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    1. Introduction

    Permian sediments within the Talchir, Mand-Raigarh and

    Korba coalfields, all part of Mahanadi Master Basin, have

    been studied intensively by numerous workers during the

    past 15 years. However, the Ib River Coalfield has never

    been investigated thoroughly for plant fossils. During sev-

    eral field trips, the Ib River Coalfield was thoroughly

    searched for plant fossils to contribute to a synthesis of

    floristic diversity, early developmental history and evolu-

    tion of the flora throughout the Mahanadi Master Basin.

    The Ib River Coalfield is located in the southeastern

    part of NW-SE trending Mahanadi Master Basin between

    2130' to 2206' N and 8337' to 8410' E. The Ib RiverCoalfield is named after the river Ib, a tributary of the

    Mahanadi River. Plant megafossils were collected from

    deposits at Gopalpur Pahar and Laxamanpur Pahar, which

    are currently mapped as Barren Measures (Middle Permian)

    and Upper Kamthi formations (Triassic) respectively (GSI

    1997; CMPDI 1987). The complete flora includes three

    genera along with scale leaves. They are Neomariopteris

    (N. hughesii), Glossopteris (Glossopteris indica, G. commu-

    nis, G. tenuifolia, G. rewaensis, G. intermedia, G. zeilleri,

    G. decipiens, G. browniana, G. gigas, G. subtilis, G. inter-

    mittens, G. karharbariensis, G. tenuinervis, G. spatulata)

    and Vertebraria (V. indica). The present study includes

    descriptions of these 16 plant taxa from the coalfield.

    Contrary to the current understanding of the geology of the

    region, the megafloral assemblages from the two study

    areas are typical of the Lower Kamthi/Raniganj Formation

    (of Late Permian age). This assessment is supported by ear-

    lier palynological work carried out by Goswami (1997),

    which recorded the presence of Late Permian palynoassem-blages in fossiliferous Gopalpur deposits (table 1). Since

    both localities are situated fairly close to regions mapped as

    Lower Kamthi Formation, it is proposed here that the expo-

    sures should be re-evaluated in terms of their currently

    accepted lithostratigraphy.

    Record of Lower Gondwana megafloral assemblage from LowerKamthi Formation of Ib River Coalfield, Orissa, India

    SHREERUP GOSWAMI*

    Birbal Sahni Institute of Palaeobotany, 53 University Road, Lucknow 226 007, India

    *P.G. Department of Environmental Science, Fakir Mohan University, Vyasa Vihar, Balasore 756 019, India

    (Fax, 91-6782-264244; E-mail, [email protected])

    Recent investigations carried out in the Ib River Coalfield, Mahanadi Master Basin, Orissa, identified some fossilif-

    erous beds in the Lower Gondwana deposits. Two exposures of the Lower Kamthi Formation yielded diverse andabundant plant remains, which include Neomariopteris, Vertebraria, and a scale leaf along with 14 Glossopteris

    species otherwise mapped as Barren Measures and Upper Kamthi formations. Glossopteris indica dominates the

    flora (22.78%) followed by G. communis (17.72%) and G. browniana (13.92%). Based on megafloral assemblages,

    different beds exposed at Gopalpur and Laxamanpur Pahar are assigned here to the Lower Kamthi Formation (Late

    Permian). The floristic composition suggests that a warm and humid climate prevailed during the Late Permian. The

    status of the Kamthi Formation in the Ib River Coalfield has been redefined in the present study.

    [Goswami S 2006 Record of Lower Gondwana megafloral assemblage from Lower Kamthi Formation of Ib River Coalfield, Orissa, India;J.Biosci.

    31 115128]

    http://www.ias.ac.in/jbiosci

    Keywords. Glossopteris; Ib River Coalfield; Kamthi Formation;Neomariopteris; Vertebraria.

    J. Biosci. 31(1), March 2006, 115128, Indian Academy of Sciences 115

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    2. Previous palaeobotanical studies

    Megafloral assemblages of Lower Kamthi Formation inTalchir Coalfield are recorded from mainly two localities viz.,Handapa and Madhupur. The complete flora have been des-cribed by Khan (1969); Surange and Maheshwari (1970);Surange and Chandra (1973a,b,c; 1974a,b,c,d; 1978); Chandraand Rigby (1981, 1983); Chandra (1984); Pant et al(1985);Singh and Chandra (1987, 2000) and Chandra and Singh

    (1986, 1988, 1989, 1992, 1995). The megafloral assemblages(having leaf, root, seed and fertile forms) recovered from these

    beds consist of six groups, namely Lycopodiales, Sphenophyl-lales, Equisetales, Filicales, Cycadales, Ginkgoales andGlossopteridales comprising 18 pteridophytes and 77 gym-nosperm taxa. The number of Glossopteris species (46) recov-ered from Lower Kamthi Formation deposits of this coalfieldis the largest of any specific diversity of the genus in anyLower Kamthi beds of Indian Gondwana. Altogether 31 plant

    Shreerup Goswami116

    J. Biosci. 31(1), March 2006

    Table 1. Stratigraphic nomenclature for Ib River Coalfield

    Recent Alluvium/ Recent gravel and conglomerate

    Laterite

    Early to Upper Upper Conglomerate, red shale with

    Middle Gondwana Kamthi Dicroidium flora and coarse ferruginous-

    Triassic =Kamthi sandstone with clasts.Estheriids such as

    Cyzicus sp. andPaleolimnoids sp. are also

    reported (Pal et al 1992) (150 m +)

    --------------------------------------------------------Uncomformity-----------------------------------------------------------------------------------------

    Late Lower Lower Fine to medium grained well sorted

    Permian Gondwana Kamthi sandstone, siltstone, clay bed, coal, shale with

    =Raniganj broad mesh forms Glossopteris species, some

    arthophytes and ferns (Goswami 2002 and the present

    study). Palynofloral assemblage is dominated by

    Striatopodocarpites, Crescentipollenites,

    Faunipollenites, Arcuatipollenites andDensipollenites

    (Maiti 1994; Meena 1998;

    Meena and Goswami 2004; Goswami 1997)

    (180 m)

    Middle Barren Grey shale, carbonaceous shale,

    Permian Measures fine to coarse-grained sandstone,

    clay and ironstone nodules/shale (250 m +)

    Early Upper Micaceous subarkosic sandstone, grey

    Permian Barakar and carbonaceous shales with plenty of

    Glossopteris, some ferns and arthrophytes,

    fireclay and thick coal seams (Goswami 2002).

    Palynofloral assemblage is dominated by

    Scheuringipollenites, Faunipollenites,

    Striatopodocarpites, Punctatisporites and

    Horriditriletes etc (Tiwari 1968; Meena 1999, 2000)

    Lower Feldspathic sandstone, grey and

    Barakar carbonaceous shales with

    Gangamopteris-Noeggerathiopsis-Euryphyllum fossil

    assemblage (Goswami 2002) and thick coal seams.

    Palynofloral assemblage is dominated by

    Brijrajisporites, Rhizomaspora, Primuspollenites,

    Lahirites, Sulcatisporites, Cuneatisporites, Platysaccus

    andApiculatisporis (Tiwari 1968) (350500 m)

    Early Permian Karharbari Conglomerate, carbonaceous sandstone with

    fresh feldspar grains containing thin coal bands

    only along the NW margin of the basin (3065 m)

    Early Permian Talchir Diamictite, greenish sandstone, olive and chocolate

    coloured needle shales and rhythmites (130 m+)

    --------------------------------------------------------Uncomformity-----------------------------------------------------------------------------------------

    Precambrian Granites, gneisses, amphibolites and migmatites

    Age Group Formation Lithology and fossil content (thickness in meters)

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    genera and 95 species are recorded. It is one of the richestpalaeoflora of Lower Kamthi Formation of Indian Gondwana.

    Contrary to this, less number of plant taxa are recorded

    from the Lower Kamthi Formation of Ib River Coalfield.

    Feistmantel (1880) reported Schizoneura gondwanensis,

    Vertebraria indica, Sphenopteris sp. cf. S. polymorpha and

    four species of Glossopteris, viz.; Glossopteris communis, G.indica, G. damudica and G. browniana from the Lower

    Kamthi sediments exposed at Garjan hill, Girundla, Kodaloi

    and Belpahar areas. Pandey and Chakraborty (1964a,b)

    reported a fossil assemblage in the Lower Kamthi beds which

    included G. indica, G. browniana, S. gondwanensis, V. indica

    and Pecopteris sp. Singh (2000) reviewed that only above

    mentioned eight taxa are recorded from Lower Kamthi

    Formation of this coalfield. Subsequently a luxuriant repre-

    sentation of the pteridophytes and gymnosperms has been

    observed in two Lower Kamthi exposures (i.e. Sitaram Pahar,

    Dungri Pahar) of this coalfield by Goswami (1997, 2002).

    The mega floral assemblage consists of Equisetales such as

    equisetaceous stems, S. gondwanensis, Filicales such asNeomariopteris hughesii, Cycadales such as Macrotaenio-

    pteris feddenii, M. wianamattae, Glossopteridales such as

    eleven Glossopteris species and V. indica. It depicts that a rel-

    atively low diversity of floral elements was known from this

    formation of the Ib River Coalfield until the work of

    Goswami (1997, 2000). Hence to strengthen the floral diver-

    sity, two more Lower Kamthi exposures (Gopalpur Pahar and

    Laxamanpur Pahar) have been extensively searched out for

    plant fossils and are included in the present study. As a result,

    altogether 30 taxa are reported from different Lower Kamthi

    exposures of this sub-basin.

    Palynological assemblage from the sediments exposed in

    Gopalpur Pahar has the dominance of Striatopodocarpitesfol-lowed by Crescentipollenites, Faunipollenites, Distriatites

    and Arcuatipollenites. The presence of Verticipollenites,

    Densipollenites magnicorpus and Striatites in low percentage

    in this assemblage is significant. The appearances of

    Goubinispora, Trabeculosporites, Klausipollentites, Chorda-

    sporites, Arcuatipollenites and Falcisporites in the assem-

    blage render Late Permian age to this Palynozone (Goswami

    1997). The overall character of the assemblage compares

    well with the Late Permian Raniganj microflora RIA assem-

    blage of Tiwari and Singh (1986), Striatopodocarpites-

    Crescentipollenites zone of Tiwari and Tripathi (1988) and

    Densipollenites magnicorpus Assemblage zone of Tiwari and

    Tripathi (1992). Similar Late Permian Raniganj microflorawere recorded in different boreholes (Boreholes IBT-2, 3, 4, 5,

    6, 7, IBSH-6, IBH-6, 16) drilled in this coalfield (Meena and

    Goswami 2004; Maiti 1994; Meena 1998, 2000).

    3. Geological observations

    The Ib River Coalfield occurs within the elongate Mahanadi

    Master Basin, which is oriented parallel to the structural grain

    of the underlying Precambrian rocks. The southern boundary

    of this master basin is defined by a major NW-SE lineament.

    The Ib River Coalfield is characterized by a northwesterly

    plunging synclinal flexure. The southwestern boundary of the

    basin is marked by a fault juxtaposing the Barren Measures

    and Raniganj formations against Precambrian rocks.

    This coalfield was first mapped by Ball (1871) and thenresurveyed by Mehta and Anandalwar (1960), Raja Rao

    (1982), CMPDI (1987), Chaudhury (1988), Mukhopadhyay

    (1987, 1989), Pal et al (1992) and GSI (1997). Based on

    these past investigations and current observations, the pro-

    posed stratigraphic nomenclature for the Ib River Coalfield

    is given in table 1.

    4. Occurrence, material and methods

    The megafossils described here were collected from two dif-

    ferent exposures i.e. Gopalpur Pahar and Laxamanpur Pahar

    (delineated as L1

    and L2

    in figure 1) within the coalfield

    (2130' to 2206' N and 8337' to 8410'E), Jharsuguda,

    Sundargarh Districts, Orissa during two excursions. They are:

    Lower Kamthi Latitude Longitude

    Formation Exposures

    (i). Gopalpur Pahar, 22 03'N 83 42' 30' 'E

    Sundargarh Dist.

    (ii). Laxamanpur Pahar, 21 41' 30' 'N 83 45' 30' 'E

    Jharsuguda Dist.

    4.1 Gopalpur Pahar and Laxamanpur Pahar

    Gopalpur Pahar is a 70 m tall hillock about 0.5 km northwest

    of Gopalpur village, Sundargarh District and just near the

    confluence of Basundhara and Chatudhara nalas streams.

    Similarly Laxamanpur Pahar is a 120 m tall hillock, about

    1 km southeast of Laxamanpur village, Jharsuguda District.

    It is predominantly the lower Kamthi Formation that is

    exposed at these localities, and the lithological successions

    at both sites are comparable. The fossiliferous beds, namely

    the pinkish red shale and yellowish shale of Gopalpur Pahar

    and the pinkish red shale, red compact shale of Laxamanpur

    Pahar, occur on the southern and eastern flanks of these

    hillocks respectively. The lithological successions that were

    documented at these localities are illustrated in figure 2.

    The total collection comprises 79 specimens, including 75

    examples of Glossopteris (14 species), two specimens of

    Vertebraria and a single specimen of Neomariopteris and a

    scale leaf (table 2). The morphological characters of the spec-

    imens were observed using a Leica low power binocular

    microscope; photographs were taken using a Nikon 35 mm

    camera. For identification and description of the Glossopteris

    species the methodology given by Chandra and Surange

    (1979) has been adopted. The number of specimens studied in

    Megafloral assemblage from Lower Kamthi Formation 117

    J. Biosci. 31(1), March 2006

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    the present collection and the percentage of each taxon has

    been given in table 3. Detailed repetitive descriptions and

    comparative accounts of the well-known species reported

    from Ib River Coalfield have been avoided. Distinguishing

    features of the species of Glossopteris are tabulated for com-

    parison in table 4. The species are classified into three broad

    categories based on visual aspects i.e. narrow mesh form,

    intermediate mesh form and broad mesh form Glossopteris

    species. The specific morphographic characters are represent-

    ed in thirteen major columns. Identification of the 14 species

    was generally based on these characters and comparison with

    the holotypes, either from photographs or the original speci-

    mens. Table 4 demonstrates that the identification up to the

    species level is not based on any single species-specific char-

    acter, but rather on diagnostic suites of characters. It was

    determined that the venation pattern and mesh type in the mid-

    dle part of the leaf tends to be more consistent, and therefore

    more reliable in the delimitation of species.

    5. Systematic palaeobotany

    The fossils identified from Gopalpur Pahar and

    Laxamanpur Pahar areN. hughesii, G. indica, G. communis,

    G. tenuifolia, G. rewaensis, G. intermedia, G. zeilleri,

    Shreerup Goswami118

    J. Biosci. 31(1), March 2006

    INDEX

    UPPER KAMTHI FM.

    LOWER KAMTHI FM.

    BARREN MEASURES FM.

    BARAKAR FM.

    TALCHIR FM.

    PRECAMBRIAN

    COAL SEAM

    FAULT

    FOSSIL LOCALITY

    STATE BOUNDARY

    BOREHOLE

    SURFACE SAMPLE SITE

    83 35' 83 40' 83 45' 83 50' 83 55' 84 0'

    22

    15'

    22

    10'

    22

    5'

    22

    0'

    21

    50'

    21

    45'

    21

    40'

    Figure 1. Geological map of Ib River Coalfield, District Sundargarh and Jharsuguda, Orissa (after GSI 1997).

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    G. decipiens, G. browniana, G. gigas, G. subtilis, G. inter-

    mittens, G. karharbariensis, G. tenuinervis, G. spatulata,

    scale leaf and V. indica (table 2).

    Neomariopteris Maithy 1974

    Division: Pteridophyta

    Order: Filicales

    Genus: Neomariopteris Maithy 1974

    Type species: Neomariopteris hughesii

    (Feistmantel) Maithy 1974

    Type Locality: Jharia Coalfield, Bihar

    (Raniganj Formation), India

    Neomariopteris hughesii (Feistmantel) Maithy 1974

    (figure 3d)

    Locality: Laxamanpur Pahar

    Horizon: Lower Kamthi Formation

    Megafloral assemblage from Lower Kamthi Formation 119

    J. Biosci. 31(1), March 2006

    L

    o

    we

    r

    K

    a

    m

    t

    h

    i

    F

    m.

    L

    o

    w

    e

    r

    K

    a

    m

    t

    h

    i

    F

    m.

    1m

    1m

    0m 0m

    BASE IS NOT EXPOSED BASE IS NOT EXPOSED

    ALLUVIUM

    SANDSTONE

    SANDSTONE WITH

    SILTY LAYERS

    PEBBLY SILT STONE

    SILTY SHALE

    SHALE

    COMPACT SHALE

    FOSSILIFEROUS BED

    CROSS BEDDED

    Figure 2. Geological sections along Gopalpur Pahar and Laxamanpur Pahar.

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    Comparison: The fronds figured here compare very

    well with the specimens of N. hughesiidescribed by Feistmantel (1882, Pl. 12,

    fig. 3) and Maithy (1974, Pl. 2, fig.7) as

    they all have similar shape. Particularly

    their pinnules are lanceolate, have

    decurrent base and show sphenopteroid

    venation pattern.

    Distribution: Neomariopteris hughesii is known to

    occur in Karharbari, Barakar, Barren

    Measures, Raniganj and Kamthi forma-

    tions of Indian Gondwana.

    Vertebraria Royle 1839

    Division: Pinophyta Meyen 1984

    Order: Dictyopteridiales Rigby 1978

    (nom. Corr. McLoughlin 1990b)

    Genus: Vertebraria Royle 1839

    Type species: Vertebraria indica Royle 1839

    Type locality: Raniganj Coalfield (Barakar

    Formation), West Bengal, India

    Vertebraria indica Royle 1839 emend. Schopf 1965

    (figure 3c)

    Locality: Laxamanpur Pahar

    Horizon: Lower Kamthi Formation

    Comparison and Morphologically the current specimens

    remarks: closely resemble Holotype specimen

    described by Royle (1839) as they all

    consist of square areas in two linearrows, separated by a median longitudi-

    nal furrow and the areas are transverse-

    ly separated by grooves.

    Distribution: In India, Vertebraria indica occurs in

    Talchir, Karharbari, Barakar, Barren

    Measures, Raniganj and Kamthi forma-

    tions.

    Scale leaf

    (figure 5j)

    Division: Pinophyta Meyen 1984

    Order: Dictyopteridiales Rigby 1978 (nom.

    Corr. McLoughlin 1990b)

    Scale leafThere is a single specimen of scale leaf in the present col-

    lection. The length of the scale leaf is 1.8 cm, while breadth

    is 1.5 cm. No seed or sporangium is found associated with

    the scale leaf. Its apex is obtuse. Base is truncated and con-

    tracted. Lateral margins are concave. Veins arise from base,

    are dichotomous and divergent with interconnections. They

    are bifurcating and reaching up to apex. Meshes are narrow

    and elongate.

    Shreerup Goswami120

    J. Biosci. 31(1), March 2006

    Table 2. Distribution of plant megafossils from Lower Kamthi

    Formation exposed around Gopalpur Pahar and Laxamanpur

    Pahar, Ib River Coalfield.

    Taxa Gopalpur Pahar Laxamanpur Pahar

    Filicales (1)

    Neomariopteris

    hughesii +

    Glossopteridales

    leaf forms (14)

    Glossopteris indica + +

    G. communis + +

    G. tenuifolia + +

    G. rewaensis +

    G. intermedia +

    G. zeilleri + +

    G. decipiens +

    G. gigas +

    G. subtilis + +

    G. intermittens + +

    G. karharbariensis +

    G. tenuinervis +

    G. spatulata +

    G. browniana + +

    Glossopteridales

    root form (1)

    Vertebraria indica +

    Miscellaneous (1)

    Scale leaf +

    Total taxa 14 10

    Table 3. Relative percentage of fossil taxa from Lower Kamthi

    Exposures around Gopalpur Pahar and Laxamanpur Pahar, Ib River

    Coalfield.

    Number of

    specimens

    Taxa studied Percentage

    Dominant (>10%)

    Glossopteris indica 18 22.78

    G. communis 14 17.72

    G. browniana 11 13.92

    Sub-dominant (510%)

    G. zeilleri 6 7.59

    G. tenuifolia 5 6.33

    Rare (15%)

    G. decipiens 3 3.80

    G. intermittens 3 3.80

    G. subtilis 3 3.80

    G. rewaensis 2 2.53

    G. intermedia 2 2.53

    G. karharbariensis 2 2.53G. tenuinervis 2 2.53

    G. gigas 2 2.53

    G. spatulata 2 2.53

    Vertebraria indica 2 2.53

    Neomariopteris hughesii 1 1.27

    Scale leaf 1 1.27

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    Comparison and Morphologically the current specimen

    remarks: closely resembles the specimens

    described by Arber (1905), Walkom

    (1922), Surange and Maheshwari

    (1970), Chandra and Surange (1977a,b)

    as their veins are dichotomous and

    divergent with interconnections.

    Locality: Gopalpur Pahar

    Horizon: Lower Kamthi Formation

    Distribution: Scale leaves occur only in Karharbari,

    Barakar, Raniganj and Kamthi forma-

    tions of Indian Gondwana.

    Glossopteris Brongniart 1822Division: Pinophyta Meyen 1984

    Order: Dictyopteridiales Rigby 1978 (nom.

    Corr. McLoughlin 1990b)

    Genus Glossopteris Brongniart 1822

    Type species Glossopteris browniana Brongniart 1828

    Locality: Laxamanpur Pahar and Gopalpur Pahar

    Horizon: Lower Kamthi Formation

    There are 14 different Glossopteris species (G. indica,

    G. communis, G. tenuifolia, G. rewaensis, G. intermedia,

    G. zeilleri, G. decipiens, G. browniana, G. gigas, G. subtilis,

    G. intermittens, G. karharbariensis, G. tenuinervis, G. spat-

    ulata) in the present collection and their detailed descrip-tions are presented in the table 4.

    6. Discussion

    The sediments in and around Gopalpur Pahar section are

    shown as a part of the Barren Measures Formation (figure 1

    after CMPDI 1987, Chadhury 1988 and GSI 1997).

    However, the megaflora recovered in this study strongly

    advocates the presence of Late Permian sediments (Lower

    Kamthi Formation) in this particular area, as evidenced by

    the occurrence of very similar plant assemblages in the

    Lower Kamthi (=Raniganj) Formation. Similarly the sedi-

    ments in and around Laxamanpur Pahar section are shown

    as a part of the Upper Kamthi Formation (figure 1). The

    megaflora that has been recovered in this area is also very

    similar to plant assemblages typical of the Raniganj

    (=Lower Kamthi) Formation. This evidence, together with

    the absence of any typical Triassic element such as

    Dicroidium, Lepidopteris,points strongly to the presence of

    Late Permian sediments (Lower Kamthi Formation) in this

    particular region. It is therefore suggested that a detailed

    programme of geological mapping be undertaken to con-strain more precisely the distribution of Lower Gondwana

    sediments in these areas.

    Table 3 demonstrates that in the present study G. indica

    dominates the flora (22.78%) followed by G. communis

    (17.72%) and G. browniana (13.92%). The Presence of

    broad mesh forms of Glossopteris, namely G. rewaensis,

    G. subtilis, G. intermedia supports the current proposal that

    the sedimentary rocks under investigations belong to the

    Lower Kamthi/Raniganj Formation (Lele 1976). Broad mesh

    forms of Glossopteris such as G. rewaensis, G. subtilis, G.

    intermedia, G. conspicua, G. karanpuraensis and G. retifera

    are widely considered to be index fossils of Lower Kamthi

    sediments depicting Late Permian age (Lele 1976; Chandra

    and Chandra 1988; Chandra and Singh 1992; Singh and

    Chandra 2000). The present megafloral assemblage of Lower

    Kamthi Formation is comparable with the flora of Sitaram

    Pahar and Dungri Pahar of this coalfield as both the assem-

    blages containN. hughesii, G. indica, G. communis, G. spat-

    ulata, and G. gigas (Goswami 1997, 2002). The flora is also

    comparable with Handapa and Madhupur localities in the

    adjoining Talchir Coalfield (Chandra and Singh 1992; Singh

    Megafloral assemblage from Lower Kamthi Formation 121

    J. Biosci. 31(1), March 2006

    Table 4. Classification of Kamthi Formation in Ib River Coalfield (Chandra and Chandra 1988; Lele 1976).

    Lithology with

    Age Formation fossil contents Vegetation Palaeoclimate

    Triassic Upper Kamthi= Conglomerate, red shale withDicroidium Dense forest, small Warm, humid,

    Kamthi flora, coarse ferruginous sandstone with trees growing under temperate with

    clasts.Estheriids such as Cyzicus sp. and neath big tree, upland dry and semi-aridPaleolimnoids sp. are also reported vegetation spells

    (Pal et al1992)

    Late Lower Kamthi= Fine to medium grained well-sorted sandstone, Thick swampy, dense Warm, humid,

    Permiar Raniganj siltstone, clay bed, coal, abundance of trilete forest with small lakes, temperate with

    mioflora. Megafloral assemblages include ponds, low lying river intermittent rain

    Schizoneura Neomariopteris and broad mesh valley falls

    forms Glossopteris species (Goswami 2002

    and present study). Palynofloral assemblage is

    dominated by Striatopodocarpites,

    Crescentipollenites, Verticipollenites and

    Densipollenites

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    Shreerup Goswami122

    J. Biosci. 31(1), March 2006

    Figure 3. (a) Glossopteris indica Schimper 1869, Specimen No. 17/6291, x 1.1, Lower Kamthi Formation, Gopalpur Pahar.

    (b) Glossopteris communis Feistmantel 1879 Specimen No. 11/6294, x 1.1, Lower Kamthi Formation, Laxamanpur

    Pahar. (c) Vertebraria indica Royle 1839, Specimen No. 2/6294, x 1, Lower Kamthi Formation, Laxamanpur Pahar.

    (d) Neomariopteris hughesii (Zeiller) Maithy 1974, Specimen No. 1/6294, x 1, Lower Kamthi Formation, Laxamanpur Pahar.

    (e) Glossopteris rewaensis Chandra and Surange 1979, Specimen No.12/6291, x 2, Lower Kamthi Formation, Gopalpur Pahar.

    (f) Glossopteris intermittens Feistmantel 1881, Specimen No. 7/6291, x 1.5, Lower Kamthi Formation, Gopalpur Pahar.

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    Megafloral assemblage from Lower Kamthi Formation 123

    J. Biosci. 31(1), March 2006

    Figure 4. (a) Glossopteris spatulata Pant and Singh K B 1971, Specimen No. 3/6291, x 1.1, Lower Kamthi Formation, Gopalpur Pahar.

    (b) Glossopteris communis Feistmantel 1879 Specimen No. 6/6294, x 1, Lower Kamthi Formation, Laxamanpur Pahar. (c) (a)

    Glossopteris decipiens Feistmantel 1879, Specimen No. 5/6291, x 1, Lower Kamthi Formation, Gopalpur Pahar. (b) Glossopteris brown-

    iana Brongniart 1828, Specimen No. 9/6291, x 1, Lower Kamthi Formation, Gopalpur Pahar. (d) Glossopteris subtilis Pant and Gupta

    1971, Specimen No. 8/6294, x 1.5, Lower Kamthi Formation, Laxamanpur Pahar.

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    Shreerup Goswami124

    J. Biosci. 31(1), March 2006

    Figure 5. (a) Glossopteris gigas Pant and Singh K B 1971, Specimen No. 24/6291, x 1, Lower Kamthi Formation, Gopalpur Pahar. (b)

    Glossopteris tenuinervis Pant and Gupta 1971, Specimen No. 14/6294, x 1.3, Lower Kamthi Formation, Laxamanpur Pahar. (c) Glossopteris

    tenuifolia Pant and Gupta 1968, Specimen No. 19/6291, x 1, Lower Kamthi Formation, Gopalpur Pahar. (d) Glossopteris karharbariensis

    Chandra and Surange 1979, Specimen No 27/6291, x 1, Lower Kamthi Formation, Gopalpur Pahar. (e) Glossopteris indica Schimper 1869,

    Specimen No. 31/6291, x 1, Lower Kamthi Formation, Gopalpur Pahar. (f) Glossopteris browniana Brongniart 1828, Specimen No.

    29/6291, x 1, Lower Kamthi Formation, Gopalpur Pahar. (g) Glossopteris zeilleri Pant and Gupta 1968, Specimen No. 26/6291, x 1, Lower

    Kamthi Formation, Gopalpur Pahar. (h) Glossopteris intermedia Feistmantel 1880, Specimen No. 36/6291, x 1.5, Lower Kamthi Formation,

    Gopalpur Pahar. (i).Apical part of the leaf, Glossopteris zeilleri Pant and Gupta, 1968, Specimen No. 21/6294, x 1.4, Lower Kamthi

    Formation, Laxamanpur Pahar. (j) Scale leaf, Specimen No. 2/6291, x 1.4, Lower Kamthi Formation, Gopalpur Pahar.

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    MS received 18 February 2005; accepted 19 December 2005

    ePublication: 14 February 2006

    Corresponding editor: VIDYANANDNANJUNDIAH