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iLLiii ?f1i1JLL iafvn?f rn?fJ1 VEGFFATTON PATTERNS IN THE TROPICAL SEMI-EVERGREEN FOREST AT SAKAERAT, NAKEION RATCHASINA 1!1 uj:;viiff:3ii SARAYUDIJ BUNYAVEJCHEWI•N f'o 4 .00 i7 1i f11 iiiih1 It LFfl n•vm4M 10900 2S3i SILVICULTURAL RESEARCH DIVISION, ROYAL FOREST DEPARTMENT (1 'I CHATUCHAK, BANGKOK 10900 m. 0. 2537
15

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Page 1: f'o - app.dnp.go.thapp.dnp.go.th/opac/multimedia/ebook/37_539/37_539.pdf · revealed that there appeared the positive relationship between basal area per hectare with magnesium, available

iLLiii ?f1i1JLL iafvn?f rn?fJ1

VEGFFATTON PATTERNS IN THE TROPICAL SEMI-EVERGREEN

FOREST AT SAKAERAT, NAKEION RATCHASINA

1!1 uj:;viiff:3ii

SARAYUDIJ BUNYAVEJCHEWI•N

f'o 4 .00

i7 1i f11 iiiih1

It LFfl n•vm4M 10900

2S3i SILVICULTURAL RESEARCH DIVISION, ROYAL FOREST DEPARTMENT

(1 'I CHATUCHAK, BANGKOK 10900

m. 0. 2537

Page 2: f'o - app.dnp.go.thapp.dnp.go.th/opac/multimedia/ebook/37_539/37_539.pdf · revealed that there appeared the positive relationship between basal area per hectare with magnesium, available

91 t ) Hf'Ii : • AflBiI1J

; , Forest Dcpavtmeit Librt'y

Page 3: f'o - app.dnp.go.thapp.dnp.go.th/opac/multimedia/ebook/37_539/37_539.pdf · revealed that there appeared the positive relationship between basal area per hectare with magnesium, available

:ann?f fliTfl?(1Ji

VEGETATION PATTERNS IN THE TROPICAL SEMI-EVERGREEN

FOREST AT SAKAERAT, NAKIION RATCHASINA

ifr#fi1 P11 I, msa-lp -xA lu

SARAYUDH BUNYAVEJCIllMIN

f'o 1!00

nIj n-2flh'U:i

1I'n flLYWi 10900 I

SILVICULTURAL RESEARCH DIVISION, ROYAL FORESTDEPARTMET1

CHAT(JCHAK, BANGKOK 10900

2537

&ni&i'ti

IIll Ilifi IID IIDHII DI III llI ID II 00003835

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VI

LL 1JLL ?1ZLL11'flZ' 11T3J1

VEGETATION PATTERNS IN THE TROPICAL SEMI-EVERGREEN

FOREST AT SAKAERAT, NAKEJON RATCHASINA

911MTh1UVwL'-)WW'-JU

9 14

SARAYLJDH BUNYAVEJCHFMIN

iQrLLJ 4iu'nitrimi1iiU rrnhUi i, n 1nwi i000 9 19

SILVIULTURAL RESEAWH DIVISION, ROYAL FORESTDEPARMEff

CHATUCHAK, BANGKOK 10900

Twelve stands of the tropical semi-evergreen forest at

Sakaerat Environmental Research Station, Nakhon Ratchasima province, NE

Thailand were studied for structure and to determine the relationship of

stand to environmental factors.

The tropical semi-evergreen forest can be divided into two

dominance-types ; Hopea ferrea type and Shorea henryana type. Basal area

and density of all stems (Dbh> 10 cm.) were slightly different between two

types ( 30 m2/ha and 562 trees/ha in Hoja ferrea type and 27 1n2/ha and

514 trees/ha in Shorea henryana type).

Size-class analysis indicated similar structure of two

dominance-types. Both were well described by a negative power curve and

negative exponential distribution. Size-class distributions of

individual species exhibited variable patterns. Regression analysis

revealed that there appeared the positive relationship between basal

area per hectare with magnesium, available moisture capacity, phosphorus

and bulk density. On the other hand silt + clay, soil pH and slope

produced the negative relationships with stand basal area.

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-2 —,

ICA ' Q# ' 4, .v , V

nLLE)3JZLn'T1

.r-5-1?fz1J1 20x50 3j. mfl1J 2 LLi1E

•mn 12 fb cluster analysis AZI 44

2 J f LJ1111 (Hopeaferrea type)

.L1Z'J L ThJ1 (Shorea henryana type) I

2 V

LL11 (dbh>1O J.) 1flfls Ql

L.

JfE ( 30 3J. /Lr1fThLW ua.t, 562 MAOMLRI 4 -, 2 V . V _

LthflJL1flJn1J 26.9 Ji. JIeIflLLW5 LLZ 514 JJ1flJLL 31

V Lfl1t h&JIJ

negative power curve cn

61 fl1t nI S ]LL1J1J 1 1 1LL.fl1JiJ

LVJ silt+clay pH 2zi iW1i

iu1nTJ 1 111iLL

-, V G V , V G 1J4'iiu1JL L n t3JcJz1EbJ . b-u . 19 V 4 V V

UUE

J- L 35 J . • 3fl 111 W_

1 V

111 19 9 JjjJjUqO 6_~ U

a LLa\l U1WSi fU1i . wl~cj"L'o~ljj I'A U

n -1bJLL9flbJ inn Uaw epiphyte "Zi £ . V 2

fl1 LLflid' LLfl]'J1CU 36% (30 fl.J. ) dE

.q_, V E J V

ThJ1 (fl1i LLCl, 2523) flL1fl1flW1 L411flJLL1 c c V V ,

UU'-j

V' 1) lu n -1,7 9 fl-j~ -I fl 2)

, Wfl11J

-5MUIT'5 wWM -M 41 f) MY 011ALM fl LlZ3J

1 LTh -1 1-I -1

6 € E V V

Lt-u11Thnl vnrninin LiHni iini•t. nn' .

V

.111?fJ1 nI flflJ1J 2526 111JILZ ..

Lc . ii J j\J LU1fl1 .

V

Korat group L1LthA clay loam., sandy loam t

Page 6: f'o - app.dnp.go.thapp.dnp.go.th/opac/multimedia/ebook/37_539/37_539.pdf · revealed that there appeared the positive relationship between basal area per hectare with magnesium, available

- 3 -

sandy clay loam 11thLLi 11240 jj.i1 iifl91th1fl 1 4M ICA 14

171IL1,11 ~Jll L IMau3ju 41nrn 26.2°C fl1LJ 21.6C IUL,

1_flfJ 28.7 C lutIMUMIMIJ Si Si

LL JN1fl1 20x50 J. 2 LLJ'bJ 111 12 M~Ul MJ@IULL~@Ifl

j J 1J Si Q,

flEflflfl91 1flI LLW1rn4flfli1 L LL LLV1]LLJ\1 L 1fl1

lOxlO 3j. 10 LL1i TJ 20 LLflJlJ fl h19hJflZ 19

•LJEJnLLLi 4.5 t.3jAUIfl

LL 111IULL~awm~UAOWIU,~IVJU 2 MM Ln1JcWnJ1

0-5 10-15 20-25 30-35 LLaw 45-50 41f.3J. nrU

flCUMAM11 (bulk density) core method XZ4 0

Ui@U 0

111LLfV51fl1 2 UZO.S i.3J. 4CM.1L1JJ1fl

Ei texture Ifl?j hydrometer method iiinin

(pH) zji:ifl %U@~111~-~U 1:1 LL i pH meter il1i 19

WakIey and Black rapid t itrat ion method exchangeable K t Na 5'/fl

flame photometer exchangeable Ca Lt Mg 1' atomicIflul"Y

absorption spectrophotometer iThnn available P Bray's II f

cation exchange capacity ammonium acetate pH 7 i13:4,ljlm available

moisture capacity 15 bars

4:111 V IoMiall 111111 1J1fl

* v 0

LL11 1L31J (fl relative) 1X1 importance value 4t#

index d @- VW 4ULL'1 LULL ZJ LJ fl1Lfl JW' L .LVfl L'JIT cluster

9 Si

analysis

CJLL -;ri i fUCJ

14 '1'' ' cV 44 2 1 LL .LJ

1 c:

:LflflJ fl LTh.J Uflfl1ffl1 jS i ti

2

( secondary growth) 1L1i1fl1 J'bJ1161 Si 9

LL1i51Jfl 11 2

26.9-30.0 J. /:L L111LL1' Lfl 1LL11J L 19

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lL -

.

.!JJ ( 27. 5 J. I L nLr) 1L UJaL1hflJ nfii Lith w1

(Table .1) L

Cw

U@Un-nolflmed~l un';WIM UaW.,3JWU*nAU'W)01 LnaLfl?_jIn4%'*U~Ifl'AULVIOn 14~. i:L Xi 19

2i l4 2 , 2 11M U Lfl1 Vfl UZL1 3J1 'flY1J L1flfl11ThF1 U

fl 151 U -L -~un -iignl±~-) ~POW 9' 2 '1t 4 Q.'

L1:11'1 fl1LV1ZJflJ LLLflWN1u1UJ1flfl I1J1CJ11 I

9,

UcJrnL1hJin u,ii LJtUWfl VI LW1flfl Lflifl V

oGV 9.' , o

im'Malll €V

1LLWTh1fl1flJ1flWfl

St...'C4"QI '0

44Bun.yaveichewin (1986) n1L1i'1 cluster analysis LL1J'

izkLfrfl?f @@nAu 2 91fl3i flEl 9Ifl3mQw,.Lfluumu (•Hop9a ferrea type) L•flJfrL (Shorea henryana type)

51fl3iflwa,

V

1. Lftrt L .L J11 L IUW~13J LflL 3J L JJtEJ 19

(Shorea henryana Pierre) fl1Jfl (Irvingia malayana 011'v. ex A. Benn . ) 1ifl

1 V

•LIill-flfl (Largerstroemia duprreana Pierre) 1 (Eugenia sp.) LLL .G q, V

(Dialium cochinchinense Pierre) (secondary dominance) ICU w

niwnrn 400-560 3j, •L11N111w:L •3j

V*j

iimnn 1zN?f1ulh sandy loam, sandy clay loam, loam i clay le

loam L9~N (dbh>10 ) 30.01 3J.?-/ L rInUOTI V I i

LLNI., 562 /:LflL J13J 28-35 J.

q1 •

20 L L 1Jt JE3' L1ilL'M UJ L OIU

(Dipt:rcarpus turbinatus Gaertn. f . ) anom (Parkia suznatranaMiq . ) LL 44 IQI Cm

1 d ThrW1LiIJ I, L 1i t, . i i 30-38

J. •3Tht11 (•dbh>10 . . ) 26.88 J. AfInL011 LL1J1LLWL (d.bh>10 1.3j.)

V9i V V Q1 4 i •

514 'rni;/Le1nLLr5 400-700 J.

•4/ 6 ,

11nflh1 Lzfli1lLnfl fl1 flJ L L1 fl Jfl L fl1

G

~114@191M G 9/

3. ecotone LLL L1]LVWLkflJ ' fl1i'J1flL

44 .-, 'r V q., 4J £ \4 V

Boo J. n1i L 1Jwd LflL11 LL LJ1 11]WW UJL1i 19 • 4, ./

t 4

1zwflJ L nfh. t. iinrnci i.

,.gV •

q-? wIDU

.LL: ecotone uvincv

ILI ' 1111 11Thw1w1 1JWt1LL11

. V • •d ' • -' .V V V44

~um4ur4-iffumm,3:

(Table 2) izjiz

fliL1flflI' LJ1L9/

L: nJ1l L1f1]flfl

'Tcn- i 30 ?J. 98 wE3vnrni 17.4 h flLLfl lufdm

/ ' Qf `Q1 V 44 • -, • •

L 3JVWJ .66 /: f flLL 1A'EWfl1J 12.7 % 1JW1LL1flJ 'Th1

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- 5 -

Table 1. Basal area (X + SD) of various forest type in Thailand

Basal area •(m2/ha)

Forest type dbh>10 cm

1. Dry dipterocarp forest (Bunyaveichewin, 1983a)

Shorea siamensis type 20.32 -I- 7.48

Shorea obtusa type 16.74 + 5.55

Dipterocarpus obtusifolius-shorea obtusa type 23.51 ± 7.96

Dipterocarpus tu1xrcu1atus-Shorea obtusa type 23.88 + 8.09

Pine-dipterocarp type 24.39 + 5.24

2. Tropical dry deciduous forest (Bunyavejchewin,

1983b)

Tectona grandis type 38.38 ± 10.53

Lagerstroemia calyculata type 33.12 + 14.30

3. Dry evergreen forest in Nain Prom basin (Sahunalu

et al., 1979) 27.5 .4. Tropical rain forest at •Khao So (ParRtoop, 1980)

valley site 25.0

hill side 32.0

ridge 41.0

5. Coniferous forest at Phu Kradund (Bunyaveichewin,

1979) 13.21

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!/

- 6

Table 2. Number of trees in each size-class of the dominance-types of the

tropical semi-evergreen forest . Number within parentheses give

the percentage within each size-class (dbh>10 cm)

dbh Hopera ferrea type Shorea henryana type

(cm ) (trees/ha) (trees/ha)

4.7- 10 592 775

10-• 20 350(62.28) 365(71.01)

20- 30 114(20.28) 83(16. 15)

30- 40 46(8.18) 25(4.86)

40- 50 28(4.98) 15(2.92)

50- 60 12(2.14) 6(1.17)

60-70 4(0.71) 8(1.56)

70- 80 1(0.18) 5(0.97) 80- 90

5(0.89) 20.39

90-100 1(0.18) 1(0.19) 100-110

- 2(0.39)

110-120 - - 120-130 - - 130-140 - 1(0.19)

140-50 - 1(0.19)

150-160 - - 160-170 - - 170-180 1(0.18) -

total (dbh> 10 cm) 562 .514

total (d.bh>4.5 cm) 19,154 19289

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- 7 -

V

4 2'. , c1:: I, flflZJ L JE3'

J1Jn 0-15 t.3j.

LL1LLn:i?i1'IAILLZ*11"~Iu Table 3 ni 1n11 pH LL

LJt E1 (I~LLOIM;m

1rn1att4 ThJ

LL.-OnUAM LLawIIL'flV3j) LLZFfl •CEC 1 J L 1JZ1Yfl flLUfl11J LfJJ

.Qd 1 Q-' Q, £ 2 , 2 • 44 1191Ernn- LI11J twinaijmurmuwflu@umi ?1flfl

9 t G L11flh1fJ LJJ1 pH AUOIIUULDI flU113AflTh.Lfl'LLflbiflJ

49, • 2i i ' U-3 UJ Ln1fl

011fl( niniirlJ i. r1

9 1W

4 , 0 , ICU

LL1Z t a an ( selectivity

mechanism) 1968) L1Jtthfli' 19

IQI Si 9 LL1Z

LThfli1LUJfl (climax) LLZbJflfl

aiiThJt L ]iJ J-shaped nirH Cr

negative exponential curve flTMUIULL~aA, 11 1fl1flJLYIfl1i

(Leak, 1965; Miii ler, 1982)

negative power curve (Fig.1) J1iWflJ

9, '12/' l6 G / '4V . :k- ii -nfl uJ1J L1Lfl 1fl11ThJ1flflThflL Ufl1J Lt?1fl1 Lu1fl1fl1ZJL1 Si J ii

E) 4 c •G a 4 q, Wa LL Qe

31 01 -1 3AUt U -1 ONI -1 3AOI'U4"eU @11'4u~'ULL

qj 9-, QI '4v4 4 O6)9iq.

UJn ii inimrnJin

Robertson

et. al (1978) negative power curve U11fl

flIlh-wiu- 19 1W

ICU IQ.

W1LLLLfl1Jfl S '1]LL1J1J

(Fig. 2) ( 1) negative power curve L ? 01 l (2) negative

exponential curve L1 flLflflZfl (Hydnocarpus ilicifolius King) ici 0 V 2i'42,':4 c

(Waisura trichosternon Miq. ) ( 3) iniminrn Un1J1nl Lan -.mnit(utuifl

Jczi mul4 0 V

1LW3jIPt aflal I.?11 L flV3JflU@I flE311' (Quercus

0

21, IQd I 0

rnyrsinifolia Bl.) (4) 1 L' flJfl L (5) 1 19 Im

41 9, I

1nnilld ifl I ZJfl L LL1Jfl t 1J31T1J1 (Lagerstroemia

duperreana Pierre)

n1,1 .

09IT131'

(3) L11.L 1 1 11Jfl

24 9,

'jti L3J LflLfl'LL1M LJLLZ

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Table •3. Mean standard deviation, and ranges (numbers within parentheses) for soil properties

(0-15 cm depth) and topographic features of the three groups in the tropical semi - evergreen forest

Soil properties and topographic features Hopea ferrea type Shorea henryana type ecotone

bulk density (gm/cc) 1.16 ± 0.06 1.04 ± 0.08 1.01 ± 0.04

(1. 1.1 - 1.25) (0.92 - 1.12) (0.98 - 1.04)

silt+clay (%) 56.40 6.25 54.60 11.12 64.50 0.71

(48.5 - 63.5) 08.5 - 64.0) (64.0 - GS)

pH 4.51 ± 0.78 3.69 ± 0.57 3.25 ± 0

(3.85 - 5.80) (3.00 - 4.20) (3.25)

cation exchange capacity .(meq/100 gm) 7.02 + 1.83 .8.49 + 2.02 6.36 ± 0.65

(4.25 - 8.45) (6.20 - 11.15) (5.90 - !6).82

organic matter (%) 3.19 ± .0.90 3.71 ± 1.55 3.14 ± 0.31

(1.74 - 3.96) (2.61 - 6.41) (2.92 - 3.)36 available phosphorus (ppm) 5.3 1.04 4.70 + 1.99 3.0 ± 0

(4.0 - 6.5) (2.0 - 7.5) (3.0)

exchangeable cation (ppm) potassium

89.50 ± 14.01 108.30 28.55 147.75 + 47.73 (68.5 - 106.5) (90.0 - 157) (114.0 - 1)81.5

calcium 114.20 ± 56.72 287.70 ± 179.47 132.0 ± 31.11

(61.0 - 201.0) (124.0 - 568.5) (110.0 - 15)4.0 magnesium 148.80 ± 55.47 209.50 101.16 122.15 ± .50.56

(88.5 - 210.0) (149.0 - 387.5) (86.5 - 158).0

sodium 16.6 ± 5.803 17.20 ± 3.78 14.25 ± 1.06

ho.o - 24.0) (13.0 - 23.0) • (13.5 - 15.0) available moisture capacity (%) 11.85 ± 1.14 8.71 ± 1.87 6.76 4- 1.13

(0.31 - 13.26) (7.58 - 11.89) (5.96 - 7.56)

elevation (m) 446 ± 20.74 534 ± 63.87 560 ± 0

(410 - 460). (460 - 610) (560 - slope (%) . (0 - 15) (5 - 35) (0)

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L4I 0°J4i f4PuOU

teqiaJ4) 1i!suaO

Page 13: f'o - app.dnp.go.thapp.dnp.go.th/opac/multimedia/ebook/37_539/37_539.pdf · revealed that there appeared the positive relationship between basal area per hectare with magnesium, available

- 10 -

(EqIsaaI3) 11!SU3O

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Page 14: f'o - app.dnp.go.thapp.dnp.go.th/opac/multimedia/ebook/37_539/37_539.pdf · revealed that there appeared the positive relationship between basal area per hectare with magnesium, available

'G

Jil LLJ Lfl J •LflJTfl1 qj

V 2' 9) 2' V '. ' 4

LJfl L1E1

w,_9- L JflWfl flLflJfl9]E L11fl Lu -

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(Y) •LLi (dbh>4. 5 t. J. ) _4 ., - ' --

(X) 0-5 ?I. J. LLflflJi L'W

Lth1

Y = 7.0370 + 0.0022 M - 0.0383 silt+clay - 1.4834 pH - 0.069

slope + 0.0853 available moisture capacity + 0.3250 P

+ 4.6698 bulk density ± 0.3717

(R4 0.949 P<0.05)

V

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clay pH LL -1J -1 w IOLIJUTMqi1@11~ I 10 -1 fl M D n -17 L -;~I L ~1 IJ

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_ 12 - :• FLft Lfl7

st.p L EF1-ZI F) -3

0 . ' 44

4flt JC?1 UUqrf fl5J1W, 1i?(1 U173jIUU1q q flfl

UUII LL @ LI 0 1 Trti. 2522. n n iii S

LZJ1TW'J 1JflJ. L3il1 63.. 91

M-1 09UM. 62 MUI.

n9va - n1, LLJL14?1 fl'fl. 2523. _ '.

xzi

- Jw LflT5 1 1J

69. 11Lnh1, fl'VYVWi. 40 MIO

n4 2523. ThJYTh. 1 %jr:bi _1j1zJ L n1Lwz9r5, fl VYVW1. 88 ln.

Black, C.A. 1968. Soil-plant relations. John Wiley and Sons, New York,

U.S.A.

Bunyaveichewin, S. 1979. Phytosociological structure and soil properties

in Nam Pong Basin. M.S. Thesis. Kasetsart Univ. Bangkok. 123 p.

Bunyavejchewin, S. 1983a. Canopy structure of the dry dipterocarp forest

of Thailand. Thai For. Bull. 14:1-93.

Bun.yavej chew in, S. 1983b. Analysis of the tropical dry deciduous forest

of Thailand, I. Characteristics of the dominance-type. Nat.

lust. Bull. Slain Soc. 31(2),.109-122.

Bunyavei chew in, S. 1986. Ecological studies in the tropical

semi-evergreen rain forest at Sakaerat, Nakhon Rachasima,

northeast Thailand, I. Vegetation patterns. Nat. Hist. Bull.

Siam Soc. 34 (1):35-57.

Leak, W.B. 1965. The J-.shaped probability distribution. Forest

Science 11:405-409.

Muller, R.N. 1982. Vegetation patterns in the mixed mesoph.ytic forest

of eastern Kentucky. Ecoloy 63:1901-1917.

Robertson, P.A., G.T. Weaver, and J.A. Cavanaugh. 1978. Vegetation and

tree species patterns near the northern terminus of the

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