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• Gas exchange, in concert with the circulatory system,provide the oxygen necessary for aerobic cellular respirationand removes the waste product, carbon dioxide.
Ventilation, which increases the flow of the respiratory medium over the respiratory surface, ensures that there is a strong diffusion gradient between the gill surface and the environment.
Crayfish and lobsters have paddlelike appendages that drive a current of water over their gills.
Fish gills are ventilated by a current of water that enters the mouth, passes through slits in the pharynx, flows over the gills, and exits the body.
• As blood moves anteriorly in a gill capillary, it becomesmore and more loaded with oxygen, but itsimultaneously encounters water with even higheroxygen concentrations because it is just beginning itspassage over the gills.
• All along the gillcapillary, there is adiffusion gradientfavoring the transferof oxygen fromwater to blood.
• Ventilation is much more complex in birds than inmammals.
• Besides lungs, birds have eight or nine air sacs that donot function directly in gas exchange, but act as bellowsthat keep air flowing through the lungs.