

stre S

chre

b.

Scat

tere

d in

to fo

rests

and

shru

blan

ds1,

2: s,

3: r

B, C

, D: s

12

(At),

3 (A

t, M

, T)

T. gl

omer

atum

L.

1, 2,

3: r

B, C

: vr

1, 2

3 (F

)T.

hirt

um A

ll.

1: r,

2: s

, 3: r

B: r,

C: s

, D: v

r1

2 (A

t, T)

, 3 (A

t, M

, T)

T. m

ediu

m su

bsp.

bal

cani

cum

Vele

n.

Mos

tly in

Que

rcus

fore

sts

1, 2,

3: s

A: s,

B: r

, C, D

: s1

(H),

2 (A

t, H

), 3

(At,

T, H

)T.

ochr

oleu

con

Huds

. M

ostly

in Q

uerc

us fo

rests

1:

s, 2

, 3: r

A, B

: vr,

C: r,

D: c

12

(At),

3 (A

t, M

, T)

T. p

igna

ntii

Fauc

hé &

Cha

ub.

Mos

tly in

Que

rcus

fore

sts

1: r

C, D

: vr

1T.

repe

ns L

. 1,

2, 3:

rC,

D: v

r1

2 (A

t), 3

(At,

M, T

)T.

subt

erra

neum

L.

Mos

tly in

ther

mop

hilu

s shr

ubla

nds a

nd fo

rests

1: s,

2: c

, 3: s

A: vr

, B: c

, C: s

12

(At),

3 (A

t, M

, T)

Vicia

cassu

bica

L.

1, 2:

r, 3

: sA:

vr, B

, C: r

, D: s

12

(At),

3 (A

t, M

, T)

V. gr

andi

flora

Sco

p.

Com

mon

spec

ies o

f for

ests

and

shru

blan

ds1,

2, 3:

cA,

B, C

: c, D

: s1

2 (A

t, T)

, 3 (A

t, M

, T)

V. h

irsut

a (L

.) S.

F. Gr

ay

Com

mon

spec

ies o

f for

ests

and

shru

blan

ds1,

2, 3:

cA:

r, B

: c, C

: vc,

D: s

12

(At),

3 (A

t, M

, T)

V. la

thyr

oide

s L.

Mos

tly in

ther

mop

hilu

s shr

ubla

nds a

nd fo

rests

1: r,

2: c

, 3: s

A: vr

, B: c

, C: s

12

(At),

3 (A

t)V.

tenu

ifolia

subs

p. d

alm

atica

(A. K

ern.

) Gre

uter

In

Que

rcus

fore

sts3:

rC:

vr3

(At,

M, T

, S)

V. te

trasp

erm

a (L

.) Sc

hreb

. 1:

r, 2

: c, 3

: rC:

s, D

: r1

2 (A

t, T)

, 3 (A

t, M

, T)

V. v

illos

a su

bsp.

varia

(Hos

t) Co

rb.

Mos

tly in

ther

mop

hilu

s shr

ubla

nds a

nd fo

rests

1, 2,

3: r

A: vr

, B: s

, C, D

: vr

12

(At),

3 (A

t, M

, T)

V. v

illos

a Ro

th su

bsp.

villo

sa

Mos

tly in

ther

mop

hilu

s shr

ubla

nds a

nd fo

rests

1: s,

2: r

, 3: s

A: r,

B: c

, C: r

12

(At),

3 (A

t, T)

Lina

ceae

Linu

m th

racic

um D

egen

1:

rC:

vr1

Mor

acea

eFi

cus c

arica

L.

Doub

tfully

nat

ural

ized.

Its o

ccur

renc

e is

conn

ecte

d w

ith h

uman

impa

ct1,

2: r

A, B

, C: v

r1,

2

Mor

us a

lba

L.

Doub

tfully

nat

ural

ized.

Its o

ccur

renc

e is

conn

ecte

d w

ith h

uman

impa

ct

2: r

A: vr

2 (F

)

Olea

ceae

Frax

inus

ornu

s L.

Very

com

mon

spec

ies i

n fo

rests

1, 2,

3: c

A: vc

, B: s

, C: v

c, D:

vc1

2 (A

t, G,

T),

3 (A

t, M

, T)

Jasm

inum

frut

icans

L.

Mos

tly in

Que

rcus

shru

blan

ds

1, 2,

3: r

A, B

: vr,

C: r

12

(At),

3 (A

t)Li

gustr

um v

ulga

re L

. Ra

re, in

fore

sts an

d sh

rubl

ands

2, 3:

sA,

B, C

: r, D

: vr

2 (A

t), 3

(At)

7• Phytol. Balcan. 14(3) • 2008

Page 14

356 Fotiadis, G. & Athanasiadis, N. • The vascular flora of Krousia and Belles Mts (NE Greece)Ta

ble 8

. C

ontin

uatio

n.ta

xaha

bita

tco

nsta

ncy

in su

bare

asco

nsta

ncy

in or

ders

new

reco

rdre

cord

ed b

y ot

her a

utho

rsOl

ea eu

ropa

ea su

bsp.

olea

ster (

Hof

fm. &

Lin

k)

Nego

di

In o

ne lo

calit

y in

Quer

cus s

hrub

land

s3:

rB:

vr3

(F)

Phill

yrea

latif

olia

L.

Mos

tly in

Que

rcus

shru

blan

ds

1, 3:

rC:

r1

3 (F

)On

agra

ceae

Circ

aea

lute

tiana

L.

In w

et p

lace

s 1,

2: r

A, D

: vr

12

(F)

Epilo

bium

ang

ustif

oliu

m L

. M

ostly

in F

agus

fore

sts1,

2, 3:

rC,

D: v

r2

1 (S

), 3

(M)

E. la

nceo

latu

m S

ebas

t. &

Mau

ri M

ostly

in F

agus

fore

sts1,

3: r

C: vr

, D: c

13

(At,

T)E.

par

viflo

rum

Sch

reb.

M

ostly

in F

agus

fore

sts1:

rA,

C: v

r1

Oxal

idac

eae

Oxal

is ac

etos

ella

L.

In w

et an

d du

mp

plac

es1,

2: r

A: s,

B: v

r1

2 (F

)Pa

pava

race

aeCh

elido

nium

maj

us L

. 1,

2, 3:

rA:

r, C

: vr

1, 2

3 (M

)Co

ryda

lis so

lida

(L.)

Swar

t In

good

qua

lity s

tand

s 2,

3: r

C, D

: vr

23

(M)

Fum

aria

par

viflo

ra L

am.

Rare

, in th

erm

ophi

lous

fore

sts an

d sh

rubl

ands

1,

2, 3:

rA:

vr, B

: vc,

C: r

1, 2,

3Pa

pave

r dub

ium

L. s

ubsp

. dub

ium

Ra

re, in

shru

blan

ds

1, 2:

rB:

r, C

: vr

12

(F)

P. rh

oeas

L.

Rare

, in sh

rubl

ands

1,

2, 3:

rA:

s, B

: c, C

: vr

12

(At),

3 (A

t, M

)Ph

ytol

acac

eae

Phyt

olac

ca a

mer

icana

L.

Mos

tly in

Aln

us st

ands

1,

2: r

A: s

12

(F)

Plan

tagi

nace

aePl

anta

go b

ellar

dii A

ll.

Rare

, in P

aliu

rus s

tand

s 1,

2: r

B: r

1, 2

P. la

nceo

lata

L.

1, 2,

3: r

A: vr

, B: s

, C: v

r1

(S&

K), 2

(At),

3 (A

t, M

)P.

maj

or L

. sub

sp. m

ajor

In

Aln

us st

ands

1, 2,

3: r

A: s

12

(F),

3 (M

)Pl

atan

acea

ePl

atan

us or

ienta

lis L

. In

pur

e or m

ixed

with

Aln

us o

r Sal

ix st

ands

, alo

ng

strea

ms a

nd w

et al

luvi

al p

lain

s1:

s, 2

, 3: r

A: vc

, C: r

12

(At),

3 (A

t, M

)

Plum

bagi

nace

aeAr

mer

ia ru

meli

ca B

oiss

. 1,

2, 3:

rA,

C: v

r1

2 (A

t), 3

(At)

Poly

gona

ceae

Falo

pia

conv

olvu

lus (

L.) Á

. Löv

e 1:

s, 3

: rA:

s, B

: vr,

C: r,

D: s

13

(At,

M, T

)Ru

mex

ace

tose

lla L

. M

ostly

in th

erm

ophi

lus s

hrub

land

s and

fore

sts1:

r, 2

: s, 3

: rA,

C, D

: vr

1 (S

), 2

(At),

3 (A

t, M

, T)

R. co

nglo

mer

atus

Mur

ray

Mos

tly in

Aln

us st

ands

1,

2: r

A: r,

B: v

r1,

2R.

crisp

us L

. In

wet

and

dum

p pl

aces

2: r

A, B

: vr

2R.

obtu

sifol

ius L

. sub

sp. o

btus

ifoliu

s 1:

rA,

C: v

r1

R. p

ulch

er L

. sub

sp. p

ulch

er

Its o

ccur

renc

e is c

onne

cted

with

hum

an im

pact

1:

s, 2

, 3: r

A: vr

, B: c

, C: v

r1

2 (A

t), 3

(At,

F)R.

sang

uine

um L

. In

good

qua

lity s

tand

s 1,

2, 3:

rA:

r, D

: vr

1, 2

3 (F

)Pr

imul

acea

eAn

agal

lis a

rven

sis L

. Ra

re, in

shru

blan

ds an

d fo

rests

1, 2:

rA,

B, C

: vr

12

(At)

Page 15

357Phytol. Balcan. 14(3) • Sofia • 2008 Ta

ble 8

. C

ontin

uatio

n.ta

xaha

bita

tco

nsta

ncy

in su

bare

asco

nsta

ncy

in or

ders

new

reco

rdre

cord

ed b

y ot

her a

utho

rsCy

clam

en h

eder

ifoliu

m S

ol.

Mos

tly in

clos

e can

opy f

ores

ts an

d sh

rubl

ands

1: s,

2, 3

: cA:

vr, B

: r, C

: c, D

: s1

2 (A

t), 3

(At,

M, T

)Ly

simac

hia

num

mul

aria

L.

In w

et p

alce

s1,

3: r

A: r,

C, D

: vr

13

(At,

T)L.

pun

ctata

L.

1, 2,

3: r

A: r,

C, D

: vr

12

(At),

3 (A

t, M

, T)

Prim

ula

acau

lis (L

.) L.

M

ostly

in Q

uerc

us an

d Fa

gus f

ores

ts1:

r, 2

: s, 3

: cA:

vr, C

, D: s

12

(At,

T), 3

(At,

M, T

)P.

veris

L.

Mos

tly in

Que

rcus

and

Fagu

s for

ests

1: s,

2: r

, 3: r

A: vr

, C: r

, D: s

1, 2

3 (M

, F)

Pyro

lace

aeM

onot

ropa

hip

opity

s L.

In F

agus

and

rare

ly in

Tili

a fo

rests

1, 3:

rD:

vr1,

3Or

thili

a se

cund

a (L

.) H

ouse

In

Fag

us fo

rests

1: r

D: vr

1Ra

nunc

ulac

eae

Anem

one p

avon

ina

Lam

. Co

mm

on in

Pal

iuru

s and

Que

rcus

shru

blan

ds1:

r, 2

: c, 3

: sA:

r, B

: c, C

: r1,

23

(M)

Calth

a pa

lustr

is L.

In

Aln

us st

ands

1: r

A: vr

1Cl

emat

is fla

mm

ula

L.

Mos

tly in

to th

erm

ophi

lus f

ores

ts an

d sh

rubl

ands

1, 2:

s, 3

: rA:

r, B

: c, C

: s1

2 (A

t), 3

(At,

M, T

)C.

vita

lba

L.

Very

com

mon

in A

lnus

stan

ds, b

ut al

so fo

und

in

othe

r for

ests

and

shru

blan

ds1:

c, 2

: r, 3

: sA:

vc, B

, C, D

: s1

2 (A

t, G)

, 3 (A

t, M

, T)

Helle

boru

s odo

rus s

ubsp

. cyc

loph

yllu

s (A.

Bra

un)

Strid

M

ostly

in Q

uerc

us fo

rests

1:

r, 2

: s, 3

: cA:

vr, B

: r, C

: s, D

: r1

2 (A

t, G,

T),

3 (A

t, M

, T)

Nige

lla d

amas

cena

L.

Rare

, in sh

rubl

ands

and

fore

sts

1, 2,

3: r

A: vr

, B: r

, C: v

r1

2 (A

t), 3

(At,

M)

Ranu

ncul

us a

rven

sis L

. 1:

rA,

C: v

r1

R. ch

ius D

C.

1, 2:

rA:

vr, B

: r, C

: vr

1, 2

R. fi

caria

subs

p. fi

carii

form

is (F

.W. S

chul

tz) R

ouy

& F

ouca

ud1:

r, 2

: c, 3

: sA,

B: r

, C: s

12

(F),

3 (M

)

R. m

illefo

liatu

s Vah

l 2:

rC:

vr2

(F)

R. n

eapo

litan

us T

en.

The m

ost c

omm

on R

anun

culu

s spe

cies o

f for

ests

and

shru

blan

ds1:

r, 2

: c, 3

: sA:

vr, B

: r, C

: vr

1, 2

3 (M

)

R. p

silos

tach

ys G

riseb

. In

azon

al fo

rests

3: r

A: vr

3 (A

t, M

)R.

repe

ns L

. In

azon

al fo

rests

1, 3:

rA:

s1

3 (M

, F)

Thal

ictru

m a

quile

gifo

lium

L.

In fo

rests

of h

igh

altit

udes

2, 3:

rC:

vr, D

: r2

(At),

3 (A

t, T)

Rese

dace

aeRe

seda

lute

a L.

Ra

re, in

shru

blan

ds an

d fo

rests

1:

rC:

vr1

Rham

nace

aePa

liuru

s spi

na-c

hrist

i Mill

. Ta

xon

form

ed sh

rubl

ands

on

Belle

s and

Kro

usia

. Re

lated

to gr

azin

g1:

s, 2

, 3: c

A: s,

B: v

c, C:

s1

2 (A

t), 3

(At,

M)

Rosa

ceae

Agrim

onia

eupa

toria

L. s

ubsp

. eup

ator

ia

1, 2,

3: r

A: vr

, B: s

, C: v

r1

2 (A

t), 3

(At,

M, T

)Ap

hane

s flo

ribun

da (M

urb.

) Rot

hm.

Rare

in sh

rubl

ands

and

fore

sts

1, 2,

3: r

B: r,

C: v

r1,

32

(F)

Arem

onia

agr

imon

ioid

es (L

.) D

C. su

bsp.

agrim

onio

ides

M

ostly

in Q

uerc

us fo

rests

1,

2: s,

3: r

A: r,

C, D

: s1

2 (A

t, G,

T),

3 (A

t, M

, T)

Crat

aegu

s mon

ogyn

a Ja

cq. s

ubsp

. mon

ogyn

a On

e of t

he m

ost c

omm

on sp

ecie

s in

fore

sts an

d sh

rubl

ands

1, 2,

3: c

A , B

, C: v

c, D:

c1

2 (A

t, T)

, 3 (A

t, M

, T)

Page 16

358 Fotiadis, G. & Athanasiadis, N. • The vascular flora of Krousia and Belles Mts (NE Greece)Ta

ble 8

. C

ontin

uatio

n.ta

xaha

bita

tco

nsta

ncy

in su

bare

asco

nsta

ncy

in or

ders

new

reco

rdre

cord

ed b

y ot

her a

utho

rsCr

atae

gus m

onog

yna

subs

p. a

zare

lla (G

riseb

.) Fr

anco

In

Que

rcus

fore

sts2:

rC:

vrRe

cord

ed M

y Gam

isans

& H

ebra

rd (1

980)

but

its

occ

urre

nce n

eeds

conf

irmat

ion

Crat

aegu

s lac

inia

ta U

cria

In

Que

rcus

fore

sts2:

rC:

vrRe

cord

ed b

y Gam

isans

& H

ebra

rd (1

980)

but

its

occ

urre

nce n

eeds

conf

irmat

ion

Filip

endu

la v

ulga

ris M

oenc

h 1,

2, 3:

rA,

B: r

, C: v

r1

2 (A

t), 3

(At)

Frag

aria

vesc

a L.

M

ostly

in F

agus

fore

sts, b

ut al

so in

Pal

iuru

s sh

rubl

ands

1: r,

2, 3

: sA:

vr, B

: c, C

: r, D

: s1

2 (A

t), 3

(At)

Geum

urb

anum

L.

Mos

tly in

azon

al fo

rests

1: s,

2: c

, 3: s

A: vc

, B, C

, D: r

12

(At),

3 (A

t)M

alus

dom

estic

a Bo

rkh.

On

ly as

juve

nile.

Its o

ccur

renc

e is r

elate

d to

hu

man

impa

ct

1, 2,

3: r

A: r,

C, D

: vr

12

(At),

3 (A

t, T)

Pote

ntill

a m

icran

tha

DC.

On

e of t

he m

ost c

omm

on sp

ecie

s in

fore

sts1,

2, 3:

cA:

c, B

: r, C

: c, D

: vc

12

(At,

T), 3

(At,

M, T

)P.

recta

L.

Mos

tly in

ther

mop

hilu

s shr

ubla

nds a

nd fo

rests

1: r,

2: c

, 3: s

A: r,

B: c

, C: r

12

(At),

3 (A

t, M

, T)

Prun

us av

ium

L.

1, 2,

3: r

A, B

, C, D

: vr

12

(At),

3 (A

t, M

, T)

P. sp

inos

a L.

M

ostly

in th

erm

ophi

lus s

hrub

land

s and

fore

sts1,

2, 3:

rA,

B: s

, C: r

12

(At),

3 (A

t, M

, T)

Pyru

s am

ygda

lifor

mis

Vill.

Ve

ry co

mm

on sp

ecie

s in

Paliu

rus s

hrub

land

s1:

r, 2

: s, 3

: rA:

s, B

: vc,

C: r,

D: v

r1

2 (A

t), 3

(At,

M, T

)P.

com

mun

is L.

M

ostly

in th

erm

ophi

lus s

hrub

land

s and

fore

sts1:

r, 2

: s, 3

: rA:

s, B

: vr,

C: r

12

(At,

G), 3

(At,

M, T

)Ro

sa a

rven

sis H

udso

n - T

axon

omy o

f Ros

a sp

ecie

s ac

cord

ing t

o Zi

elińs

ki (1

990)

Co

mm

on in

fore

sts1:

s, 2

: c, 3

: sA:

c, B

: r, C

, D: c

12

(At,

T, Z)

, 3 (A

t, T,

Z)

R. ca

nina

L.

Scat

tere

d th

roug

h sh

rubl

ands

and

fore

sts1:

r, 2

, 3: s

A: s,

B: v

r, C,

D: s

1 (S

, Z),

2 (A

t, Z)

, 3: (

At, T

, Z)

R. ga

llica

L.

1, 2,

3: r

A, B

: r, C

: vr

1 (Z

), 2

(At,

Z), 3

(At,

M, T

, Z)

Rubu

s can

dica

ns R

chb.

M

ostly

in fo

rests

of h

igh

altit

udes

1: s,

2, 3

: rA,

B: v

r, C:

r, D

: s1

2 (A

t), 3

(At,

M, T

)R.

cane

scen

s DC.

M

ostly

in fo

rests

1, 2:

r, 3

: sA,

C, D

: r1

2 (A

t), 3

(At,

T)R.

hirt

us W

alds

t. &

Kit.

1:

s, 2

: r, 3

: sA:

c, B

: s, C

: r, D

: c1

2 (A

t), 3

(At,

M, T

)R.

idae

us L

. Fo

und

only

in F

agus

fore

sts1:

rD:

vr1

R. sa

nctu

s Sch

reb.

M

ostly

in fo

rests

and

shru

blan

ds o

f low

altit

udes

1: s,

2: c

, 3: s

A: vc

, B: s

, C: r

, D: v

r1

2 (A

t), 3

(At,

T)Sa

ngui

sorb

a m

inor

subs

p. m

urica

ta (S

pach

) Briq

. 1:

r, 2

: c, 3

: rA:

r, B

: s, C

: s, D

: vr

12

(At),

3 (A

t, M

, T)

Sorb

us d

omes

tica

L.

Mos

tly in

Que

rcus

fore

sts

1: r,

2, 3

: sC:

s, D

: vr

12

(At,

G, T

), 3

(At,

T)S.

torm

inal

is (L

.) Cr

antz

M

ostly

in Q

uerc

us fo

rests

1,

2: r,

3: s

A: vr

, C, D

: s1

2 (A

t, T)

, 3 (A

t, M

, T)

Rubi

acea

eCr

ucia

nella

ang

ustif

olia

L.

Rare

, in sh

rubl

ands

and

fore

sts

1, 3:

rC:

vr1

3 (A

t)Cr

ucia

ta la

evip

es O

piz.

Usua

lly in

azon

al fo

rests

and

Paliu

rus s

hrub

land

s1:

r, 2

, 3: s

A: s,

B: s

, C: r

12,

3: A

tC.

ped

emon

tana

(Bell

ardi

) Ehr

end.

M

ostly

in th

erm

ophi

lus s

hrub

land

s and

fore

sts1:

r, 2

, 3: c

A: r,

B, C

: c1

2 (A

t), 3

(At,

M)

Galiu

m a

parin

e L.

Very

com

mon

in sh

rubl

ands

and

fore

sts1,

2, 3:

cA:

c, B

: vc,

C: vc

, D: c

12

(At),

3 (A

t, M

)G.

div

arica

tum

Lam

. Ra

re, in

shru

blan

ds an

d fo

rests

1:

r, 2

: s, 3

: rC:

r1

2 (F

), 3

(F)

G. ex

alta

tum

Kre

ndl.

- Tax

onom

y acc

ordi

ng to

Kr

endl

(198

8)M

ostly

in Q

uerc

us, T

ilia

and

Fagu

s for

ests

1: r,

2, 3

: cA,

B: r

, C: s

, D: s

12

(At,

T), 3

(At,

T)

G. la

coni

cum

Boi

ss.

In Q

uerc

us fo

rests

2: r

C: vr

Reco

rded

by G

amisa

ns &

Heb

rard

(198

0) b

ut

its o

ccur

renc

e nee

ds co

nfirm

atio

n G.

odor

atum

(L.)

Scop

. In

good

qua

lity s

tand

s 1,

3: r

C: vr

, D: c

13

(M)

Page 17

359Phytol. Balcan. 14(3) • Sofia • 2008 Ta

ble 8

. C

ontin

uatio

n.ta

xaha

bita

tco

nsta

ncy

in su

bare

asco

nsta

ncy

in or

ders

new

reco

rdre

cord

ed b

y ot

her a

utho

rsG.

spur

ium

L.

Rare

, in sh

rubl

ands

and

fore

sts

3: r

C: r

3 (A

t, T)

Galiu

m ve

rum

L. s

ubsp

. ver

um

Mos

tly in

ther

mop

hilu

s shr

ubla

nds a

nd fo

rests

1: s,

2: c

, 3: s

A: r,

B: s

, C: c

12

(At),

3 (A

t, M

, T)

Sher

ardi

a ar

vens

is L.

M

ostly

in th

erm

ophi

lus s

hrub

land

s and

fore

sts1,

2, 3:

sA:

s, B

: vc,

C: r

12

(F),

3 (F

)Sa

licac

eae

Popu

lus t

rem

ula

L.

1, 3:

rD:

vr1

3 (A

t, M

, T)

Salix

alb

a L.

subs

p. a

lba

In p

ure o

r mix

ed w

ith S.

am

plex

icaul

is sta

nds,

alon

g stre

ams a

nd w

et al

luvi

al p

lain

s mai

nly i

n Kr

ousia

1, 2,

3: r

A: r

12

(At),

3 (A

t, M

)

S. am

plex

icaul

is Bo

ry &

Cha

ub.

In m

ixed

stan

ds w

ith S

alix

1,

2, 3:

rA:

r1

2 (A

t), 3

(At)

S. ca

prea

L.

1, 3:

rD:

vr1

3 (M

)Sa

xifra

gace

aeSa

xifra

ga ca

rpet

ana

subs

p. gr

aeca

(Boi

ss. &

H

eldr.)

D.A

. Web

b M

ostly

in th

erm

ophi

lus s

hrub

land

s and

fore

sts1:

r, 2

: s, 3

: rA:

vr, B

, C: r

12

(At),

3 (A

t, M

)

S. ro

tund

ifolia

L.

Mos

tly in

Tili

a sta

nds

1: r

D: r

1Sc

roph

ular

iace

aeDi

gita

lis gr

andi

flora

Mill

. 1,

2, 3:

rA:

vr, C

: r, D

: s1

2 (A

t, T)

, 3 (A

t, T)

D. la

eviga

ta su

bsp.

grae

ca (I

vani

na) W

erne

r In

Tili

a sta

nds

1: r

D: vr

D. la

nata

Ehr

h.

Mos

tly in

fore

st1,

2, 3:

sA,

B: r

, C, D

: s1

2 (A

t, G)

, 3 (A

t, M

, T)

D. v

iridi

flora

Lin

dl.

Mos

tly in

fore

sts o

f hig

h al

titud

es1,

2, 3:

rC:

vr, D

: r1

2 (A

t), 3

(At,

M)

L. ge

nisti

folia

(L.)

Mill

. Ve

ry ra

re in

fore

sts an

d sh

rubl

ands

1, 2:

rB,

C, D

: vr

1 (S

), 2

(At,

M, T

)L.

peli

sseria

na (L

.) M

ill.

Mos

tly in

ther

mop

hilu

s shr

ubla

nds a

nd fo

rests

1, 2,

3: r

B: s,

C: v

r1

2 (A

t), 3

(At)

Scro

phul

aria

cani

na L

. sub

sp. c

anin

a Ra

re, in

Que

rcus

shru

blan

ds1:

rC:

vr1

S. no

dosa

L.

In go

od q

ualit

y sta

nds

1, 2,

3: r

A: vr

, C: s

12

(At),

3 (A

t, T)

S. sc

opol

ii H

oppe

M

ostly

in F

agus

and

Abies

fore

sts1,

3: r

A, C

: vr,

D: s

1 (S

), 3

(At,

M, F

)Ve

rbas

cum

nig

rum

L.

In fo

rests

1, 3:

rA,

C: v

r, D:

s1

3 (M

, F)

V. p

hoen

iceum

subs

p. fl

avid

um (B

oiss

.) Bo

rnm

. M

ostly

in th

erm

ophi

lus f

ores

ts an

d sh

rubl

ands

1, 2,

3: c

A: r,

B: s

, C: c

, D: v

r1

2 (A

t), 3

(At,

M, T

)Ve

roni

ca a

rven

sis L

. Ra

re, in

shru

blan

ds an

d fo

rests

1,

3: r

A, B

, C: v

r1,

3V.

aus

triac

a L.

Ra

re, in

Que

rcus

shru

blan

ds1:

rC:

vr1

V. ch

amae

drys

L. s

ubsp

. cha

mae

drys

On

e of t

he m

ost c

omm

on sp

ecie

s of f

ores

ts an

d sh

rubl

ands

1, 2,

3: c

A: s,

B: c

, C, D

: vc

12

(At,

T), 3

(At,

M, T

) - T

he re

cord

of V

. vin

do-

bone

nsis

M.A

.Fisc

h. b

y Gam

isans

& H

ebra

rd

(198

0) is

pro

babl

y err

oneo

usV.

hed

erifo

lia L

. sub

sp. h

eder

ifolia

M

ostly

in th

erm

ophi

lus s

hrub

land

s and

fore

sts1,

2, 3:

rA:

vr, B

: r, C

: vr

12

(At),

3 (A

t, M

, T)

V. of

ficin

alis

L.

Mos

tly in

Tili

a sta

nds

1, 3:

rD:

s1

3 (A

t, T)

V. tr

iphy

llos L

.3:

rA:

vr3

(F)

V. u

rticif

olia

Jacq

. M

ostly

in m

ixed

Abi

es-F

agus

stan

ds1:

rC:

vr, D

: r1

V. ve

rna

L.

1, 2:

rC,

D: v

r1

2 (A

t)So

lana

ceae

Sola

num

dul

cam

ara

L.

3: r

C: vr

3 (A

t, M

, T)