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Potential ATBI-sites in Poland: Bia owie a Primeval Forest and Bieszczady Mountains Jacek Szwedo Department of Systematics and Zoogeography Museum and Institute of Zoology Polish Academy of Sciences Wilcza 64 PL 00-679 Warszawa, Poland E-mail: [email protected] National Parks in Poland Procedures for conducting research in the national parks: Proposal of the project presented on the form to the Directorate of the national park group to be investigated state of knowledge and necessity of research methodology (non-invasive collection of data is preferable) proposed plots, etc. Proposal is presented to Scientific Advisory Board of the Park, discussed and voted After acceptance, the Director of the Park is presenting special permits Head of the project have to present annual reports Any paper resulting from the research must be offered to the Park authorities The voucher specimens must be deposited in the collection of the Park In some areas there are more strict rules 50 51 53 54 50 51 52 53 54 49 14 15 16 17 18 19 20 21 22 23 24 14 15 16 17 18 19 20 21 22 23 24 49 25 25 52 Bia owie a Primeval Forest 50 51 53 54 50 51 52 53 54 49 14 15 16 17 18 19 20 21 22 23 24 14 15 16 17 18 19 20 21 22 23 24 49 25 25 52 Bia owie a Primeval Forest Bia owie a National Park • Geographic position: NE Poland, W Belarussia; 23°31’- 24°21’E 52°29’ – 52°37’N • Area: about 1500 km 2 • Geobotanic division: Holarctic region; Euro-Siberian area; Mid European province; Mid-European subprovince; Masurian-Belarussian division; North Podlasie country, Bia owie a subcountry; Bia owie a circle • Physiographic division: 84 East Baltic-Belarussian Lowlands province; 843 Podlasie-Belarussia Lowland macroregion; 843.37 Bielska Plain mesoregion • Protection: Bia owie a National Park; Bialoviezhskij Zapoviednik; several Nature Reserves Bia owie a National Park: erected 1947 (1932); IUCN category: II Area (total): 105.02 km 2 strictly protected: 47.47 km 2 partly protected: 48.47 km 2 landscape protection: 9.08 km 2 FE – Member of EUROPARC Federation; MaB – Biosphere Reserve, UNESCO list; WH – World Heritage, UNESCO list;
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Potential ATBI-sites in Poland: Bia· owie�* a Primeval Forest and Bieszczady Mountains

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Page 1: Potential ATBI-sites in Poland: Bia· owie�* a Primeval Forest and Bieszczady Mountains

Potential ATBI-sites in Poland:

Bia owie a Primeval Forest

and Bieszczady Mountains

Jacek Szwedo

Department of Systematics and Zoogeography

Museum and Institute of Zoology

Polish Academy of Sciences

Wilcza 64

PL 00-679 Warszawa, Poland

E-mail: [email protected]

National Parks in Poland

Procedures for conducting research in the national parks:

Proposal of the project presented on the form to the Directorate of the

national park

• group to be investigated

• state of knowledge and necessity of research

• methodology (non-invasive collection of data is preferable)

• proposed plots, etc.

Proposal is presented to Scientific Advisory Board of the Park, discussed

and voted

After acceptance, the Director of the Park is presenting special permits

Head of the project have to present annual reports

Any paper resulting from the research must be offered to the Park

authorities

The voucher specimens must be deposited in the collection of the Park

In some areas there are more strict rules

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Bia owie a Primeval Forest

Bia owie a National Park

• Geographic position: NE Poland, W Belarussia; 23°31’- 24°21’E 52°29’ – 52°37’N

• Area: about 1500 km2

• Geobotanic division: Holarctic region; Euro-Siberian area; Mid European province;

Mid-European subprovince; Masurian-Belarussian division;

North Podlasie country, Bia owie a subcountry; Bia owie a circle

• Physiographic division: 84 East Baltic-Belarussian Lowlands province;

843 Podlasie-Belarussia Lowland macroregion;

843.37 Bielska Plain mesoregion

• Protection: Bia owie a National Park; Bialoviezhskij Zapoviednik;

several Nature Reserves

Bia owie a National Park: erected 1947 (1932); IUCN category: II

Area (total): 105.02 km2

strictly protected: 47.47 km2

partly protected: 48.47 km2

landscape protection: 9.08 km2

FE – Member of EUROPARC Federation; MaB – Biosphere Reserve, UNESCO list;

WH – World Heritage, UNESCO list;

Page 2: Potential ATBI-sites in Poland: Bia· owie�* a Primeval Forest and Bieszczady Mountains

Habitat: the last remnants of European Lowland primeval wood,

forest communities:

Vaccinio myrtilli-Pinetum, Vaccinio uliginosi Pinetum, Sphagnetum medii

pinetosum, Querco-Pinetum, Querco-Carpinetum, Circaeo-Alnetum, Fraxino-

Alnetum, Alnetum glutinosae, Saliceto-Franguletum.

Plants: 1070 species of vascular plants

Fungi: estimated diversity 3000-4000 species

Lichens: about 600 species

Animals: over 12000 species (of about 30000 species recorded in Poland)

Infrastructure:

Bia owie a Directorate of the Park, several scientific institutions

Bia owie a NP museum

Tourist base, shopping and restaurants

Hajnówka railway station, bus station, several institutions

and authorities managing Bia owie a Primeval Forest

Bielsk Podlaski railway station, bus station

Bia ystok railway station, bus station

Series „Bibliography of Bia owie a Primeval Forest” (1969, 1976, 1983, 1991, 1997)

Three journals are published in Bia owie a:

"Acta Theriologica" - an international zoological and ecological journal (MammalResearch Institute - Zak ad Badania Ssaków),

"Phytocoenosis" - a multilingual geobotanical bulletin (Bia owie a GeobotanicalStation of Warsaw University)

„Parki Narodowe i Rezerwaty Przyrody” [National Parks and Nature Reserves] -a quarterly with articles in Polish and English summaries (the Bia owie aNational Park).

Important Monographs

Krasi ska M., Krasi ski Z.A., 2004. Monografia przyrodnicza UBR. Studium Fotografii

Przyrodniczej HAJSTRA. Warszawa - Bia owie a: 1-312.

Pucek Z. ( ed .); Pucek Z., Belousova I.P., Krasinska M., Krasinski Z.A. and Olech W. (comps .),

2004. European bison. Status survey and conservation action plan. IUCN/SSC Bison Specialist

Group. IUCN, Gland, Switzerland and Cambridge, UK. ix +54pp.

Bernadzki E., Bolibok L., Brzeziecki B., Zaj czkowski J., ybura H., 1997. Zmiany sk adu

gatunkowego drzewostanów naturalnych w Bia owieskim Parku Narodowym (1936-1993). Parki

nar. Rez. przyr . 16(2): 3-25.

Borowski S., Oko ów Cz., 1988. Birds of the Bia owie a Forest. Acta zool. Kraków. 31,2: 65-114.

Cie li ski S., Tobolewski Z., 1988. Lichens ( Lichenes ) of the Bia owie a Forest and its western

Foreland. Phytocoenosis , Bia owie a, suppl. N.S. 1.

Fali ska K., 1973. Dynamika sezonowa runa zbiorowisk le nych Bia owieskiego Parku

Narodowego. Phytocoenosis 2(1): 3-120.

Fali ski J.B., (red.), 1968. Park Narodowy w Puszczy Bia owieskiej. Warszawa.

Fali ski J.B., 1986. Vegetation dynamics in temperate lowland primeval forest. Ecological

studies in Bia owie a Forest. W. Junk. Dordrecht.

Fali ski J.B., 1994. Concise geobotanical atlas of Bia owie a Forest. Phytocoenosis Vol. 6 (N.S.)

Supplemetum Cartographiae Geobotanicae 6: 3-34.

Important Monographs, cntd

Fali ski J.B. (red.), 1996. Cryptogamous plants in the forest communities of Bia owie a

National Park, Functional group analysis and general synthesis (Project Crypto 3). Polish

Botanical Society. Warszawa - Bia owie a.

Fali ski J.B., Mu enko, W. (red.) 1997. Cryptogamous plants in the forest communities of

Bia owie a National Park, Ecological atlas (Project Crypto 4). Phytocoenosis Vol. 9 (N.S.)

Supplemetum Cartographiae Geobotanicae 7.

Gutowski J. M., Jaroszewicz B. (red.) 2001. Katalog fauny Puszczy Bia owieskiej. Instytut

Badawczy Le nictwa. Warszawa.

J drzejewska B., J drzejewski W. 1998. Predation in vertebrate communities. The Bia owie a

Primeval Forest as a case study . Springer. Berlin - Tokyo.

Kwiatkowski W. 1994. Krajobrazy ro linne Puszczy Bia owieskiej (mapa w skali 1:50 000 z

tekstem obja niaj cym). Phytocoenosis Vol. 6 (N.S.) Supplemetum Cartographiae

Geobotanicae 6: 35-87.

Matuszkiewicz W., 1952. Zespo y le ne Bia owieskiego Parku Narodowego. Ann. Univ. M.-C.

Sk odowska . Lublin, suppl. 6.

Olszewski J.L. 1986. Rola ekosystemów le nych w modyfikacji klimatu lokalnego Puszczy

Bia owieskiej. Ossolineum, Warszawa.

Page 3: Potential ATBI-sites in Poland: Bia· owie�* a Primeval Forest and Bieszczady Mountains

Important Monographs, cntd

Paczoski J. 1930. Lasy Bia owie y. Pa stwowa Rada Ochrony Przyrody, Pozna .

Prusinkiewicz Z., Michalczuk Cz., 1998. Gleby Bia owieskiego Parku Narodowego (z map 1:20

000). Phytocoenosis Vol. 10 (N.S.) Supplemetum Cartographiae Geobotanicae 10.

Pugacewicz E., 1997. Ptaki l gowe Puszczy Bia owieskiej. Pó nocnopodlaskie Towarzystwo

Ochrony Ptaków, Bia owie a.

Soko owski A.W. 1993. Fitosocjologiczna charakterystyka zbiorowisk le nych Bia owieskiego

Parku Narodowego. Parki Nar. Rez. Przyr . 12(3): 5-188 (z map ).

Soko owski A.W. 1995. Flora ro lin naczyniowych Puszczy Bia owieskiej . Bia owieski Park

Narodowy, Bia owie a.

Tomia oj L., Weso owski T., Walankiewicz W. 1984. Breeding bird community of a primeval

temperate forest (Bia owie a National Park, Poland). Acta orn . 20: 241-310.

Wi cko E. 1984. Puszcza Bia owieska . PWN, Warszawa.

The most numerous vascular plant families in the Bia owie a Forest are:

• the aster family (Asteraceae) - 108 species,

• the grass family (Poaceae) - 85

• the sedge family (Cyperaceae) - 66

• the rose family (Rosaceae) - 59

• the Fabaceae family (Fabaceae) - 54

• the pink family (Caryophyllaceae) - 43

• the figwort family (Scrophulariaceae) - 41

• the mint family (Lamiaceae) - 41

• the Brassicaceae family (Brassicaceae) - 40

• the buttercup family (Ranunculaceae) - 35

• the Apiaceae family (Apiaceae) - 32

• the orchid family (Orchidaceae) - 24

• the buckwheat family (Polygonaceae) - 24

331Spiders

51Crustacea

10Tarchigrades

20Leeches

50Molluscs

111Earth worms

2Thorny-headed worms

324Roundworms

49Gastrotriches

97Rotifers

84Flatworms

86Protozoa

58Mammals

250Birds

7Reptiles

13Amphibians

32Fish

9284Insects

66Springtails

3Coneheads

31Myriapods

10Pseudoscorpionida

14Harvestmen

576Acarina

Number of species observed in the Bia owie a Forest

Belarussian part of the Bia owie a Primeval Forest is poorly investigated

An Invertebrate monitoring project has been conducted

in Bia owie a Primeval Forest since 1988 and continuing

First summary of results has been published in separate issue of journal:

Prace Instytutu Badawczego Le nictwa, 1995: seria A, nr 790-800

Recent summaries:

Gutowski J.M. 2004. [Invertebrates as a biological monitoring object in Bia owie a Primeval

Forest]. Lesne Prace Badawcze, 1: 23-54.

Gutowski J.M., Jaroszewicz B. 2004. [Bia owie a Primeval Forest as a refuge of European

entomofauna]. Wiadomo ci entomologiczne, 23, Supplement 2: 67-87.

Series of papers resulting from monitoring in various journals, concerning e.g.

Coleoptera (Sphaeritdae, Histeridae, Lymexylidae, Cerylidae, Colydiidae, Leiodidae,

Scydmaenidae, Staphylinidae, Clambidae, Scirtidae, Byrrhidae, Trogossitidae, Cleridae,

Sphindidae, Monotomidae, Phalacridae, Erotylidae, Corylophidae, Mycetophagidae, Ciidae,

Mordellidae, Tenebrionidae, Pythidae, Anthicidae, Aderidae, Scraptiidae, Kateretiidae,

Nitidulidae, Cryptophagidae, Latridiidae, Scolytidae, Chrysomelidae), Thysanoptera, Araneae

Which taxa need to be investigated?

Protists (in most cases only parasitic taxa were recorded)

Numerous groups of invertebrates, e.g.

Nematoda (parasitic and free-living)

Annelida

Crustacea

Chelicerata

Hexapoda: Protura and Collembola

: Pterygota – various orders and families

Bia owie a National Park

17-230 Bia owie a, Park Pa acowy 11

tel. 0-85 6829700

tel./fax 085 6812306; tel. 085 6812360, 085 6812275

e-mail: [email protected]

http://www.bpn.com.pl

Managing Director

mgr in . Józef Popiel

tel. 085 6812306 (110)

e-mail: [email protected]

Page 4: Potential ATBI-sites in Poland: Bia· owie�* a Primeval Forest and Bieszczady Mountains

Museum and Education Centretel. 085 6829730 tel. 085 6812275 fax 0-85 6812306 (113, 116) mgr El bieta Jab o ska - Main Natural Education specialisttel. 0-85 6822913; tel./fax 085 6812306, 085 6812360 (113) e-mail: [email protected]

Natural History Museumtel. 085 6829702, 085 6812275 fax 085 6812306 (116) e-mail: [email protected]

mgr in . Mateusz Szymuratel. 085 6829711; 085 6812275 (111) e-mail: [email protected]

Invertebrates monitoring data:

Prof. dr hab. Jerzy M. Gutowski

Zak ad Lasów Naturalnych

Instytut Badawczy Le nictwa

[Department of Natural Forests

Forest Research Institute]

Park Dyrekcyjny 6, 17-230 Bia owie a,

tel. 085 6812396, fax. 085 6812203,

e-mail: [email protected]

Other scientific institutions

Bia owie a Geobotanical Station of Warsaw University [Bia owieska Stacja

Geobotaniczna Uniwersytetu Warszawskiego] - dynamics of plant communities,

succession phenomena, population and demographic research, phenology, invasions of

external species; address: ul. Sportowa 19, 17-230 Bia owie a, tel. 085 6812548; e-mail:

[email protected]; Contact person: dr Bogdan Jaroszewicz

Mammal Research Institute of the Polish Academy of Sciences PAN [Zak ad Badania

Ssaków PAN] [ http://bison.zbs.bialowieza.pl/] - ecology of mammals (rodents, hoofed

mammals and predators), genetics; address: ul. Waszkiewicza 1, 17-230 Bia owie a, tel.

085 6812278. e-mail: [email protected]

Contact person: prof. dr hab. W odzimierz J drzejewski,

e-mail: [email protected]

Forest Research Institute Natural Forest Department [Zak ad Lasów Naturalnych

Instytutu Badawczego Le nictwa] - phytosociology, syntaxonomy, entomology, selection

of forest trees, air and water chemical monitoring: [ http://las.ibles.waw.pl/]; address:

ul. Park Dyrekcyjny 6, 17-230 Bia owie a, tel. 085 6812396. e-mail:

[email protected]

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Bieszczady National Park

Bieszczady Mountains: border of Poland, Slovakia and Ukraine

Page 5: Potential ATBI-sites in Poland: Bia· owie�* a Primeval Forest and Bieszczady Mountains

Bieszczady Mountains i.e. the area between the U ocka Pass in the east, the

upkowska Pass in the west, Polish state border with Slovakia and Ukraine in the

south, and the valley of San River in the north

• Geographic position: SE Poland, NE Slovakia, W Ukraine; 49°-49°50’ N, 22°-23°E

• Area: about 2000 km2

• Geology - typical for the majority of the Carpathian Range, with sandstone, and

shale being the dominating base rocks.

• Physiographic division: 52 Eastern Carpathians province; 522 Eastern Beskidy

macroregion; 522.1 Forest Beskidy; 522.12 Western Bieszczady mezoregion

• Geobotanic division: Holarctic region; Euro-Siberian area; Mid European Montane

province; Carpathian subprovince; Eastern Carpathians division; Forest Carpathians

circle, Bieszczady subcircle

Elevations vary between about 500 m a.s.l. in main valleys and 1346 m a.s.l. at

the highest peak - Tarnica.

Weather - strongly influenced by a continental climate of Ukraine and western

Russia. An average air temperature is +5.1°C, (annual range from -40 to +35),

and annual precipitation 1200 - 1300 mm. Snow cover lasts here from 110 to

160 days.

Bieszczady National Park: erected 1973; UNESCO Biosphere Reserve

Area (total): 292,02 km²

forests 247,24 km²

managed reas 21,12 km²

water 0,76 km²

Protection: strictly protected: 184,25 km²

partly protected: 107,77 km²

Since 1992 part of International Reserve

of Biosphere „Eastern Carpathians”

together with Polish Cisniansko-Wetlinski

Landscape Park and The Dolina Sanu

Landscape Park, Slovak Poloniny National

Park, Ukrainian Uzhansky National Park

and Dolina Sanu Landscape Park

Since 2005 member of PAN-Parks net

Forested in over 60% with beech-fir natural forest. Forest consists in 60 - 75%

of beech Fagus silvatica and fir Abies alba, supplemented with alder, spruce, sycamore,

pine, larch, aspen, and willows. Large part of former agricultural land belonging to

bankrupt state farms, remains abandoned under various stages of secondary succession,

in some cases up to 50 years old.

• More than 60 various non-forest plant associations and communities.

• The most widely occurring forest association in the Bieszczady National Park is the

Carpathian beech wood Dentario glandulosae-Fagetum. Another common type of beech

forest - acidophilous beech woods Luzulo nemorosae-Fagetum

Vertical vegetation zones

— submountain (colline) zone up to ca 500 m a.s.l.,

— lower montane zone up to ca 1150 m a.s.l.,

— “polonina” zone i.e. zone of subalpine and alpine communities, occurring above the

upper forest line up to the highest summits

Flora:

• Vascular plants: about 700 species

• Moss: 200 species

• Lichens: 300 species

Page 6: Potential ATBI-sites in Poland: Bia· owie�* a Primeval Forest and Bieszczady Mountains

Fauna:

284 vertebrate species includes all large European predators (wolf, brown bear,

lynx, wildcat), and almost all herbivorous species: European bison, red and roe

deer, moose, and wild boar

invertebrates (very patchy knowledge, number of species not known, estimated

over 7000 species

— 112 species occurring in Poland exclusively within the limits of the Polish part

of the International Reserve of Biosphere “Eastern Carpathians”,

— 115 species new to Poland, known from other countries,

— 48 East-Carpathian endemics,

— 14 East-South-Carpathian endemics,

— 3 West-Carpathians sub-endemics,

— 21 species new to science,

— 19 species new to science with loci typici within the area,

— 2 species known only as locus typicus in the Bieszczady Mountains.

The highest and least affected by an antropogenic influence part of Bieszczady

(over 27 thousand hectares) has been included in 1973 into the Bieszczadzki

National Park. In 1992 a Biosphere Reserve „Eastern Carpathians”, encompassing

the national park and two landscape parks, has been established over the area of

108 thousand hectares. With adjoining Slovak and Ukrainian protected areas it

forms an International Biosphere Reserve covering over 160 thousand hectares. The

density of human population ranges between 5 - 10 people per sq. km, and is

among the lowest in the country.

Long Term Ecological Research in the Bieszczady Mountains

Ecological projects have started in Bieszczady in the late fifties with a study on the

population of small mammals

Recent project: Application of long-term data in studies on populations of large

mammals and the evaluation of anthropogenic effects at Bieszczady Mountains,

Poland by dr hab. Kajetan Perzanowski (MIIZ)

Types and lenghts of data sets:

• history of land-use: back to 1850

• population data on rodents: since the mid-fifties

• invertebrate studies: since the sixties

• floristic studies: since the sixties

• forest structure and composition: since the mid-sixties

• meterology: since 1960

• population data on large mammals: since 1980

Series:Monografie Bieszczadzkie [Bieszczady Monographs]

13 volumes concerning

1 – Forest communities2 - Soils3 – Vascular plants4 – Poloniny plant communities5 – Non forest plant communities of the valleys6 – Species composition and structure of the tree assemblages7 – Invertebrates part 1 8 – Invertebtrates part 2 9 - Vertebrates10 – Ecological and biogeographical background

for protection purposes11 – Landscape protection12 – Data on history of settlemetns13 – Catalogue of cultural values

Roczniki Bieszczadzkie [Bieszczady Annals]

Numerous papers on soil, climate, flora, vegetation and fauna of thearea:

Adamczyk B., Zarzycki K. 1963. Gleby bieszczadzkich zbiorowisk le nych. Acta agr. et silv. 3: 133-175.Bobek B., Perzanowski K., Zielinski J., 1986: Red deer population census in mountains -testing of an alternative method. Acta theriol., 31: 423-431.Bobek B., Perzanowski K., mietana W., 1992: The influence of snow cover on the selection pattern within red deer population by wolves in Bieszczady Mountains. in: Trans. XVIIIthIUGB Congr. Vol. 2: 341-348.Bro ek S. 1993. Przekszta canie górskich gleb porolnych przez olsz szar (Alnus incana(L.) Moench). Rozpr. habil. nr 184, Zesz. Naukowe AR w Krakowie.Buchalczyk T., Markowski J., 1979. Ssaki Bieszczadow Zachodnich.Ochr. Przyr.42:119-150.Burakowski B. 1971. Spr yki (Elateridae) Bieszczadów. Fragm. Faun.17:185-272.Cicho M., Zaj c T. 1991. Avifauna of Bieszczady National Park (SE Poland) in 1987 and 1988 - quantitative and qualitative data. Acta Zool. Cracov. 34,2: 497-517.Feliksik E. 1995. Próba oceny zagro enia lasów beskidzkich przez emisje przemys owe w oparciu o analizy dendrologiczne. In: Mat. Konf. "Ekologiczne i ekonomiczne uwarunkowania rozwoju gospodarczego Karpat p d-wsch" Bieszczady 1995, CEEW Krosno: 447 ppFr ckowiak W. 1997. Diet and food habits of the brown bear (Ursus arctos L.) in Polish eastern Carpathians. J. Wildl. Res. 2,2: 154-160 G owaci ski Z. 1996. Protection and regulation of ungulate populations in the BieszczadyNational Park. Roczniki Bieszczadzkie 5: 117 - 132.

G owaci ski Z., Profus P. 1996. L gowe zespo y ptaków buczyny i olszynki nadpotokowej w Bieszczadzkim Parku Narodowym. Roczniki Bieszczadzkie 5: 109-116.G owaci ski Z., Profus P., Wuczy ski A. 1995. Zarys awifauny Bieszczadzkiego parku Narodowego i problemy jej ochrony. Roczn. Bieszczadzkie 4: 259-263.G owaci ski Z. and Witkowski Z. 1969. Fauna Bieszczadów Zachodnich. Ochr. Przyrody 34:127-160.Grodzinski W., 1956: Swiat ro lin i zwierz t w Bieszczadach Polskich. Wierchy, 25: 168-172.Grodzi ski W., 1957: Materia y do fauny kr gowcow Bieszczad Zachodnich. Zesz. Nauk. UJ Zool. 10, 1: 177-221.Grodzi ski W., Górecki A., Janas K., Migula P. 1966. Effect of rodents on the primary productivity of alpine meadows in Bieszczady Mountains. Acta theriol. 11, 20: 419-431.Hess M. 1965. Pi tra klimatyczne w polskich Karpatach Zachodnich. Prace geogr. nr 123, IGiPZPAN.J dryczkowski W. 1979. Krocionogi (Diplopoda) Bieszczadów. Fragm. Faun. 25: 78-93.J dryczkowski W. 1987. Zaleszczotki (Pseudoscorpiones) Bieszczadów. Fragm. Faun. 30: 342-349.Kanzaki N., Perzanowski K. 1997. The potential role of wolf predation in regulating wild boar population in Bieszczady, Poland. Wildl. Conserv. Japan 2,4: 205-212.Kasprzyk S.1972.Lasy w Bieszczadach. Stan aktualny-potrzeby, perspektywy. Las Polski 20: 19. Kondratyuk A., Coppins.B.A, Zelenko S., Khodosovcev A., Coppins A., Wolseley P. 1997. Lobarion lichens as indicators of primeval forests in the Ukrainian part of the International Biosphere Reserve "Eastern Carpathians": distribution, ecology, long-term monitoring and reccommendations for conservation. Roczniki Bieszczadzkie 6: 65-88.Kostecka J., Skocze S. 1993. Earthworm (Lumbricidae) populations in four types of beech wood Fagetum carpaticum in the Bieszczady National Park. Acta Zool. Cracov. 36: 1-13.

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Kowalski K., 1960: Pitymus Mc. Mutrie, 1831 (Microtidae, Rodentia) in the Northern Carpathians. Acta theriol., 4, 6: 81-91.Le niewicz K., Perzanowski K., 1989: The winter diet of wolves in Bieszczady Mountains. Actatheriol., 34: 373-380.Michalik S. 1993. Zbiorowiska le ne Bieszczadzkiego Parku Narodowego, ich waloryzacja i problemy ochrony. Roczniki Bieszczadzkie 2: 51-62Michalik S., Szary A. 1997. Zbiorowiska le ne Bieszczadzkiego Parku Narodowego. Monografie Bieszczadzkie 1: 175 pp.Michna E., Paczos S. 1972. Zarys klimatu Bieszczadów Zachodnich. Lubelskie Tow. Nauk. Ossolineum, Wroc aw - Warszawa - Kraków.Nowosad M. 1993. Dyskusja nad koncepcj bada klimatologicznych w Bieszczadach. Roczniki Bieszczadzkie 2: 165-172.Nowosad M. 1994. Zarys charakterystyki pokrywy nie nej w Bieszczadach. Annales UMCS, sec.B.49: 197-215.Nowosad M. 1995. Zarys klimatu Bieszczadzkiego Parku Narodowego. Roczniki Bieszczadzkie 4: 163-183.Perzanowski K., Pucek T., Podyma W., 1986: Browse supply and its utilisation by deer in Carpathian beechwood (Fagetum carpaticum). Acta theriol., 31: 107-118.Perzanowski K. 1993: The economic aspects of wolf predation in Bieszczady Mountains. in: Wolves in Europe - status and perspectives. C. Promberger, W. Schroder (eds). Proc. Workshop Wolves in Europe - current status and prospects. Oberammergau 1992. 126-129.Perzanowski K., Bobek B., Frackowiak W., Gula R., Kabza B., Merta D. 1996. The management of large mammals in the Eastern Carpathians Biosphere Reserve. in: Biodiversity Conservation in Conserv. in Transboundary Protected Areas. Bieszczady-Tatry 1994. A. Breymeyer and R. Noble (eds). National Academy Press: 178-185.

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Taxa to be investigated:

various groups of invertebrates – present state

of knowledge is very patchy, summarized

in the two volumes of the series:

Monografie Bieszczadzkie,

Vol. 7 and Vol. 8 edited by J. Paw owski

published in 2000

Published resources:

Paw owski J. (ed.), 2000. Bezkr gowce Bieszczadów Zachodnich ze szczególnym uwzgl dnieniem Bieszczadzkiego Parku Narodowego - cz I. Monografie Bieszczadzkie, 7: 1-260.Paw owski J. (ed.), 2000. Bezkr gowce Bieszczadów Zachodnich ze szczególnym uwzgl dnieniem Bieszczadzkiego Parku Narodowego - cz II. Monografie Bieszczadzkie, 8: 1-369.

Bieszczady National Park

Directorate:

38-714 Ustrzyki Górne

tel/fax 013 461 06 50, 013 461 06 10

e-mail: [email protected];

[email protected]

http://www.bdpn.pl

Field Stations in Lutowiska and Ustrzyki Dolne

Carpathians Fauna Research Station

in Ustrzyki Dolne

Muzeum & Institute of Zoology PAS

Ustrzyki Górne

Contact person. dr hab. Kajetan Perzanowski,

e-mail: [email protected]

Thank you for the attention

and welcome to Poland