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(K. F. Kenneally 5412) (phot. K. Liedt).

142

Figurc L Ptilotus kcnneullyanus Benl.

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turbinate ca I mn long, up to 0.4 mm wide, surrounded at base by a densetuft of simple bristly hairs about 0.7 mm long and more or less i iregularlyseptate.

, Tepals conspicuous with their glabrous red shining apices not exceeded bythe copiolls external snow-white hairs, 3-nerved with a distinct median veinand two fainter marginal ones; the broad ridge above the tube glabrous in itslower part of about 1 2 mm. Outer tepals 7-1 5 mm long and 0 8 mm broadnear the middle, the apex 2.8 3.1 mm long, glabrous ir its upper 2 mm,subspathulately dilated io 1 2 mm, apex rounded to sub-trulcate, minutelyserrate; dorsal hairs articulate (finely denticulate to vertici l late), up to 2. 8 mmlong and markedly decreasing in length towards the base (Figure 2 D); internallyglabrous tlrroughott. Inner tepols 5 8,6.5 mm long and ca 0.5 mm broadnear the midd le ; the apex I . 2 I .8mmlong,g labrous ,d i la ted to0 .8( -1 .0 ) m[1 ,but mostly inrolled, subacute, scarcely reached by the dorsal vestiture; insidewith a faint beard of thinner, crisped. nodose hairs along both (rarely one)edges above the tube; the basal ridge nore sharply bordered (Figure 2 E).

Two adj acent ,r/ameirs consistently ferti le, f i laments flattened, 1.62.2 mmlong, ca 0 1 mm broad in the middle, subulate above and markedly (to 0.3-0 45 nim) dilated below; the thtee stqminocles mostly shorter subequal, bearinga mirrute buttonJike rudinentary anther, filiform and curling upwards at tineiand often more abruptly broadened downwards to the same extelt as the fertileones (Figure 2 F). Stamens and staminodes fused into a turbinate cupulastrongly adlate to the pedanth tube, with a free membranous, sometimesasymmetrical ring to 0. 25 mm high, with fascicles of curly nodose hairs 1 .2 mrrrlong outside at the base; sinuses acllte; no pseudo-stamino d,jal lobes. Anthers\ e r ) b r o a d l y e l l i p t i c . c a 0 . 1 x 0 . 2 m r n .

Prslil moderately stipitate. O}a/.)' more or less ch"rb-shaped to subglobose,finally up to 2.4 nrn long, the 0.3-0 4 rrrm lorg stipes included; pubescenceof rigid articulate hairs up to 0.3 mm long, occupying the upper halfofthejuvenile ovary, l imited toward its top in mature stage. Style1.0,l.3 mmlong,to 0.1 mrn in diameter, eccentrical and always shortly hairy on its thickenedbase (Figure 2 G). Stigma capitellate becorning inconspicr.rous with the age.Habitat. "Common on red, pisolit ic ironstone amongst spinifex clumps"(K. F. Kenneally).

Specimens exaninedApart from the type material (five large specimens each consisting of a branchsuch as shown in Figure I A and two small specimens such as in Figure I B)tlere is a previous collection of J. S. Beard (4105, 13 May 1965, DampierDowns, Edgar Ranges), comprising four specimens, two of them (on a singlesheet) Iodged at NSW, one at PERTH and one at K. These samples areteduced variants ('tmall shrub, 3 in. 6 in.") of the new taxon, evidently grownin a more barren soil, dillering from Kenneally's younger specimens in liavinga thicker stem (NSW) and larger leaves (K, NSW, PERTH); the pubescenc-and floral features match those of the type material.

Discussion. The resemblance to P, oboy(ttus (Gaud.) F. Muell. seems to bemerely a supcrficial one, as already indicated in the diagnosis above. It isdifficult to assess whether the plants are perennial, since the smaller isotype(7. 5 cm tall, 10 cm across) has a considerable number of inflorescences (FigureI B), and the taxor thus displays a rarge of variation in size urcommon jnpereuials in general and unknown in P. oboratus in particular.

The upper floriferous branchlets may form a right angle as is often thecase with P, oboyatus which, however, produces only terminal corymbosepanicles, not isolated spikes in lower parts of the branches as shown in Figure1A. InP. oboyatus yar. oboyatus with jts dense. persistent tomentum the h;irs

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346

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Peridnth elongated, rigid, erect but ldter divergerlt, the exposed glabroustcpal ends coloured, base constricted lo c \rnrl i turbinute rube, hardened,0 8 I mm long. obscured by rhe stif l dorsal hairs (to 2 mrn long) of the basaiDLlOeSCenCe

, Tepals l itear, invested outside along the certre with firrn, straight, joiltedand subdenfcLrlate ltairs, up to 7.5 mm long in lower half but never ieachingthe. apices (Figurc 4 C and ,tr D); strongly short-cil iatc at the margins wit[rlcl icate hairs ca l-l .5 rrm long (increasing somewhat i l length toward thebase) arld shurply de nd ro id- r errici l latc; these underlying the dorsal vestiture forthe most part and o1len turned in from the margils against inner surfirce of thetepals; t luee nerves fairly conspicnous inside especially in the thickened base.

Otrter tepals (Figttte 4 C) J.5 I .7 cm long and up to I .3 mm broad abovethe tube; apex glabrous, reddish, ca 2.5 mm long, often prolonged anotherI 1'5 mnr down tlte margins, subspathulately dilated to about 2 mm butmostly incurved, ;rpically erose-serrulate with a centraj mLrcro; thc laninagreenish, glabrous inside except for the marginal cil ia meltioned above, thecentral nerve prominent inside.

Inner tepqls (Figure 4 D) somewhat sn.raller, usuiri ly strongly inrolledtowards the apices hence appearing ntore acute in outl ine bearlrrg insideabove 1he tube a flexuose indumentun of crisped weakly nodose hairi up to7 8 mm long, twistcd between the nodes and more or less intricate rvith eachother, rising marginally on one or on either side.

. Only two adjacent stanens fertile (Figure 4 E), their slightly liattenedupright f i laments 8 5 to 12 mm iong and up to 0. 3 mm w jdc ln tire m jddle,abruptly subulate below the antlters and r.rot dilated i[ lower portions exceptfor the very base; stontinodes reduced to l-3 mm in length and hidden in partamong tlre copious hairlets up to l 3 mm long and distinctly nodose, surround-rng.the strminrl cup (ca L.b nrrr high) on its outer distal f 'ace, regularlybordering the free ring up to 0.6 mnr high ard not oblique. No pieudo-staminodes prcsent. ,.1rlrelJ ycllow, almost giobose,0 9 I.0 mm in diameter.

_ PLrll l shortly stalked, the sripes 0 6 0.8 mn] in length. Ovary club-shaped en t i re ly g labrous , 2 .2 2 .8 x 1 .0 l 4 min l inc lud ing the s t ipes) . Sry /elateral (Figure 4 F), straight, slendcr. up to 1l nrnr long and ca 0 l2 mm diam.in the mlddle, scarcely thickened downwards. S/Eri4 capitellate, papil lose,nore or less level with the anthers

Distussiort. The new taxon is based on one collcction consisting of f ivefragments. It bears some resemblance to other low, shrubby species with woodystems i.e. P. parvifolius (F. Muell.) F. Muell., P. polukii F. Muell.. p. remoti-.f lorus Benl characterised by spalsely tomcntose slioors, more or less persistentbases of thc leaves, a hirsutc staminal cup and a[ eccentrjc style. However,none of these plants ever develops the large long-pedunculale spikes brolderthan long and eviclently not becoming ovoid, as shown by the lew taxon; inaddition, the bracteoles of the lew species are considerably larger than the bractsand the tepals conspicuously narrower than in any all ied species. The cobweb-like indumentum does not occur in related species.

. P. rcmotif lorus from the Grcgory North District (Queensland) is unique forrts extremely heterogereous pubescence, and its ovary being distinctly hirsr"rtetoward the summit. ln P. polakii the staninal ring is very oblique, its ovarylong-stipitate, the pubescelce of t lte shoots is more or less restricted to leafaxils; arrd the leaves are larger and broadcr. P. purviJblius has glaucousbranches and (sub)glabrous bracls. In th'e tar. parvifolius the tepals aie cil iatcat the margins rvith shorter hairs, and the dense dorsal indurnentirm projects toor beyond the apices of the tepals, thus rendering the flowers very dif lerent fromtlrose of P. beqrdii. It P. porriJblius var. /actil.! thc tepals are broadly limbate.

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l l j i , j t^:::],1,"^*ard its base (,Figure b C). Bracreotes strorrer (0.7 O.q mm)DUt b roader (U.9 1 l mrn) . g labrous ar rd uho l l y t ransp l ren t (F igures 6 D a l ldo L) . tmbncate . compleLe ly embrac ing the per ian lh i r i i t s Io \ . \es i p l r t ; m idr ibnor promtnent.

. Perianth mauve to reddish with a very low and somewhat notched basal9i',.^Plld 9t lf.

lowermosL porrions of .r-he repal.: baie catiose insiOe: apicesL,l?-.,?,j1.:..1!uye.. A| .repals .f inall1 subequal.in Iength. glabrous i lrernally.? : : r ln9 1

sca l ' I ou le r a p rca | . pube(c-ence : rhe ha i rs , r ra ighr .0 . . )_0 .4 mm lon ! .slout. tutr-erculale. soon breaking off. Ou,ter tepals rigid. bort_shaped lFiguiel^i,,:,rT l., i :, -\ | mm. scarcely or slighrll margined. more or iess sharplyKee leo and d t \e rg tng rn an thes is . thesc i ln ry pube,cence res t r i c red ro the upp i r -most par t . fnner t tpa ls some\ \ha l b roade i t r rd f la r re r , ru ry jnq in dera i l i o t5nape bu t regu la r ly J - r ibbed in Lhe i r r red ian area . the mem'bra-nous marg ina lr^::""^"8,:0 4:]rm,wjde: beside.the apical hairlets there are somiwhat lo"nger{up ro u.o mm) but thrnner ones (i.e. looking less tuberculate under the micio_scope) developed along the margin, more sca'itered and appressed lFigure 6 Gy.. All sla.me.ns consrantly perfect; filaments slightly flattened and dilatedrowrrq rnelr base, 1.4_2 mm long, to 0.2 mm wide near the middle andPl",r g-T.d,l-. 91 4 0 o m,m. fu,ed bilow inro a compararirety t,igl, t.u r ,nlnrand I ree . .n tembrano ls . b road ly tu rb in te cup ( f igure o H j , a t iached to thepeflanrn crbc by rts towest portjon (ca 0.3 mm). Anthers broadly ell iptical,0 . 1 8 x 0 . 1 2 m n r -

^ l i t l i ! ( - f ig , r r . . O l ) a lmost sess i le uhen fu l ty dere loped. en t i re ly g labrous .

?r : r ! , y? , " l j .T . . , to "q and 0 .6 mm acros , . Sr .y /e cer r r ra l . i r ra ighr . car ' z mm rong and lh tckened downward \ . S l8za s r rbg lobose.

Spec nens examinedWestern Australia: N. W. Coast, De. Bouley, s.n., no date BM. This p)allt.was misidentified as "Ptilotus roseo-.tlbus Faimar,,'with reference to the tfpe atKew Herbarium. But the type specimen of this taxon ctearlv stiows the closel]i]iT: j9 r,. gomphrmoides F. Muell. ex Benth., having a dense woollyrndu mentum on the per ian th .Lat. 18'10', Long. 12226', A. Forrcst & T. Carel.t, s.n., lg79_MEL (77349)He;wood ls lands{southern is landt .B^o ] rupa- r teArch ipe lago. . .g rowing inAcac iascrub on sandsrone". P. G. t4 i lson 10q04. ZZ VaV f bli '-pf

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Byam Martin Island, Bonaparte Archipelago, 124.22,E, 15.24,5, p. G. llilson11513 (typus), l l Julv 1973-M. PERTH.

-

Discussion. To some extent the above described platt approaches p. conicusR. Br . ,and P. corymbosus R. Br . in genera l hab i t . p . ' c r r io ,s . -no"erer , l smarkedly separcted from all other species of the genus by the peculiar structureof,the androecium, y{19. p, coryibosus lwith iis strongei "5iy-Uor. urp""t1be longs ro a Eroup o f p t i lo t iu iLh b ipa f l i te inner tepa ls .

- "

. . Ha i rs resem.b l ing lhe roug l ) -ua l led ones o f the new spec ies are to be .eenrn the ye l loursh IndumeDtum o l p . a r t l to lqs ius F . Mue l l . They hare a s imi la rscabrid appearance, but are clearly septate rlith sometilnes imooth walts ofthe internodes especially toward tireir

.apices_and are -or" oi-G, restricted

to stens and foliage. . The perianth is characterized by long silky trichomes up: : ,^1: ]T i i l . lc , l rnd uidety erceeding r t re repal \ . i t , i , ip. . i . i i i not ctoset irelated to p. Lkcelva 6. The satne is the case uitlt p. forristi i f. Muell. all ieilto P. arthrolasius.

, WJth_regard to the pubesc:nce ofthe tepals there seems to be some resemb_tance Io 1:..tophotrichus tsenl. In the latter species the tepals ltave an apicaljiJr"ol J:liriyely. fy trajrs. Bur in that species'the apicar h;ir; a;;;uch ldnserl,o. z Til. JorDred and.smooth: the outer and inner _tepals bear short crispedharrs at tt)err external base, the inner ones_markedly narrowed below- lare

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