-10 1637 10 1918Jones 1934 0.4% 1944 56% 10 1956 1960 1963 10 10 10 10 10 10Stuber(1992)RFLP(QTL) 10 10 10 P152 10 10 10 1 10 2 10 3 10 10 10 10 10 10 10 2 10 10 10 2 10 10 10 10 10 10 10 10 10 F1 10 10 10 2 10 3 10 2 (test-crossing),(tester)(test cross variety) 10 10 10 10 10 10 10 A123……n1× A2× A3× A……n× AA×1 A×2A×3…… A×n GCA 10 10 10 10 10 10 10 10 10 10 1 10 10 10 3A×B×C 10 3 (double cross hybrid) 42A×B×C ×D 10 10 10 10 10 “” 10 10 10 × × 1 2