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Outline
Pelagic fishes defined
Two main groups
Large pelagics
Swordfishes, billfishes (Family Xiphiidae andIstiophoridae)
Dolphinfish (Family Coryphaenidae)
A focus on TUNAS, Thunnus spp. (Family
Scombridae)
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Outline
Small pelagics
Belonidae
Caesionidae
Carangidae
Clupeidae
Engraulidae
Exocoetidae Hemiramphidae
Module 5. Pelagic Fishes
Menidae
Scombridae
Sphyraenidae
Trichiuridae
Note: The dom inant famil ies and representat ive
species may be modif ied to be region - or area-
speci f ic
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Pelagic fishes: Definition
Pelagic f ish es are diverse group of f ish es that l ive atthe upp er layer of the water co lumn
Module 5. Pelagic Fishes
Use simplified
diagram
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Pelagic fishes: Two main classifications
Large pelagics Small pelagics
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LARGE PELAGICS
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Pelagic fishes: Large pelagics
Family Xiphiidae
Family Istiophoridae
Upper jaw prolonged into
a long bill, f lat oval in
cross section
Upper jaw prolonged into
a long spear, roundincross section
Module 5. Pelagic Fishes
pelvic fin caudal keels
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Diagnost ic
character ist ics
Upper jaw p rolonged
into a lon g bi l l , f lat oval
in cros s-sect ion
Two wid ely separate
do rsal f ins in adults, the
f irst w ith 38 to 45 so ft
rays and the secon dwith 4 or 5 so f t rays
A large median keel
present on each sid e of
caudal pedunc le
A large and d eep no tch
on bo th upper and
lower prof i les of caudal
peduncle
Pelvic f ins absent
Xiphias gladiusSwordfish
(malasugi)
Family Xiphiidae
One large
median keel
on each side
Pelagic fishes: Large pelagics
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Family Istiophoridae
Is t iopho rus platypterus
Indo-pacific sailfish(malasugi)
Ist iomplax ind ica (Makaira ind ica)
Black marlin
(local name)
Makaira mazara
Indo-pacific blue marlin
(local name)about 15 obscurever t ical bars on b ody
pectoral f ins rigid,cannot be folded back
to sides of body
sail l ike first dorsal f in
Pelvic fin rays
long, almost
reaching to anus
Pelagic fishes: Large pelagics
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Family Coryphaenidae
Coryp haena equisel is
Pompano dolphinfish(dorado)
Coryphaena hippurus
Common dolphinfish
(dorado)
Pelagic fishes: Large pelagics
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Diagnost ic
character ist ics
bod y elongate and
fus i fo rm
two d orsa l f ins
f in lets p resent behind
dorsal and anal f ins
caudal f in deeplyforked
at least 2 sm all keels
on each sid e of caudal
peduncle, a larger k eel
in between in m any
species
in som e species,
corselet developed
(area behind h ead and
around pectoral f ins
covered with
moderately large, thick
Family Scombridae
Module 5. Pelagic Fishes
*60 cm FL *60-80 cm FL
*150 cm FL
*90 cm FL
Tunas
Other Scombrids *20 cm FL
*Common Length
Pelagic fishes: Large pelagics
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A focus on TUNAS, Thunnus spp.
Pelagic fishes: Large pelagics
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Thunnus albacares and Thunnusobesus
Pelagic fishes: Large pelagics
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a
Diagnost ic
character ist ics
a. pectoral f ins are
straight and st i f f
Thunnus albacares, Yellowfin tuna
(bariles/tambakol)
Pelagic fishes: Large pelagics
Module 5. Pelagic Fishes
b . two dist inct ly
raised r idg es
present in caudal
f in that form the
V or M notch
c . br ight yel low
f in lets wi th no or
s l ight black
edging
Dorsal view
b
c
M d l 5 P l i Fi h
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Diagnost ic
character ist ics
Thunnus albacares, Yellowfin tuna
(bariles/tambakol)
Pelagic fishes: Large pelagics
Module 5. Pelagic Fishes
e. r ight lobe of l iver
longer and
th inner than
round ed media l
and lef t lobes;
lobes sm ooth , nostr iat ions
d. swim bladder
occu pies ent i re
body cav ity
d e
M d l 5 P l i Fi h
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Diagnost ic
character ist ics
a. pectoral f ins formsmoo th arc wi th
floppy tips
Thunnus obesus, Big-eye tuna
(bariles/tambakol)
Pelagic fishes: Large pelagics
Module 5. Pelagic Fishes
a
b. centr al area of
caudal f in f lat with
two
inconsp icuous low
mou nds present
c. yel low ish f in lets
edged with black
b
c
Dorsal view
M d l 5 P l i Fi h
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Diagnost ic
character ist ics
d. sw im bladderoccu pies ent i re
body cav ity
Thunnus obesus, Big-eye tuna
(bariles/tambakol)
Pelagic fishes: Large pelagics
Module 5. Pelagic Fishes
e. l iver with 3
round ed lobes of
abou t equal size;
ventral surface
str iated
d
e
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To the untrained eye, Yellowfin and Bigeye
Tunas are often difficult to tell apart!
Dont worry, there arefeatures that we can look
at to help us in their
identification
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Yellowfin vs Bigeye Tunas: Comparison by
external characteristics
Bigeye (~ 34 cm) Longer, pointed pectoral fin
Irregular, white lines across
body
Large head, deep body, large
eye
Dusky colored tail
Yellowfin (~ 33 cm)
Short, blunt pectoral fin
Closely spaced markings of
lines and rows of dots in
chevron pattern extending to
insertion of pectoral fin
Shorter, smaller head, small,round eye
Yellowish tail
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Yellowfin vs Bigeye Tunas: Comparison by
external characteristics
Yellowfin (~ 56 cm)
Long, narrow body, small head, small eye
Closely spaced, chevron pattern of alternating lines
and rows of spots clearly visible to below pectoral fin
Bigeye (~ 56cm)
Large, deep head, large eye, deeply rounded body Long pectoral fin with thin, pointed tip
Vertical, widely spaced irregular white lines
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Yellowfin vs Bigeye Tunas: Comparison by
external characteristics
Bigeye (93 cm)
Rounded, deep body outline, large, deep head, largeeye
Long pectoral fin, thin tip pointing ventrally
Flat trailing edge of tail
Yellowfin (96 cm)
Long, narrow body, straight outline behind 2nddorsal fin
Thick, relatively short, blade-like pectoral fin
Small head and eye
Notch in center of tail
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Yellowfin vs Bigeye Tunas: Comparison by
external characteristics
Bigeye (99 cm)
Deep, rounded body outline, large, deep head, large
eye
Long pectoral fin, thin, pointed, wavy tip
Trailing edge of caudal fin flat
Yellowfin (104 cm)
Long, narrow body, straight behind 2nddorsal fin, small head and eye
Evenly spaced lines and rows of uniform dots
Noticeable V notch in caudal fin with two raised areas
2nddorsal and anal fins beginning to elongate
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Thunnus alalunga and Thunnustonggol
Pelagic fishes: Large pelagics
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Diagnost ic
character ist ics
a. colour less
pectoral f ins long ,
reach w ay past
second do rsal
and anal f ins
Thunnus alalunga,Albacore tuna
(tambakol)
Pelagic fishes: Large pelagics
Module 5. Pelagic Fishes
b. back edge of
caudal f in wh i te
or color less
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Diagnost ic
character ist ics
a. pectoral f ins barely
reach space
between dorsal f ins
Thunnus tonggo l, Longtail tuna
(tambakol)
Pelagic fishes: Large pelagics
Module 5. Pelagic Fishes
b. second d orsal fin
tal ler th an f irst
dorsal f in
c . 9 dorsal f in lets and
8 anal fin lets
d. body slender,
especial ly towardscaudal f in
e. fain t rows o f
co lour less spots
along bel ly
livr
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Other Scombridae
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Family Scombridae
Rastrel l iger brachysoma
Short-bodied mackerel
(hasa-hasa)
Rastrel l iger faughn iIsland mackerel
(alumahan)
Rastrel l iger kanagu rta
Indian mackerel
(alumahan)
Body very deep; head
length equal to body
depth; Gil l rakers very
long, vis ib le when
mouth is opened
Body moderately
deep; head length
longer than body
depth; Gil l rakers very
long, vis ib le when
mouth is opened
Body sl im; Gi l l rakers
shor t , not vis ib le
when mouth is
opened
2 rows of prominent spo ts at base of f i rst dorsal
f in , vis ib le from above
black blotch behind
pectoral-f in base
1 or 2 rows of sm al l , dark spots on sides
of dors al-f in bases
black spot on body n ear lower
margin of pectora l f insPelagic fishes: Small pelagics
Module 5. Pelagic Fishes
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Family Scombridae
Scomber austra lasicus
Spotted chub mackerel
(alumahang bato)
Scomber japonicu s
Chub mackerel
(alumahang bato)
obl ique l ines wh ich zigzag
crossed faint wavy l ines
thin wavy, broken lines which appear as speckling
sometimes with numerous dusky, rounded blotches
Pelagic fishes: Small pelagics
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Family Scombridae
Aux is rochei rocheiBullet tuna
(tulingan)
Auxis thazard thazard
Frigate tuna
(tulingan)
Corselet is well developed in its posterior part(more than 6 sc ales wide un der second do rsal-
f in origin)
Corselet is well developed and narrow in its posterior
par t (no m ore than 5 scales wide under second dorsal-
f in or ig in) OR does not reach second dorsal f in or ig in
Dark spot connected
Dark spot
separated
Pattern of 15 or mor e fair ly
broad, nearly vert ical dark
bars
Pattern of 15 or more narrow, obliqu e
to nearly horizontal, dark wavy lines
Pelagic fishes: Small pelagics
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4 to 6 horizontal lines on
belly
Family Scombridae
Euthynnu s aff in is
Kawakawa
(local name)
Katsuwonus pelamis
Skipjack tuna
(gulyasan)
Back spots
Narrow interspace
Pelagic fishes: Small pelagics
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Family Scombridae
Scomberomo rus commerson
Narrow-barred spanish mackerel(tangige)
Scomberomo rus gut tatus
Indo-pacific king mackerel(tangige)
Acanthocyb ium so landr i
Wahoo
(tangige)
Maxil la exposed
Maxil la conc ealed
Pelagic fishes: Small pelagics
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Family Scombridae
Sarda oriental isStriped bonito
(tambakol)
Gymnosarda unico lorDogtooth tuna
(tambakol)5 to 11 dark dorsallongi tudinal str ipes
Very w avy lateral
l ine
Pelagic fishes: Small pelagics
g
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Grammatorcynu s bi l inetaus
Double lined-mackerel(Alumahang Bato)
Family Scombridae
Scomberomorus queenslandic
Queensland school mackerel(Tanigue)
Scomberomorus semifasciatus
Broad-barred king mackerel
(Hasa-hasa)
Scomberomo rus munro i
Australian spotted mackerel
(Bariles)
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SMALL PELAGICS
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Family Belonidae
Tylosurus acus melanotus
Agujon needlefish(balo/kambabalo)
Ablennes hians
Flat needlefish
(balo/kambabalo)
Tylosurus crocodi lus crocodi lus
Hound needlefish(balo/kambabalo)
Strongy lura leiura
Yellowfin needlefish
(balo/kambabalo)
Bony appendage of some
specimens
Dist inct b lack lateral
keel on caudal
peduncle
Teeth point forw ard
No lateral keel on
caudal peduncle
Pelagic fishes: Small pelagics
g
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Platybelone argalus platyura
Keeltail needlefish
Tylos uru s gavialo ides
Stout longtomKambabalo
Family Belonidae
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Family Caesionidae
Caesio cun ing
Redbelly yellowtail fusilier(dalagang bukid)
Pelagic fishes: Small pelagics
Caesio lu nar is
Lunaris fusilier(dalagang bukid)
Caesio caerulaurea
Blue and gold fusilier
(dalagang bukid)
Caesio teres
Yellow and blueback fusilier
(dalagang bukid)
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Family Caesionidae
Pterocaesio chrysozona
Goldband fusilier(local name)
Pelagic fishes: Small pelagics
Pterocaesio dig ramma
Doublelined fusilier(local name)
Pterocaesio marr i
Marrs fusilier
(local name)
Pterocaesio pisang
Banana fusilier
(local name)
Bright yel low band direct ly
below lateral l ine
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Family Caesionidae
Pterocaesio randall i
Randalls fusilier(local name)
Pelagic fishes: Small pelagics
Dipterygonotus balteatus
Mottled fusilier(local name)
Pterocaesio ti le
Darkbanded fusilier
(local name)
Pterocaesio tr i l ineata
Threestripe fusilier
(local name)
Lateral l ine covered by
black strpe
Large yellow blotch
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Family Carangidae
6 Common Decapterusspecies, Roundscads (local:
galonggong) caught in Philippine waters
Decapterus macrosom a
Shortfin scad
Decapterus russel l i
Indian scad
Decapterus macarel lus
Mackerel scad
Decapterus m aruadsi
Japanese scad
Decapterus tabl
Roughear scad
Decapterus ku rro ides
Redtail scad
Dusky brown tail Yellowish tail Reddish tail
Pelagic fishes: Small pelagics
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Family Carangidae
Pelagic fishes: Small pelagics
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Family Carangidae
Pelagic fishes: Small pelagics
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Family Carangidae
Pelagic fishes: Small pelagics
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Family Carangidae
Pelagic fishes: Small pelagics
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Family Carangidae
Caranx ignob i l is
Giant trevallyMaliputo
Caranx melampygus
Bluefin trevallyTalakitok/Maliputo
Caranx p apuensis
Brassy trevallyPampano/Talakitok
Caranx sexfasciatus
Bigeye trevally
Maliputo/Talakitok
Caranx ti l le
Tille trevally
Bulubukto/Talakitok
Decapterus akaadsi
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Family CarangidaeDecapterus muro adsi
Amberstripe scad
Galunggong
Elagat is b ip innulata
Rainbow runner
Salmon
Megalaspis cordy la
Torpedo scad
Oriles/Pampano
Pseudocaranx d entex
White trevally
Duhay/Laparan
Scomb eroides lysan
Doublespotted queenfish
Lapis
Scomb eroides tala
Barred queenfish
Lapis
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Family CarangidaeScomb eroides tol
Needlescaled queenfish
Lapis/Talang-talang
Serio la dumeri l i
Needlescaled queenfish
Lapis/Talang-talang
Seriola lalandi
Yellowtail amberjackTrachinotu s bai l loni i
Small spotted dart
Pampano/Talakitok
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Family Carangidae
Ulua mental isLongrakered trevally
Piaka/Talakitok
Scombero ides comm ersonn ianusLongrakered trevally
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Family Carangidae
6 Common Decapterusspecies, Roundscads (local:
galonggong) caught in Philippine waters
Decapterus macrosoma
Shortfin scad
Decapterus ru ssel l i
Indian scad
Dusky brown tail
Pelagic fishes: Small pelagics
supramaxillamaxilla
supramaxilla
maxilla
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Family Carangidae
6 Common Decapterusspecies, Roundscads (local:
galonggong) caught in Philippine waters
Decapterus m acarel lus
Mackerel scad
Decapterus maruads i
Japanese scad
Yellowish tail
Pelagic fishes: Small pelagics
White membrane in upper jaw palate
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Family Carangidae
6 Common Decapterusspecies, Roundscads (local:
galonggong) caught in Philippine waters
Decapterus tabl
Roughear scad
Decapterus kurroid es
Redtail scad
Reddish tail
Pelagic fishes: Small pelagics
supramaxilla
maxilla
supramaxilla
maxilla
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Family Carangidae
Selar boop s
Oxeye scad(matangbaka)
Selar
crumenophtha lmus
Bigeye scad
(matangbaka)
Scutes thick and long
Scutes smal l and sh ort
Big eye
Pelagic fishes: Small pelagics
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Family CarangidaeAlepes djedaba, Shrimp
scad(salaysalay)
Atu le mate, Yellowtail scad
(kalapato)
Megalasp is co rdyla, Hardtail
(oriles)
Selaroides leptolepis, Yellowstripe
scad(salaysalay ginto)
Yellow str ipeBlack opercular
spot
Term inal ray of
dorsal and anal f ins
not f inlet- l ike
Term inal ray of
dorsal and anal f ins
f inlet- l ike
Large scutes
Finlets
Pelagic fishes: Small pelagics
Family Carangidae
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Family Carangidae
Jacks, Trevally (local: talakitok)
Alect is indicu s
Indian threadfish
(local name)
Caranx
ignobi l is
Trevally
(local name)
Carangoides
ferdau
Trevally
(local name)
Ulua mental is
Longrakered trevally(local name)
Gnathanodon
spec iosus
Golden trevally
(local name)
Uraspis uraspis
Whitemouth jack
(local name)
Pelagic fishes: Small pelagics
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Elagatis bipin nu lata
Rainbow runner
(salmon)
Scomberoides lysan
Queenfish
(lapis, dorado)
Seriol ina nig rofasc iata
Blackbanded trevally(tonto)
Naucrates ductor
Pilotfish(local name)Parastromateus niger
Black pomfret
(local name)
Family Carangidae
Pelagic fishes: Small pelagics
No scutes
No scutes
No scutes
No scutes
18-19 weak s cutes ,
form ing sl ight keel
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Family Clupeidae
Sardinel la gibb osa
Goldstripe sardinella(tamban)
Pelagic fishes: Small pelagics
Sardinel la f imb r iata
Fringescale sardinella(tamban, lapad)
Amblygaster si rm
Spotted sardinella
(tunsoy)
Sardinel la lemuru
Bali sardinella
(tunsoy)
Dark spot on
dorsa l f in or ig in
Blackish caudal f in
Dark black spot at gi l l cov er edge;
go lden spot beh ind opercu lumLateral series of 10 to 20
dark b lu ish go ld spots
golden midlatera l
l ine down f lank
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Family Clupeidae
Dussum ier ia acuta
Rainbow sardine(tulis)
Pelagic fishes: Small pelagics
Dussum ier ia elopso ides
Slender rainbow sardine(tulis)
Anod ontostoma chacunda
Chacunda gizzard shad
(kabasi)
Nematolosa n asus
Blochs gizzard shad
(kabasi)
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Family Clupeidae
Sardinel la longic eps
Indian oil sardine
Tamban/Tunsoy
Amblygaster le iogaster
Smoothbelly sardinella
Tamban
Herlotsichthys quadrimaculatus
Bluestripe herring
Tunsoy
Spratel lo ides robustus
Fringe-scale round herring
Spratel lo ides graci l i
Silver-stripe rounde herring
Dilis
Sardinella melanu ra
Blacktip sardinella
Tamban/Lao-lao
Family EngraulidaeModule 5. Pelagic Fishes
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Family Engraulidae
Anchovies
(dilis, tuakang)
Pelagic fishes: Small pelagics
Stolephorus
Pre-pelvic scu tesPointed
maxi l la
Engraul is
No pre-pelvic scutesRoundedmaxi l la
Encrasichol ina Thryssa
Pre-pelvic scutes
Pointed
maxi l la
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Family Engraulidae
Sto lephorus commersonn i i
Commersons anchovy
Munamon/Dilis
Sto lephorus ind icus
Indian anchovy
Dilis/Munamon
Stolephorus ronqu i l lo i
Ronquillos anchovy
Tuakang/Dilis
Encrasichol ina punct i fer
Buccaneer anchovy
Bolinao
Encrasichol ina devisi
Devis anchovy
Bulinao/Dilis
Stolephorus heteroloba
Shorthead anchovy
Dilis
Family Exocoetidae
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y
Flyingfishes
(bolador)
ParexocoetusExocoetus
HirundichthysChei lopogon, Cypselurus,
Progn ich thys
Origin o f anal f in 3 rays ormore behind or ig in of d orsal
f in
Or ig in of anal f in sl ight ly
before, und er or not
more than 2 rays behindor ig in of dorsal f in
6-7 Scale rowsPelv ic f ins shor t
Pelagic fishes: Small pelagics
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Family Exocoetidae
Chei lopogon cyanopterus
Margined flying fish
Bolador
Chei lopogon furcatus
Spotfin flying fish
Bolador/Iliw
Chei lopogon nigr icans
Blacksail flying fish
Bolador/Iliw
Chei lopogon poeci lopterus
Yellow flying fish
Bolador/Iliw
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Family HemiramphidaeRhynchorhampus georg i i
Long billed half beak
Bugiw/Siriw
Hemirhampus far
Black-barred half beak
Bugiw/Susay
Hemirhampus lutkei
Lutkes halfbeak
Buging/Kansusuwit
Hemirhampus robu stus
Three-by-two garfish
Soroy/Sasa
Hemirhampus af f in is
Tropical halfbeak
Siriw/Kansusuwit
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Family Menidae
Mene maculata
Moonfish(chabeta)
Pelagic fishes: Small pelagics
Body ext remely comp ressed
Pelvic f ins
Anal f in
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Family Sphyraenidae
Sphy raena barracud a
Great barracuda(torcillo/barracuda)
Sphy raena obtu sata
Obtuse barracuda
(torcillo/barracuda)
Sphyraena jel lo
Pickhandle barracuda
(torcillo/barracuda)
Sphy raena pu tnamae
Sawtooth barracuda
(torcillo/barracuda)
Double
emarginate
caudal f in
Caudal f in
yel lowish
Caudal f in
yel lowish
Caudal f in
largely
black
Dark m arkings crossing latera l l ine on body
Dark bars cross ing latera l l ine on body
Obl ique dark bars on upper hal f of
bod y, not cro ssin g lateral l ine
Pelagic fishes: Small pelagics
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Family Trichiuridae
Tr ich iurus lepturus
Largehead hairtail
(espada)
Pelagic fishes: Small pelagics
Ribbon-l ike
Caudal f in absent
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Family Trichiuridae
Tr ich iurus ganget icus
Ganges hairtail
(espada)
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END of LECTURE
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REVIEW GAME(modify using important fish families and species in
area)-work as a group
-correctly identify the species shown and accumulate
points!
To which FAMILY do these f ishes belong? (1
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To which FAMILY do these f ishes belong? (1
point)
1
T hi h FAMILY d th f i h b l ? (1
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To which FAMILY do these f ishes belong? (1
point)
2
Give the (a) Fam ily, (b) Common Eng lish name,Module 5. Pelagic Fishes
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and (c) Scient i f ic name of the f ish shown (3
points)
3
Give the (a) Fam ily, (b) Common Eng lish name,Module 5. Pelagic Fishes
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and (c) Scient i f ic name of the f ish shown (3
points)
4
Gil l rakers very lo ng and visib le
when mouth is opened
1 or 2 rows of sm al l , dark spots on sides
of dors al-f in bases
black spot on b ody near lowermargin of pectora l f ins
Head length longer than bod y depth
T hi h FAMILY d th f i h b l ? (1
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To which FAMILY do these f ishes belong? (1
point)
5
Give the (a) Fam ily, (b) Common Eng lish name,Module 5. Pelagic Fishes
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and (c) Scient i f ic name of the f ish shown (3
points)
6
Give the (a) Fam ily, (b) Common Eng lish name,Module 5. Pelagic Fishes
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and (c) Scient i f ic name of the f ish shown (3
points)
7
To which FAMILY do these f ishes belong? (1
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To which FAMILY do these f ishes belong? (1
point)
8
To which FAMILY do these f ishes belong? (1
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To which FAMILY do these f ishes belong? (1
point)
9
Give the (a) Fam ily, (b) Common Eng lish name,Module 5. Pelagic Fishes
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and (c) Scient i f ic name of the f ish shown (3
points)
10
pectoral f ins are
straight and st i f f
two dist inct ly ra ised
r idges present in
caudal f in that form
the V or M notch
br ight y ellow f in lets
wi th no o r s l igh t
b lack edging
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HOW DID YOU DO?